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<document id="9CF84CFF60D56E171FD98BD8A09CF315" ID-DOI="10.11646/zootaxa.4103.4.1" ID-GBIF-Dataset="cbd5532b-82f3-4fc1-a7f3-cd1a0e795c49" ID-ISSN="1175-5326" ID-Zenodo-Dep="264759" ID-ZooBank="B6428194-1C7A-4A9F-9038-EE4A78222A06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460614658210" checkinUser="plazi" docAuthor="Liu, Xingyue, Lu, Xiumei &amp; Zhang, Weiwei" docDate="2016" docId="03FBD81AAF28FFA9FF58FA28AEC0A689" docLanguage="en" docName="zootaxa.4103.4.1.pdf.imf" docOrigin="Zootaxa 4103 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhynchoraphidia Liu, Lu &amp; Zhang, 2016, gen. nov." docType="treatment" docVersion="9" lastPageNumber="312" masterDocId="FFC2A062AF2FFFA2FFCFFFC0AF18A724" masterDocTitle="New genera and species of the minute snakeflies (Raphidioptera: Mesoraphidiidae: Nanoraphidiini) from the mid Cretaceous of Myanmar" masterLastPageNumber="324" masterPageNumber="301" pageNumber="308" updateTime="1698329959600" updateUser="plazi">
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<mods:title id="3E0436731B147608D6FAFE6EE9A885DE">New genera and species of the minute snakeflies (Raphidioptera: Mesoraphidiidae: Nanoraphidiini) from the mid Cretaceous of Myanmar</mods:title>
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<mods:namePart id="25E9232CC021ED0E0B04F5D52276C6C1">Liu, Xingyue</mods:namePart>
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<mods:namePart id="03477F6CC5638BC799CDA55AC592D801">Lu, Xiumei</mods:namePart>
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<mods:namePart id="B5284F4C775215E131148DF659AF29E8">Zhang, Weiwei</mods:namePart>
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<mods:date id="0D83307628B28AF7FF9D62262EECBF43">2016</mods:date>
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<treatment id="03FBD81AAF28FFA9FF58FA28AEC0A689" ID-DOI="http://doi.org/10.5281/zenodo.6055081" ID-GBIF-Taxon="119568745" ID-Zenodo-Dep="6055081" LSID="urn:lsid:plazi:treatment:03FBD81AAF28FFA9FF58FA28AEC0A689" httpUri="http://treatment.plazi.org/id/03FBD81AAF28FFA9FF58FA28AEC0A689" lastPageId="11" lastPageNumber="312" pageId="7" pageNumber="308">
<subSubSection id="C3483A87AF28FFA5FF58FA28AE09A107" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="8BED690CAF28FFA5FF58FA28AD2FA126" blockId="7.[151,567,1511,1571]" box="[151,567,1511,1538]" pageId="7" pageNumber="308">
<heading id="D0A5DE60AF28FFA5FF58FA28AD2FA126" bold="true" box="[151,567,1511,1538]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<emphasis id="B926B51EAF28FFA5FF58FA28AD2FA126" bold="true" box="[151,567,1511,1538]" pageId="7" pageNumber="308">
Genus
<taxonomicName id="4C52128FAF28FFA5FF21FA27AEDAA125" box="[238,450,1511,1537]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF28FFA5FF21FA27AEDAA125" bold="true" box="[238,450,1511,1537]" italics="true" pageId="7" pageNumber="308">Rhynchoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2150865AF28FFA5FE06FA28AD2FA126" box="[457,567,1512,1538]" pageId="7" pageNumber="308" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED690CAF28FFA5FF58F9CAAE09A107" blockId="7.[151,567,1511,1571]" box="[151,273,1546,1571]" pageId="7" pageNumber="308">
(
<figureCitation id="13697589AF28FFA5FF50F9CAAE11A107" box="[159,265,1546,1571]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1816,1838]" captionStartId-1="10.[151,250,1383,1405]" captionTargetBox-0="[168,1428,390,1784]" captionTargetBox-1="[315,1270,200,1348]" captionTargetId-0="figure@9.[151,1435,375,1795]" captionTargetId-1="figure@10.[303,1284,193,1362]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 5. Rhynchoraphidia burmana gen. et sp. nov., holotype EMTG BA- 002256. A. Habitus in dorsal view; B. Head and thorax in dorsal view; C. distal parts of left wings; D. Drawing of head and prothorax in dorsal view. Arrow indicating pterostigmal crossvein. Scale bar = 1.0 mm." captionText-1="FIGURE 6. Rhynchoraphidia burmana gen. et sp. nov. A. Holotype EMTG BA- 002256, drawing of right fore- and hindwing; B. same specimen, drawing of left fore- and hindwing. Scale bar = 1.0 mm." httpUri-0="https://zenodo.org/record/264764/files/figure.png" httpUri-1="https://zenodo.org/record/264765/files/figure.png" pageId="7" pageNumber="308">Figs. 56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF28FFA5FF58F995AD8FA14F" box="[151,663,1618,1643]" pageId="7" pageNumber="308" type="materials_examined">
