621 lines
134 KiB
XML
621 lines
134 KiB
XML
<document id="550F289C0A7D37919C28D6D09182D6C1" ID-CLB-Dataset="36382" ID-DOI="10.11646/zootaxa.3828.1.1" ID-GBIF-Dataset="8c1cda9b-0d82-41f3-a1c1-d8eb574a1ba7" ID-ISSN="1175-5326" ID-Zenodo-Dep="286513" ID-ZooBank="B2FEAE60-7E51-45FA-A38F-930A084A5AAA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460452080076" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="03F68791FFF218228FFE3F4301B4FE63" docLanguage="en" docName="zt03828p076.pdf" docOrigin="Zootaxa 3828 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Spinosciacharis Jałoszyński, 2014, gen. n." docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="FFCFFFE9FFD718088F693A1C034CFC3C" masterDocTitle="' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="76" masterPageNumber="1" pageNumber="38" updateTime="1698338417320" updateUser="ExternalLinkService">
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<mods:titleInfo id="50DF70CA9CB7403919DFA31F7B61439F">
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<mods:title id="70B60E5802D6E7513646D6509624E485">' The curse of Horaeomorphus ': taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="EFA552475B89CC2731DB6414E10297FF" type="personal">
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<mods:roleTerm id="FFEB1F5999D87D5C7B79F99A778BB201">Author</mods:roleTerm>
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<mods:namePart id="8A3C22C26E190CDABBADB18FC107E78F">Jałoszyński, Paweł</mods:namePart>
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<mods:typeOfResource id="CCE2CC162EE8394822990FC3DCC66FAB">text</mods:typeOfResource>
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<mods:title id="779144999D1655B2BAA1F73EE3B22941">Zootaxa</mods:title>
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<mods:part id="4D64FED82D6D50EA38E8836E969F7DD9">
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<mods:date id="2C653450FD311092EF27B46F14C3F0CB">2014</mods:date>
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<mods:detail id="48023E33F3964C352C0FC9E9419E32E2" type="volume">
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<mods:number id="280ECB2103ECBC793E65BF7A66A2CA25">3828</mods:number>
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<mods:classification id="7E04B21119F3367B7A60D6390E3020C9">journal article</mods:classification>
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<mods:identifier id="3BB0B49FC0D066E5D5B9A4843F529317" type="CLB-Dataset">36382</mods:identifier>
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<mods:identifier id="A7DE72108678AD11D14C3E5B550591A8" type="DOI">10.11646/zootaxa.3828.1.1</mods:identifier>
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<treatment id="03F68791FFF218228FFE3F4301B4FE63" ID-DOI="http://doi.org/10.5281/zenodo.6141076" ID-GBIF-Taxon="127711757" ID-Zenodo-Dep="6141076" LSID="urn:lsid:plazi:treatment:03F68791FFF218228FFE3F4301B4FE63" httpUri="http://treatment.plazi.org/id/03F68791FFF218228FFE3F4301B4FE63" lastPageId="42" lastPageNumber="41" pageId="37" pageNumber="38">
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<subSubSection id="C345650CFFF2182D8FFE3F4302FCF946" box="[151,432,1375,1402]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="8BE03687FFF2182D8FFE3F4302FCF946" blockId="37.[151,432,1375,1402]" box="[151,432,1375,1402]" pageId="37" pageNumber="38">
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<heading id="D0A881EBFFF2182D8FFE3F4302FCF946" bold="true" box="[151,432,1375,1402]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
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<emphasis id="B92BEA95FFF2182D8FFE3F4302FCF946" bold="true" box="[151,432,1375,1402]" pageId="37" pageNumber="38">