<paragraph id="8BED690CAF28FFA5FF58F995AD8FA14F" blockId="7.[151,663,1618,1643]" box="[151,663,1618,1643]" pageId="7" pageNumber="308">
<typeStatus id="54E9D7AEAF28FFA5FF58F995AFD1A14E" box="[151,201,1621,1642]" pageId="7" pageNumber="308">Type</typeStatus>
species:
<taxonomicName id="4C52128FAF28FFA5FEE9F994AD20A14E" box="[294,568,1620,1642]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="burmana" status="sp. nov.">
<emphasis id="B926B51EAF28FFA5FEE9F994AD20A14E" box="[294,568,1620,1642]" italics="true" pageId="7" pageNumber="308">Rhynchoraphidia burmana</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF28FFA5FDF2F992AD8FA14F" bold="true" box="[573,663,1618,1643]" pageId="7" pageNumber="308">
<taxonomicNameLabel id="A2150865AF28FFA5FDF2F992AD8FA14F" box="[573,663,1618,1643]" pageId="7" pageNumber="308" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF28FFA5FF58F957AC47A0D0" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="8BED690CAF28FFA5FF58F957AC47A0D0" blockId="7.[151,1437,1687,2037]" pageId="7" pageNumber="308">
<emphasis id="B926B51EAF28FFA5FF58F957AE09A194" bold="true" box="[151,273,1687,1712]" pageId="7" pageNumber="308">Diagnosis.</emphasis>
Minute size. Head ovoid, with portion posterior to compound eyes shorter than eye length and tapering caudad; clypeus slightly prolonged and distinctly narrowed, nearly as long as compound eye length; compound eyes large and ovoid; ocelli absent; antennae possibly long. Pronotum slightly shorter than head, with a constant height along entire length. All tibiae swollen, lacking specialized spines; tibial spurs absent. Wings narrowly elongate. Forewing: Costal space distinctly broadened at its midpoint; pterostigma elongate, ~1/4× wing length, medially with a pterostigmal crossvein, proximally closed by a crossvein and distally closed by a veinlet of RA; Scp terminating into C slightly proximad wing midpoint; posterior veinlet of RA continuous with stem of RA;
<collectingCountry id="F345299CAF28FFA5FAB5F8B0AA84A0A3" box="[1402,1436,1904,1927]" name="Poland" pageId="7" pageNumber="308">RP</collectingCountry>
with anterior branch proximally not angled; two radial cells, one discal cell, and one discoidal cell present. Hindwing: Stem of MA proximally fused with MP; CuA distally bifurcated; an oblique proximal mp-cua present; three radial cells, one discal cell, and one discoidal cell present.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF27FFAAFF08FF57AB79A7F1" pageId="8" pageNumber="309" type="etymology">
<paragraph id="8BED690CAF27FFAAFF08FF57AB79A7F1" blockId="8.[151,1436,151,536]" pageId="8" pageNumber="309">
<emphasis id="B926B51EAF27FFAAFF08FF57AE56A794" bold="true" box="[199,334,151,176]" pageId="8" pageNumber="309">Etymology.</emphasis>
From “
<emphasis id="B926B51EAF27FFAAFE60FF58AD12A794" box="[431,522,152,176]" italics="true" pageId="8" pageNumber="309">rhyncho</emphasis>
-”, rostrate, and
<taxonomicName id="4C52128FAF27FFAAFD00FF58AC2EA794" box="[719,822,152,176]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF27FFAAFD00FF58AC2EA794" box="[719,822,152,176]" italics="true" pageId="8" pageNumber="309">Raphidia</emphasis>
</taxonomicName>
, a common genus-group name of
<taxonomicName id="4C52128FAF27FFAAFB1BFF57AA6EA794" box="[1236,1398,151,176]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="order">Raphidioptera</taxonomicName>
, in reference to the prolonged and narrowed clypeus, of the new genus. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF27FFAAFF08FF1FAE09A5BE" pageId="8" pageNumber="309" type="discussion">
<paragraph id="8BED690CAF27FFAAFF08FF1FAEDCA53C" blockId="8.[151,1436,151,536]" pageId="8" pageNumber="309">
<emphasis id="B926B51EAF27FFAAFF08FF1FAE23A7DC" bold="true" box="[199,315,223,248]" pageId="8" pageNumber="309">Remarks.</emphasis>
The new genus appears to be closely related to
<emphasis id="B926B51EAF27FFAAFC9EFF20ACEBA7DC" box="[849,1011,224,248]" italics="true" pageId="8" pageNumber="309">Alavaraphidia</emphasis>
Pérez-de la Fuente, Peñalver, Delclòs &amp; Engel,
<quantity id="4CAAC4E9AF27FFAAFECAFEC4AE43A639" box="[261,347,260,285]" metricMagnitude="1" metricUnit="m" metricValue="5.11048" pageId="8" pageNumber="309" unit="in" value="2012.0">2012 in</quantity>
having similar prolonged clypeus and swollen tibiae, but can be separated from the latter genus by the absence of ocelli (three ocelli present in
<emphasis id="B926B51EAF27FFAAFD76FEE8AC43A664" box="[697,859,296,320]" italics="true" pageId="8" pageNumber="309">Alavaraphidia</emphasis>
) and the tarsomere 3 without digitiform processes (six to eighth digitiform processes present on bilobed extentions in
<emphasis id="B926B51EAF27FFAAFC63FE8CAB56A640" box="[940,1102,332,356]" italics="true" pageId="8" pageNumber="309">Alavaraphidia</emphasis>
). Among the nanoraphidiine genera,
<taxonomicName id="4C52128FAF27FFAAFF3FFEB0AEBCA6AC" box="[240,420,368,392]" class="Insecta" family="Mesoraphidiidae" genus="Rynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF27FFAAFF3FFEB0AEBCA6AC" box="[240,420,368,392]" italics="true" pageId="8" pageNumber="309">Rynchoraphidia</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF27FFAAFE64FEAFAD0DA6AC" bold="true" box="[427,533,367,392]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2150865AF27FFAAFE64FEAFAD0DA6AC" box="[427,533,367,392]" pageId="8" pageNumber="309" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
also shares the presence of pterostigmal crossvein with the contemporary genus
<taxonomicName id="4C52128FAF27FFAAFF58FE54AE5CA688" box="[151,324,404,428]" class="Insecta" family="Mesoraphidiidae" genus="Burmoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF27FFAAFF58FE54AE5CA688" box="[151,324,404,428]" italics="true" pageId="8" pageNumber="309">Burmoraphidia</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF27FFAAFE83FE54AEAEA689" bold="true" box="[332,438,404,429]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2150865AF27FFAAFE83FE54AEAEA689" box="[332,438,404,429]" pageId="8" pageNumber="309" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
while it can be distinguished from the latter genus based on the absence of ocelli (three ocelli present in
<taxonomicName id="4C52128FAF27FFAAFEA9FE78AD0BA6F4" box="[358,531,440,464]" class="Insecta" family="Mesoraphidiidae" genus="Burmoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF27FFAAFEA9FE78AD0BA6F4" box="[358,531,440,464]" italics="true" pageId="8" pageNumber="309">Burmoraphidia</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF27FFAAFDEEFE77AD89A6F4" bold="true" box="[545,657,439,464]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2150865AF27FFAAFDEEFE77AD89A6F4" box="[545,657,439,464]" pageId="8" pageNumber="309" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), the pterostigma proximally closed by a crossvein (pterostigma proximally diffused in
<taxonomicName id="4C52128FAF27FFAAFE75FE1CAD7FA6D0" box="[442,615,476,500]" class="Insecta" family="Mesoraphidiidae" genus="Burmoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF27FFAAFE75FE1CAD7FA6D0" box="[442,615,476,500]" italics="true" pageId="8" pageNumber="309">Burmoraphidia</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF27FFAAFDB6FE1CADF5A6D1" bold="true" box="[633,749,476,501]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2150865AF27FFAAFDB6FE1CADF5A6D1" box="[633,749,476,501]" pageId="8" pageNumber="309" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
), and the swollen metatibia (metatibia not swollen in
<taxonomicName id="4C52128FAF27FFAAFF58FDC0AE5CA53C" box="[151,324,512,536]" class="Insecta" family="Mesoraphidiidae" genus="Burmoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF27FFAAFF58FDC0AE5CA53C" box="[151,324,512,536]" italics="true" pageId="8" pageNumber="309">Burmoraphidia</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF27FFAAFE85FE3FAEABA53C" bold="true" box="[330,435,511,536]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2150865AF27FFAAFE85FE3FAEABA53C" box="[330,435,511,536]" pageId="8" pageNumber="309" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8BED690CAF27FFAAFF58FD9EAE09A5BE" blockId="8.[151,577,606,666]" pageId="8" pageNumber="309">
<taxonomicName id="4C52128FAF27FFAAFF58FD9EAEF9A55C" box="[151,481,606,632]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="burmana" status="sp. nov.">