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<taxonomicName id="4C5F4D04FFF2182D8FFE3F43021BF945" ID-CoL="8XPYR" box="[151,343,1375,1401]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B92BEA95FFF2182D8FFE3F43021BF945" bold="true" box="[151,343,1375,1401]" italics="true" pageId="37" pageNumber="38">Spinosciacharis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21857EEFFF2182D8E373F7C02FCF946" box="[350,432,1376,1402]" pageId="37" pageNumber="38" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
|
||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C345650CFFF2182D8FFE3FBB00E1F980" box="[151,941,1446,1469]" pageId="37" pageNumber="38" type="materials_examined">
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<paragraph id="8BE03687FFF2182D8FFE3FBB00E1F980" blockId="37.[151,941,1446,1469]" box="[151,941,1446,1469]" pageId="37" pageNumber="38">
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<typeStatus id="54E48825FFF2182D8FFE3FBB0385F980" box="[151,201,1447,1468]" pageId="37" pageNumber="38">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="4C5F4D04FFF2182D8E4F3FBA01BDF980" authority="Franz, 1975: 142" authorityName="Franz" authorityPageNumber="142" authorityYear="1975" box="[294,753,1446,1469]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="puncticeps">
|
||
<emphasis id="B92BEA95FFF2182D8E4F3FBA0172F981" box="[294,574,1446,1469]" italics="true" pageId="37" pageNumber="38">Horaeomorphus puncticeps</emphasis>
|
||
<bibRefCitation id="EFCE4B76FFF2182D8D2D3FBB018DF980" author="Franz" box="[580,705,1446,1468]" pageId="37" pageNumber="38" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312." type="journal article" year="1975">Franz, 1975</bibRefCitation>
|
||
: 142
|
||
</taxonomicName>
|
||
(here designated).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFF2182E8FFE3FF50060FDC8" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="diagnosis">
|
||
<paragraph id="8BE03687FFF2182E8FFE3FF50060FDC8" blockId="37.[151,1437,1513,2006]" lastBlockId="38.[151,1437,151,1977]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
|
||
<emphasis id="B92BEA95FFF2182D8FFE3FF5025DFA3E" bold="true" box="[151,273,1513,1538]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
|
||
Body (
|
||
<figureCitation id="13642A02FFF2182D8E183FF50119FA3E" box="[369,597,1513,1538]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="37" pageNumber="38">Figs. 112, 114–116</figureCitation>
|
||
) elongate and flattened; head short and convex; vertex not projecting dorsocaudad, its posterior margin slightly to distinctly concave; supraantennal tubercles distinct but weakly raised and not accompanied posteriorly by pits; frontoclypeal groove present; eyes located in anterior part of head; tempora long and strongly curved toward occipital constriction; tempora, genae and postgenae with dense bristles; 'neck region' distinctly demarcated from anterior part of head capsule by constriction and as narrow as nearly half width of vertex; hypostomal ridges (
|
||
<figureCitation id="13642A02FFF2182D8D453C810133FA8A" box="[556,639,1693,1718]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="37" pageNumber="38">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8DE23C8101EFFA8A" box="[651,675,1693,1718]" italics="true" pageId="37" pageNumber="38">hr</emphasis>
|
||
) sharply marked and long, reaching nearly to posterior tentorial pits but not connected in middle; posterior tentorial pits (
|
||
<figureCitation id="13642A02FFF2182D8C643CDD002AFAE6" box="[781,870,1729,1754]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="37" pageNumber="38">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8C1F3CDE00D6FAE5" box="[886,922,1730,1753]" italics="true" pageId="37" pageNumber="38">ptp</emphasis>
|
||
) elongate and located distinctly in front of transverse groove demarcating 'neck' ventrally; submentum (
|
||
<figureCitation id="13642A02FFF2182D8C233CF900D3FAC2" box="[842,927,1765,1790]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="37" pageNumber="38">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8CC53CFA0095FAC1" box="[940,985,1766,1789]" italics="true" pageId="37" pageNumber="38">smn</emphasis>