<emphasis id="B926B51EAF27FFAAFF58FD9EAEF9A55C" bold="true" box="[151,481,606,632]" italics="true" pageId="8" pageNumber="309">Rhynchoraphidia burmana</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF27FFAAFE27FD9FAD59A55C" bold="true" box="[488,577,607,632]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A2150865AF27FFAAFE27FD9FAD59A55C" box="[488,577,607,632]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13697589AF27FFAAFF50FD41AE11A5BE" box="[159,265,641,666]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="9.[151,250,1816,1838]" captionStartId-1="10.[151,250,1383,1405]" captionTargetBox-0="[168,1428,390,1784]" captionTargetBox-1="[315,1270,200,1348]" captionTargetId-0="figure@9.[151,1435,375,1795]" captionTargetId-1="figure@10.[303,1284,193,1362]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURE 5. Rhynchoraphidia burmana gen. et sp. nov., holotype EMTG BA- 002256. A. Habitus in dorsal view; B. Head and thorax in dorsal view; C. distal parts of left wings; D. Drawing of head and prothorax in dorsal view. Arrow indicating pterostigmal crossvein. Scale bar = 1.0 mm." captionText-1="FIGURE 6. Rhynchoraphidia burmana gen. et sp. nov. A. Holotype EMTG BA- 002256, drawing of right fore- and hindwing; B. same specimen, drawing of left fore- and hindwing. Scale bar = 1.0 mm." httpUri-0="https://zenodo.org/record/264764/files/figure.png" httpUri-1="https://zenodo.org/record/264765/files/figure.png" pageId="8" pageNumber="309">Figs. 56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF27FFAAFF58FD09AEC0A5C6" box="[151,472,713,738]" pageId="8" pageNumber="309" type="diagnosis">
<paragraph id="8BED690CAF27FFAAFF58FD09AEC0A5C6" blockId="8.[151,1437,713,2034]" box="[151,472,713,738]" pageId="8" pageNumber="309">
<emphasis id="B926B51EAF27FFAAFF58FD09AE09A5C6" bold="true" box="[151,273,713,738]" pageId="8" pageNumber="309">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF27FFAAFF08FD2DAEA0A0A2" pageId="8" pageNumber="309" type="description">
<paragraph id="8BED690CAF27FFAAFF08FD2DAB60A40E" blockId="8.[151,1437,713,2034]" pageId="8" pageNumber="309">
<emphasis id="B926B51EAF27FFAAFF08FD2DAE41A422" bold="true" box="[199,345,749,774]" pageId="8" pageNumber="309">Description.</emphasis>
<typeStatus id="54E9D7AEAF27FFAAFEAEFD2DAED1A422" box="[353,457,749,774]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
EMTG BA-002256. Female. Body length
<quantity id="4CAAC4E9AF27FFAAFC7BFD2DAB14A421" box="[948,1036,749,774]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="309" unit="mm" value="5.1">5.1 mm</quantity>
(measured without ovipositor) and
<quantity id="4CAAC4E9AF27FFAAFF58FCD2AFF5A40D" box="[151,237,786,810]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="8" pageNumber="309" unit="mm" value="7.4">7.4 mm</quantity>
(measured with ovipositor). Body generally blackish brown except for pale legs.
</paragraph>
<paragraph id="8BED690CAF27FFAAFF08FCF5ACDFA302" blockId="8.[151,1437,713,2034]" pageId="8" pageNumber="309">
Head ovoid,
<quantity id="4CAAC4E9AF27FFAAFEAFFCF6AEA2A469" box="[352,442,822,846]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="309" unit="mm" value="0.9">0.9 mm</quantity>
long, with portion posterior to compound eyes shorter than eye length and tapering caudad; clypeus distinctly narrowed and slightly prolonged, nearly as long as compound eye length. Compound eyes large and ovoid, separated by distance slightly shorter than compound eye length. Ocelli absent. Antennae inserted at position slightly posteriad anterior tangent of compound eyes, with only proximal parts preserved; preserved part of antenna about 1.0 mm; scape subcylindrical, strongly robust; pedicel subcylindrical, about half length of scape, and slightly thicker than neighboring flagellomere; flagellum damaged, entire length unknown, but possibly long. Mouthparts not clearly visible, with short maxillary palps.
</paragraph>
<paragraph id="8BED690CAF27FFAAFF08FBF1AE4DA392" blockId="8.[151,1437,713,2034]" pageId="8" pageNumber="309">
Prothorax
<quantity id="4CAAC4E9AF27FFAAFE8FFBF1AE82A36D" box="[320,410,1073,1098]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="8" pageNumber="309" unit="mm" value="0.6">0.6 mm</quantity>
long, meso- plus metathorax
<quantity id="4CAAC4E9AF27FFAAFD36FBF2AC4CA36D" box="[761,852,1074,1098]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="309" unit="mm" value="0.9">0.9 mm</quantity>
long; pronotum slightly shorter than head, with a constant height along its entire length, and with anterolateral portions slightly expanded. Legs short, densely setose; all tibiae swollen and lacking spines; tarsomere 3 bilobed; pretarsal claws simple, with a basal enlargement; arolium reduced.
</paragraph>
<paragraph id="8BED690CAF27FFAAFF08FB01AE71A03E" blockId="8.[151,1437,713,2034]" pageId="8" pageNumber="309">
Wings hyaline, veins dark, with short setae. Forewing length
<quantity id="4CAAC4E9AF27FFAAFC4CFB02ACC5A3FD" box="[899,989,1218,1242]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="309" unit="mm" value="4.4">4.4 mm</quantity>
, maximum width
<quantity id="4CAAC4E9AF27FFAAFB62FB02AA1FA3FD" box="[1197,1287,1218,1242]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="309" unit="mm" value="0.8">0.8 mm</quantity>
; costal space distinctly broadened at its midpoint (at widest point costal space ~1.4× width of pterostigma); three costal crossveins present; ScP terminating into C slightly proximad wing midpoint; pterostigma elongate (
<quantity id="4CAAC4E9AF27FFAAFB33FACAAA4BA205" box="[1276,1363,1290,1314]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="309" unit="mm" value="1.1">1.1 mm</quantity>
long), ~1/4× forewing length, with constant width along its entire length, and medially with an oblique pterostigmal crossvein; pterostigma starting
<quantity id="4CAAC4E9AF27FFAAFE35FA92AD4AA24D" box="[506,594,1362,1386]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="mm" value="0.3">0.3 mm</quantity>
(~4.0× pterostigmal width) beyond termination of ScP, proximally closed by a crossvein and distally closed by anterior veinlet of RA; RA distally bifurcated, with posterior branch continuous with its stem;
<collectingCountry id="F345299CAF27FFAAFE73FA5AAEC6A295" box="[444,478,1434,1457]" name="Poland" pageId="8" pageNumber="309">RP</collectingCountry>
simple or with two short branches, anterior branch not zig-zagged with stem of
<collectingCountry id="F345299CAF27FFAAFABDFA5AAA80A295" box="[1394,1432,1434,1457]" name="Poland" pageId="8" pageNumber="309">RP</collectingCountry>
; MA with two branches; two radial cells and one discal cell present; MP with two long, distally bifurcated branches; one discoidal cell present; CuA distally bifurcated, with anterior branch partially fused with posterior branch of MP. Hindwing length
<quantity id="4CAAC4E9AF27FFAAFE42F9C5AEFCA139" box="[397,484,1541,1566]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="309" unit="mm" value="3.6">3.6 mm</quantity>
, maximum width
<quantity id="4CAAC4E9AF27FFAAFD60F9C6AC1FA139" box="[687,775,1542,1566]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="309" unit="mm" value="0.7">0.7 mm</quantity>
; costal space distinctly narrower than that of forewing; two costal crossveins present; pterostigma elongate (1.0 mm long), with constant width along its entire length, and with indistinct trace of a pterostigmal crossvein at distal 1/3; pterostigma starting
<quantity id="4CAAC4E9AF27FFAAFBD2F98EAB6FA141" box="[1053,1143,1614,1638]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="mm" value="0.3">0.3 mm</quantity>
(slightly more than 3.0× pterostigmal width) beyond termination of ScP, proximally closed by a crossvein and distally closed by anterior veinlet of RA; RA distally bifurcated, with posterior branch continuous with its stem;
<collectingCountry id="F345299CAF27FFAAFB91F956AB98A189" box="[1118,1152,1686,1709]" name="Poland" pageId="8" pageNumber="309">RP</collectingCountry>
with two short branches; MA with two branches, stem of MA proximally fused with MP; three radials cells and one distal cell present; MP with two long, distally bifurcated branches; one discoidal cell present; an oblique proximal mp-cua present; CuA distally bifurcated.
</paragraph>
<paragraph id="8BED690CAF27FFAAFF08F8E5AEA0A0A2" blockId="8.[151,1437,713,2034]" pageId="8" pageNumber="309">
Abdomen length
<quantity id="4CAAC4E9AF27FFAAFE40F8E5AEF0A019" box="[399,488,1829,1854]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="309" unit="mm" value="2.6">2.6 mm</quantity>
(measured without ovipositor) and
<quantity id="4CAAC4E9AF27FFAAFC49F8E6ACC7A019" box="[902,991,1830,1854]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="8" pageNumber="309" unit="mm" value="4.4">4.4 mm</quantity>
(measured with ovipositor). Genitalia: Genital sclerites not clearly visible; ectoproct small; ovipositor
<quantity id="4CAAC4E9AF27FFAAFCA4F889ACDBA045" box="[875,963,1865,1890]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="309" unit="mm" value="2.5">2.5 mm</quantity>
long and
<quantity id="4CAAC4E9AF27FFAAFBF8F88AAB96A045" box="[1079,1166,1866,1890]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="309" unit="mm" value="0.1">0.1 mm</quantity>
thick, distally tapering, with digitiform gonostyli.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF27FFAAFF08F851AD94A0D5" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="8BED690CAF27FFAAFF08F851AD94A0D5" blockId="8.[151,1437,713,2034]" pageId="8" pageNumber="309">