|
||
) without lateral sutures; each mandible with robust subapical tooth (
|
||
<figureCitation id="13642A02FFF2182D8E8E3D150100FB1E" box="[487,588,1801,1826]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="37" pageNumber="38">Fig. 100</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8D353D160130FB1D" box="[604,636,1802,1825]" italics="true" pageId="37" pageNumber="38">sat</emphasis>
|
||
) and densely setose dorsomesal prostheca (
|
||
<figureCitation id="13642A02FFF2182D8B173D1507A8FB1E" box="[1150,1252,1801,1826]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="37" pageNumber="38">Fig. 100</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8B9D3D160659FB1D" box="[1268,1301,1802,1825]" italics="true" pageId="37" pageNumber="38">pst</emphasis>
|
||
); maxillary palpomeres III and IV (
|
||
<figureCitation id="13642A02FFF2182D8EC43D31015EFB7A" box="[429,530,1837,1862]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="37" pageNumber="38">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8D483D32013FFB7A" box="[545,627,1838,1862]" italics="true" pageId="37" pageNumber="38">mxt3–4</emphasis>
|
||
) slender, strongly elongate; pronotum with one pair of small internal antebasal pits (
|
||
<figureCitation id="13642A02FFF2182D8E293D4D02EEFB56" box="[320,418,1873,1898]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="37" pageNumber="38">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8EC63D4E0289FB55" box="[431,453,1874,1897]" italics="true" pageId="37" pageNumber="38">ip</emphasis>
|
||
) which are separated by narrow longitudinal wrinkle or carina (
|
||
<figureCitation id="13642A02FFF2182D8BF33D4D07B0FB56" box="[1178,1276,1873,1898]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="37" pageNumber="38">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8A603D4E0666FB55" box="[1289,1322,1874,1897]" italics="true" pageId="37" pageNumber="38">mc</emphasis>
|
||
), external pair of diffused impressions (
|
||
<figureCitation id="13642A02FFF2182D8E8E3D690106FBB2" box="[487,586,1909,1934]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="37" pageNumber="38">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8D313D690124FBB2" box="[600,616,1909,1934]" italics="true" pageId="37" pageNumber="38">li</emphasis>
|
||
) and distinct sublateral carinae (
|
||
<figureCitation id="13642A02FFF2182D8CB63D69070EFBB2" box="[991,1090,1909,1934]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="37" pageNumber="38">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8B393D690723FBB2" box="[1104,1135,1909,1934]" italics="true" pageId="37" pageNumber="38">slc</emphasis>
|
||
) extending from posterior pronotal corners to 1/5–1/4 PL; prosternum laterally demarcated from hypomera by pronotosternal sutures (
|
||
<figureCitation id="13642A02FFF2182D8A063D850388FBEA" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="37" pageNumber="38">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8FBD3DA203B4FBE9" box="[212,248,1982,2005]" italics="true" pageId="37" pageNumber="38">nss</emphasis>
|
||
), with indistinct intercoxal carina; procoxal sockets (
|
||
<figureCitation id="13642A02FFF2182D8C073DA10098FBEA" box="[878,980,1981,2006]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="37" pageNumber="38">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF2182D8C8D3DA20745FBE9" box="[996,1033,1982,2005]" italics="true" pageId="37" pageNumber="38">pcs</emphasis>
|
||
) closed by posterolateral lobes of prosternum; prothoracic hypomera (
|
||
<figureCitation id="13642A02FFF1182E8D513A8B01D1FC8C" box="[568,669,151,176]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8DC43A8B0184FC8C" box="[685,712,151,176]" italics="true" pageId="38" pageNumber="39">hy</emphasis>
|
||
) with large internal (adcoxal) parts demarcated from sides of pronotum by complete hypomeral ridges (
|
||
<figureCitation id="13642A02FFF1182E8D023AA00187FCE8" box="[619,715,188,213]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8DBF3AA001B7FCE9" box="[726,763,188,213]" italics="true" pageId="38" pageNumber="39">hyr</emphasis>
|
||
); sides of pronotum with dense thick bristles (
|
||
<figureCitation id="13642A02FFF1182E8B963AA00611FCE8" box="[1279,1373,188,213]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
) well visible in dorsal view; mesoventrite with short anterior ridge (
|
||
<figureCitation id="13642A02FFF1182E8C333AC300F0FCC4" box="[858,956,223,248]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="38" pageNumber="39">Fig. 105</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8CA33AFD00AFFCC4" box="[970,995,225,248]" italics="true" pageId="38" pageNumber="39">ar</emphasis>
|
||
) medially connected with mesoventral process (
|
||
<figureCitation id="13642A02FFF1182E8E683B180225FD21" box="[257,361,260,285]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="38" pageNumber="39">Fig. 105</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E123B1902F9FD20" box="[379,437,261,284]" italics="true" pageId="38" pageNumber="39">msvp</emphasis>
|
||
) which is narrow and weakly expanding ventrally and reaching only to middle of mesocoxal cavities, so that mesocoxae are indistinctly separated; ventrolateral foveae (
|
||
<figureCitation id="13642A02FFF1182E8BF23B3B064DFD7C" box="[1179,1281,295,320]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="38" pageNumber="39">Fig. 110</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8A7B3B3B0662FD7C" box="[1298,1326,295,320]" italics="true" pageId="38" pageNumber="39">vlf</emphasis>
|
||
) present, dorsolateral foveae absent; anterior metaventral process (
|
||
<figureCitation id="13642A02FFF1182E8C483B5000C8FD59" box="[801,900,332,357]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="38" pageNumber="39">Fig. 106</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8CFB3B510082FD58" box="[914,974,333,356]" italics="true" pageId="38" pageNumber="39">amvp</emphasis>
|
||
) present; metaventral intercoxal process (
|
||
<figureCitation id="13642A02FFF1182E8FF63B7303B3FDB4" box="[159,255,367,392]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="38" pageNumber="39">Fig. 111</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E653B6D020EFDB4" box="[268,322,369,392]" italics="true" pageId="38" pageNumber="39">mtvp</emphasis>
|
||
) broadly subtriangular, with small median notch; metacoxae (
|
||
<figureCitation id="13642A02FFF1182E8B683B73072FFDB4" box="[1025,1123,367,392]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="38" pageNumber="39">Fig. 109</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8B063B6C07DAFDB4" box="[1135,1174,368,392]" italics="true" pageId="38" pageNumber="39">cx3</emphasis>
|
||
) narrowly separated in middle; each elytron with two deep and relatively large asetose basal foveae (
|
||
<figureCitation id="13642A02FFF1182E8B433B8807C3FD90" box="[1066,1167,404,429]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 104</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8BF73B88078CFD91" box="[1182,1216,404,429]" italics="true" pageId="38" pageNumber="39">bef</emphasis>
|
||
) not connected by groove and well visible in dry-mounted specimens; metafemora in males with ventral projection (
|
||
<figureCitation id="13642A02FFF1182E8B833BAB06C3FDEC" box="[1258,1423,439,464]" captionStart="FIGURES 117 – 120" captionStartId="45.[151,264,696,719]" captionTargetBox="[161,1431,193,663]" captionTargetId="figure@45.[151,1436,193,675]" captionTargetPageId="45" captionText="FIGURES 117 – 120. Spinosciacharis spp. distal part of left metafemur in posterior view. Spinosciacharis puncticeps (Franz) comb. n. (117); Spinosciacharis sassy sp. n. (118), Spinosciacharis clydensis sp. n. (119), and Spinosciacharis acheron sp. n. (120)." httpUri="https://zenodo.org/record/286538/files/figure.png" pageId="38" pageNumber="39">Figs. 117–120</figureCitation>
|
||
); aedeagus (
|
||