<emphasis id="B926B51EAF27FFAAFF08F851AE6DA08E" bold="true" box="[199,373,1937,1962]" pageId="8" pageNumber="309">
<typeStatus id="54E9D7AEAF27FFAAFF08F851AE1AA08E" box="[199,258,1937,1962]" pageId="8" pageNumber="309">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54E9D7AEAF27FFAAFEB4F851AEFBA08E" box="[379,483,1937,1962]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
<collectionCode id="ED43F1C9AF27FFAAFE24F852AD22A08E" box="[491,570,1938,1962]" pageId="8" pageNumber="309">EMTG</collectionCode>
BA-002256: amber piece preserving a complete adult female of
<taxonomicName id="4C52128FAF27FFAAFADEF852AA8DA08D" box="[1297,1429,1937,1961]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="burmana">
<emphasis id="B926B51EAF27FFAAFADEF852AA8DA08D" box="[1297,1429,1937,1961]" italics="true" pageId="8" pageNumber="309">R. burmana</emphasis>
</taxonomicName>
, a thripid, and a midge; it is polished in the form of a subtriangular cabochon, slightly obscure, with length×width about 24.1×
<quantity id="4CAAC4E9AF27FFAAFED3F819AE9BA0D5" box="[284,387,2009,2034]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="8" pageNumber="309" unit="mm" value="16.2">16.2 mm</quantity>
, height about
<quantity id="4CAAC4E9AF27FFAAFDEEF81AAD90A0D5" box="[545,648,2010,2034]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="8" pageNumber="309" unit="mm" value="10.7">10.7 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF26FFABFF08FF57AE56A7F1" pageId="9" pageNumber="310" type="etymology">
<paragraph id="8BED690CAF26FFABFF08FF57AE56A7F1" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="310">
<emphasis id="B926B51EAF26FFABFF08FF57AE56A794" bold="true" box="[199,334,151,176]" pageId="9" pageNumber="310">Etymology.</emphasis>
The specific epithet “
<taxonomicName id="4C52128FAF26FFABFD8DFF58ADBFA794" box="[578,679,152,176]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="burmana">
<emphasis id="B926B51EAF26FFABFD8DFF58ADBFA794" box="[578,679,152,176]" italics="true" pageId="9" pageNumber="310">burmana</emphasis>
</taxonomicName>
” refers to the occurrence of the new species in the Early Cretaceous Burmese amber.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF26FFA8FF08FF1FAD5DA156" lastPageId="10" lastPageNumber="311" pageId="9" pageNumber="310" type="discussion">
<paragraph id="8BED690CAF26FFABFF08FF1FAE0DA664" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="310">
<emphasis id="B926B51EAF26FFABFF08FF1FAE23A7DC" bold="true" box="[199,315,223,248]" pageId="9" pageNumber="310">Remarks.</emphasis>
By having the narrow wings and pterostigmal crossvein, the new species somewhat resembles
<taxonomicName id="4C52128FAF26FFABFA4EFF21AFC4A638" class="Insecta" family="Mesoraphidiidae" genus="Dolichoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="engeli" status="sp. nov.">
<emphasis id="B926B51EAF26FFABFA4EFF21AFC4A638" italics="true" pageId="9" pageNumber="310">D. engeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2150865AF26FFABFF2DFEC4AE24A639" box="[226,316,260,285]" pageId="9" pageNumber="310" rank="species">
<emphasis id="B926B51EAF26FFABFF2DFEC4AE24A639" bold="true" box="[226,316,260,285]" pageId="9" pageNumber="310">sp. nov.</emphasis>
</taxonomicNameLabel>
, but it can be distinguished from the latter species by the narrowly prolonged clypeus and the shape of
<collectingCountry id="F345299CAF26FFABFF7AFEE8AFCFA61B" box="[181,215,296,319]" name="Poland" pageId="9" pageNumber="310">RP</collectingCountry>
fork.
</paragraph>
<caption id="DF2D3984AF26FFABFF58F8D8AD4DA048" httpUri="https://zenodo.org/record/264764/files/figure.png" pageId="9" pageNumber="310" targetBox="[168,1428,390,1784]" targetPageId="9">
<paragraph id="8BED690CAF26FFABFF58F8D8AD4DA048" blockId="9.[151,1436,1816,1900]" pageId="9" pageNumber="310">
<emphasis id="B926B51EAF26FFABFF58F8D8AE0AA00A" bold="true" box="[151,274,1816,1838]" pageId="9" pageNumber="310">FIGURE 5.</emphasis>
<taxonomicName id="4C52128FAF26FFABFED6F8D8AD33A00A" box="[281,555,1816,1838]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="burmana">
<emphasis id="B926B51EAF26FFABFED6F8D8AD33A00A" box="[281,555,1816,1838]" italics="true" pageId="9" pageNumber="310">Rhynchoraphidia burmana</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF26FFABFDFEF8D9ADD6A00A" bold="true" box="[561,718,1817,1838]" pageId="9" pageNumber="310">gen. et sp. nov.</emphasis>