<figureCitation id="13642A02FFF1182E8E643BC002FCFDC8" box="[269,432,476,501]" captionStart="FIGURES 121 – 128" captionStartId="46.[151,264,1877,1900]" captionTargetBox="[168,1425,220,1825]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURES 121 – 128. Aedeagi of Spinosciacharis spp. in ventral (121, 123, 125, 127) and lateral (122, 124, 126, 128) views. Spinosciacharis puncticeps (Franz) comb. n. (121 – 122); Spinosciacharis sassy sp. n. (123 – 124), Spinosciacharis clydensis sp. n. (125 – 126), and Spinosciacharis acheron sp. n. (127 – 128)." httpUri="https://zenodo.org/record/286539/files/figure.png" pageId="38" pageNumber="39">Figs. 121–128</figureCitation>
|
||
) with free and slender parameres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFF118208FAE3BE302FBFE9B" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="description">
|
||
<paragraph id="8BE03687FFF1182E8FAE3BE301FAFE00" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
<emphasis id="B92BEA95FFF1182E8FAE3BE30215FE24" bold="true" box="[199,345,511,536]" pageId="38" pageNumber="39">Description.</emphasis>
|
||
Body (
|
||
<figureCitation id="13642A02FFF1182E8EC33BE301CCFE24" box="[426,640,511,536]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="38" pageNumber="39">Figs. 112, 114–116</figureCitation>
|
||
) moderately large (BL
|
||
<quantity id="4CA79B62FFF1182E8CEC3BE30767FE24" box="[901,1067,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="2.115" metricValueMax="2.45" metricValueMin="1.78" pageId="38" pageNumber="39" unit="mm" value="2.115" valueMax="2.45" valueMin="1.78">1.78–2.45 mm</quantity>
|
||
), flattened and slender, with long appendages, pigmentation brown, cuticle setose.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE385B067CFF04" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Head (
|
||
<figureCitation id="13642A02FFF1182E8E7E385B0236FE5C" box="[279,378,583,608]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Figs. 97</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFF1182E8EE238540165FE5C" box="[395,553,583,608]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="38" pageNumber="39">112, 114–116</figureCitation>
|
||
) with occipital constriction (
|
||
<figureCitation id="13642A02FFF1182E8CEB385B0096FE5C" box="[898,986,583,608]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8C823855075DFE5C" box="[1003,1041,585,608]" italics="true" pageId="38" pageNumber="39">occ</emphasis>
|
||
) much narrower than vertex and dividing the head capsule into exposed anterior part and narrow posterior 'neck region' retracted into prothorax; eyes small and slightly oval, located in anterior part of head; tempora long and in posterior half strongly curved posteromesally, with sparse bristles; vertex transverse, uniformly convex, not projecting dorso-caudad, with posterior margin slightly concave and distinctly demarcated from occiput; frons confluent with vertex, transverse and subtrapezoidal, convex, strongly but not abruptly lowering toward labrum; frontoclypeal groove present; antennal insertions broadly separated, located beneath distinct but weakly raised supraantennal tubercles.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE395807AEF931" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Labrum (
|
||
<figureCitation id="13642A02FFF1182E8E46395802CCFF60" box="[303,384,836,861]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 99</figureCitation>
|
||
) transverse and semioval with slightly emarginate anterior margin, with small number of long dorsal setae. Mandibles symmetrical, subtriangular, each with broad base and large subapical tooth (
|
||
<figureCitation id="13642A02FFF1182E8B94397B0613FFBC" box="[1277,1375,871,896]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 100</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8A02397506C0FFBC" box="[1387,1420,873,896]" italics="true" pageId="38" pageNumber="39">sat</emphasis>
|
||
), also distal mesal margin of basal part forms broad and blunt tooth in front of densely setose dorsomesal prostheca reaching mandibular base. Each maxilla composed of small cardo (
|
||
<figureCitation id="13642A02FFF1182E8CE539B300A2FFF4" box="[908,1006,943,968]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8C9339B30759FFF4" box="[1018,1045,943,968]" italics="true" pageId="38" pageNumber="39">cd</emphasis>
|
||
) bearing three setae; subtriangular basistipes (
|
||
<figureCitation id="13642A02FFF1182E8E7A39C80238FFD0" box="[275,372,980,1005]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E1639C802D3FFD1" box="[383,415,980,1005]" italics="true" pageId="38" pageNumber="39">bst</emphasis>
|
||
); elongate mediostipes (
|
||
<figureCitation id="13642A02FFF1182E8DDB39C8005FFFD0" box="[690,787,980,1005]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8C7739C90009FFD0" box="[798,837,981,1004]" italics="true" pageId="38" pageNumber="39">mst</emphasis>
|
||
); elongate galea (
|
||
<figureCitation id="13642A02FFF1182E8B6439C80722FFD0" box="[1037,1134,980,1005]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8B1039C807D0FFD1" box="[1145,1180,980,1005]" italics="true" pageId="38" pageNumber="39">gal</emphasis>
|
||
) and lacinia (
|
||
<figureCitation id="13642A02FFF1182E8A5F39C806DBFFD0" box="[1334,1431,980,1005]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8FFE39EB03F5F82C" box="[151,185,1015,1040]" italics="true" pageId="38" pageNumber="39">lac</emphasis>
|
||
); large, elongate palpifer (
|
||
<figureCitation id="13642A02FFF1182E8E8D39EB0109F82C" box="[484,581,1015,1040]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8D3839E40139F82C" box="[593,629,1016,1040]" italics="true" pageId="38" pageNumber="39">pfp</emphasis>
|
||
); and strongly elongate and moderately large maxillary palpus (
|
||
<figureCitation id="13642A02FFF1182E8A2D39EB06DBF82C" box="[1348,1431,1015,1040]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8FFE3E01038AF808" box="[151,198,1053,1076]" italics="true" pageId="38" pageNumber="39">mxp</emphasis>
|
||
) composed of elongate palpomere I (
|
||
<figureCitation id="13642A02FFF1182E8DF03E00004EF808" box="[665,770,1052,1077]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8C7F3E00001FF808" box="[790,851,1052,1076]" italics="true" pageId="38" pageNumber="39">mxp1</emphasis>
|
||
), strongly elongate, slightly clavate and slender palpomere II (
|
||
<figureCitation id="13642A02FFF1182E8E503E2302D0F864" box="[313,412,1087,1112]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8EC03E5C02A9F864" box="[425,485,1088,1112]" italics="true" pageId="38" pageNumber="39">mxp2</emphasis>
|
||
), large and strongly elongate, slender palpomere III (
|
||
<figureCitation id="13642A02FFF1182E8B2F3E2307E5F864" box="[1094,1193,1087,1112]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8BDF3E5C07BEF864" box="[1206,1266,1088,1112]" italics="true" pageId="38" pageNumber="39">mxp3</emphasis>
|
||
) broadest near distal third, and small, slender, subconical and pointed palpomere IV (
|
||
<figureCitation id="13642A02FFF1182E8CC23E780741F840" box="[939,1037,1124,1149]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 101</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8B703E78071AF840" box="[1049,1110,1124,1148]" italics="true" pageId="38" pageNumber="39">mxp4</emphasis>
|
||
). Labium (
|
||
<figureCitation id="13642A02FFF1182E8BBD3E780617F840" box="[1236,1371,1124,1149]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Figs. 97–98</figureCitation>
|
||
) with large submentum (
|
||
<figureCitation id="13642A02FFF1182E8E053E9B028EF89C" box="[364,450,1159,1184]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8EB93E9502B1F89C" box="[464,509,1161,1184]" italics="true" pageId="38" pageNumber="39">smn</emphasis>
|
||
) not demarcated laterally from postcardinal parts of hypostomae; subtrapezoidal mentum (
|
||
<figureCitation id="13642A02FFF1182E8E6B3EB00218F8F8" box="[258,340,1196,1221]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E093EB102CEF8F8" box="[352,386,1197,1220]" italics="true" pageId="38" pageNumber="39">mn</emphasis>