, holotype EMTG BA-002256. A. Habitus in dorsal view; B. Head and thorax in dorsal view; C. distal parts of left wings; D. Drawing of head and prothorax in dorsal view. Arrow indicating pterostigmal crossvein. Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF2D3984AF25FFA8FF58FAA7AC8BA2BA" httpUri="https://zenodo.org/record/264765/files/figure.png" pageId="10" pageNumber="311" targetBox="[315,1270,200,1348]" targetPageId="10">
<paragraph id="8BED690CAF25FFA8FF58FAA7AC8BA2BA" blockId="10.[151,1436,1383,1438]" pageId="10" pageNumber="311">
<emphasis id="B926B51EAF25FFA8FF58FAA7AE0AA258" bold="true" box="[151,274,1383,1405]" pageId="10" pageNumber="311">FIGURE 6.</emphasis>
<taxonomicName id="4C52128FAF25FFA8FED6FAA7AD33A259" box="[281,555,1383,1405]" class="Insecta" family="Mesoraphidiidae" genus="Rhynchoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="burmana">
<emphasis id="B926B51EAF25FFA8FED6FAA7AD33A259" box="[281,555,1383,1405]" italics="true" pageId="10" pageNumber="311">Rhynchoraphidia burmana</emphasis>
</taxonomicName>
<emphasis id="B926B51EAF25FFA8FDFFFAA7ADD5A258" bold="true" box="[560,717,1383,1404]" pageId="10" pageNumber="311">gen. et sp. nov.</emphasis>
A. Holotype EMTG BA-002256, drawing of right fore- and hindwing; B. same specimen, drawing of left fore- and hindwing. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BED690CAF25FFA8FF58FA32AE06A109" blockId="10.[151,538,1521,1581]" pageId="10" pageNumber="311">
<emphasis id="B926B51EAF25FFA8FF58FA32AD02A12F" bold="true" box="[151,538,1521,1548]" pageId="10" pageNumber="311">
Genus
<taxonomicName id="4C52128FAF25FFA8FF21FA31AEB1A12F" box="[238,425,1521,1547]" class="Insecta" family="Mesoraphidiidae" genus="Burmoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B926B51EAF25FFA8FF21FA31AEB1A12F" bold="true" box="[238,425,1521,1547]" italics="true" pageId="10" pageNumber="311">Burmoraphidia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2150865AF25FFA8FE7FFA32AD02A12F" box="[432,538,1522,1547]" pageId="10" pageNumber="311" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13697589AF25FFA8FF50F9D4AE0FA109" box="[159,279,1556,1581]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStartId-0="11.[151,250,1925,1947]" captionStartId-1="12.[151,250,1932,1954]" captionStartId-2="13.[151,250,859,881]" captionStartId-3="16.[151,250,1842,1864]" captionStartId-4="17.[151,250,1498,1520]" captionTargetBox-0="[170,1428,703,1889]" captionTargetBox-1="[335,1252,201,1875]" captionTargetBox-2="[322,1265,197,830]" captionTargetBox-3="[164,1423,210,1805]" captionTargetBox-4="[165,1424,200,1445]" captionTargetId-0="figure@11.[151,1436,693,1904]" captionTargetId-1="figure@12.[331,1262,193,1884]" captionTargetId-2="figure@13.[316,1271,193,837]" captionTargetId-3="figure@16.[151,1435,195,1820]" captionTargetId-4="figure@17.[153,1438,193,1465]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="16" captionTargetPageId-4="17" captionText-0="FIGURE 7. Burmoraphidia reni gen. et sp. nov., female. A. Holotype EMTG BA- 001272, habitus in dorsal view; B. Paratype EMTG BA- 002135, habitus in dorsal view; C. Paratype EMTG BA- 002260, habitus in dorsal view; D. Paratype EMTG BA- 0 0 2254, habitus in dorsal view. Arrows indicating pterostigmal crossvein. Scale bar = 1.0 mm." captionText-1="FIGURE 8. Burmoraphidia reni gen. et sp. nov., male. A. Paratype EMTG BA- 001772, habitus in dorsal view; B. Paratype EMTG BA- 002158, habitus in ventral view; C. Paratype EMTG BA- 002255, habitus in ventral view; D. Paratype EMTG BA- 0 0 2197, habitus in laterocaudal view. Scale bar = 1.0 mm." captionText-2="FIGURE 9. Burmoraphidia reni gen. et sp. nov., holotype EMTG BA- 001272, drawing of fore- and hindwing. Scale bar = 1.0 mm." captionText-3="FIGURE 10. Male genitalia of Burmoraphidia reni gen. et sp. nov. A. Paratype EMTG BA- 001772, photo in lateral view; B. Paratype EMTG BA- 001772, drawing in lateral view; C. Paratype EMTG BA- 002158, photo in ventral view; D. Paratype EMTG BA- 002158, drawing in ventral view; E. Paratype EMTG BA- 002197, photo in caudal view; F. Paratype EMTG BA- 0 0 2197, drawing in caudal view. Arrows indicating distal forking of the central structure. T: tergum; S: sternum; gx: gonocoxite; gp: gonapophysis; e: ectoproct. Scale bar = 0.5 mm." captionText-4="FIGURE 11. Female genitalia of Burmoraphidia reni gen. et sp. nov. A. Paratype EMTG BA- 002260, photo in ventral view; B. Paratype EMTG BA- 002260, drawing in ventral view; C. Paratype EMTG BA- 002135, photo in lateral view; D. Paratype EMTG BA- 002135, drawing in lateral view; E. Paratype EMTG BA- 002254, drawing in lateral view. T: tergum; S: sternum; sg: subgenitale; gst: gonostylus; op: ovipositor; e: ectoproct. Scale bar = 0.5 mm." httpUri-0="https://zenodo.org/record/264766/files/figure.png" httpUri-1="https://zenodo.org/record/264767/files/figure.png" httpUri-2="https://zenodo.org/record/264768/files/figure.png" httpUri-3="https://zenodo.org/record/264769/files/figure.png" httpUri-4="https://zenodo.org/record/264770/files/figure.png" pageId="10" pageNumber="311">Figs. 711</figureCitation>
)
</paragraph>
<paragraph id="8BED690CAF25FFA8FF58F99DAD5DA156" blockId="10.[151,581,1628,1650]" box="[151,581,1628,1650]" pageId="10" pageNumber="311">
<typeStatus id="54E9D7AEAF25FFA8FF58F99DAFD1A156" box="[151,201,1629,1650]" pageId="10" pageNumber="311">Type</typeStatus>
species:
<taxonomicName id="4C52128FAF25FFA8FEE9F99CAEE8A156" box="[294,496,1628,1650]" class="Insecta" family="Mesoraphidiidae" genus="Burmoraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reni" status="sp. nov.">
<emphasis id="B926B51EAF25FFA8FEE9F99CAEE8A156" box="[294,496,1628,1650]" italics="true" pageId="10" pageNumber="311">Burmoraphidia reni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2150865AF25FFA8FE39F99DAD5DA156" box="[502,581,1629,1650]" pageId="10" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF25FFA9FF58F95FADAAA639" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311" type="diagnosis">
<paragraph id="8BED690CAF25FFA9FF58F95FADAAA639" blockId="10.[151,1437,1695,2008]" lastBlockId="11.[151,1437,151,429]" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311">
<emphasis id="B926B51EAF25FFA8FF58F95FAE09A19C" bold="true" box="[151,273,1695,1720]" pageId="10" pageNumber="311">Diagnosis.</emphasis>
Minute size. Head ovoid, with portion posterior to compound eyes shorter than eye length and tapering caudad, and with clypeus slightly shorter than compound eye length; compound eyes large, ovoid; three small ocelli present on posterior half of head; antenna much longer than head plus prothorax, but shorter than entire body length, with ~40 flagellomeres. Pronotum slightly shorter than head, with a constant height along entire length. Pro- and mesotibiae swollen, but metatibia slender, tibial spurs present. Forewing: Costal space distinctly broadened (at widest point costal space ~1.5× width of pterostigma); pterostigma elongate, ~1/3× wing length, proximally diffused but not closed by a crossvein, starting proximad termination of Scp, with an oblique pterostigmal crossvein slightly distad midpoint of pterostigma, distally closed by a veinlet of RA; ScP terminating into C slightly distad wing midpoint; posterior veinlet of RA prolonged and zig-zagged with stem of RA;
<collectingCountry id="F345299CAF25FFA8FAB5F800AA84A0F3" box="[1402,1436,1984,2007]" name="Poland" pageId="10" pageNumber="311">RP</collectingCountry>
simple; two radial cells, one discal cell, and two discoidal cells present. Hindwing: Configuration of pterostigmal area same to that of male; stem of MA proximally fused with RA; an oblique proximal mp-cua present; two radial cells, one discal cell, and one discoidal cell present; A1 distally bifurcated and medially fused with A2 for a short distance, proximally forming an ovoid anal cell.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF24FFA9FF08FEE7AB9CA641" pageId="11" pageNumber="312" type="etymology">
<paragraph id="8BED690CAF24FFA9FF08FEE7AB9CA641" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="312">
<emphasis id="B926B51EAF24FFA9FF08FEE7AE56A664" bold="true" box="[199,334,295,320]" pageId="11" pageNumber="312">Etymology.</emphasis>
From
<collectingCountry id="F345299CAF24FFA9FE52FEE8AEF1A61B" box="[413,489,296,319]" name="Myanmar" pageId="11" pageNumber="312">Burma</collectingCountry>
and
<taxonomicName id="4C52128FAF24FFA9FDEAFEE8AD94A664" box="[549,652,296,320]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="B926B51EAF24FFA9FDEAFEE8AD94A664" box="[549,652,296,320]" italics="true" pageId="11" pageNumber="312">Raphidia</emphasis>
</taxonomicName>
, a common genus-group name of
<taxonomicName id="4C52128FAF24FFA9FBD5FEE7ABA3A664" box="[1050,1211,295,320]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="order">Raphidioptera</taxonomicName>
, in reference to the occurrence of the new genus from the mid Cretaceous Burmese amber. Gender: feminine.
</paragraph>
</subSubSection>
<subSubSection id="C3483A87AF24FFA9FF08FEAFAEC0A689" pageId="11" pageNumber="312" type="discussion">
<paragraph id="8BED690CAF24FFA9FF08FEAFAEC0A689" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="312">
<emphasis id="B926B51EAF24FFA9FF08FEAFAE23A6AC" bold="true" box="[199,315,367,392]" pageId="11" pageNumber="312">Remarks.</emphasis>
The new genus differs from all other nanoraphidiine genera by the pterostigma with a pterostigmal crossvein and a diffuse base.
</paragraph>
</subSubSection>
</treatment>
</document>