|
||
); and short prementum bearing narrowly separated at bases long 3-segmented labial palpi and a pair of median bristles inserted on small subtrapezoidal ligula (
|
||
<figureCitation id="13642A02FFF1182E8CF83ED300A4F8D4" box="[913,1000,1231,1256]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 98</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8C9E3ED30741F8D4" box="[1015,1037,1231,1256]" italics="true" pageId="38" pageNumber="39">lg</emphasis>
|
||
). Hypostomal ridges (
|
||
<figureCitation id="13642A02FFF1182E8A7A3ED30626F8D4" box="[1299,1386,1231,1256]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8A133ED306DFF8D4" box="[1402,1427,1231,1256]" italics="true" pageId="38" pageNumber="39">hr</emphasis>
|
||
) sharply marked, long and extending nearly to posterior tentorial pits but not connecting in middle.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE3F0B0131F944" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Gular plate (
|
||
<figureCitation id="13642A02FFF1182E8E3C3F0B02E5F90C" box="[341,425,1303,1328]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8EDF3F05029EF90C" box="[438,466,1305,1328]" italics="true" pageId="38" pageNumber="39">gp</emphasis>
|
||
) large and subtrapezoidal, with rapidly narrowed anterior part; gular sutures (
|
||
<figureCitation id="13642A02FFF1182E8A2A3F0B06DBF90C" box="[1347,1431,1303,1328]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8FFE3F2103FCF968" box="[151,176,1341,1364]" italics="true" pageId="38" pageNumber="39">gs</emphasis>
|
||
) superficial; posterior tentorial pits (
|
||
<figureCitation id="13642A02FFF1182E8D093F2001FBF968" box="[608,695,1340,1365]" captionStart="FIGURES 97 – 101" captionStartId="39.[151,264,1576,1599]" captionTargetBox="[166,1429,193,1556]" captionTargetId="figure@39.[166,1431,193,1556]" captionTargetPageId="39" captionText="FIGURES 97 – 101. Morphology of Spinosciacharis acheron sp. n. Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp 1 – 3, labial palpomere 1 – 3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp 1 – 4, maxillary palpomere 1 – 4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum." httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="38" pageNumber="39">Fig. 97</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8DAE3F2101A7F968" box="[711,747,1341,1364]" italics="true" pageId="38" pageNumber="39">ptp</emphasis>
|
||
) narrow and straight, located clearly in front of transverse groove ventrally demarcating 'neck region'.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE3F980211F9FC" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Antennae (
|
||
<figureCitation id="13642A02FFF1182E8E2A3F980151F9A1" box="[323,541,1412,1437]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="38" pageNumber="39">Figs. 112, 114–116</figureCitation>
|
||
) long and slender, gradually but only slightly thickening distally; antennomere XI unremarkable.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE3FD0007CFA80" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Prothorax (
|
||
<figureCitation id="13642A02FFF1182E8E213FD002FBF9D8" box="[328,439,1484,1509]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Figs. 102</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFF1182E8EAF3FD0012EF9D9" box="[454,610,1484,1509]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="38" pageNumber="39">112, 114–116</figureCitation>
|
||
) in dorsal view weakly to strongly elongate and broadest near anterior third, with rounded anterior margin and anterior parts of lateral margins, with barely marked, blunt and strongly obtuse anterior corners; posterior corners distinct, obtuse; posterior margin shallowly bisinuate. Base of pronotum with distinct sublateral carinae (
|
||
<figureCitation id="13642A02FFF1182E8D693C2B012FFA6C" box="[512,611,1591,1616]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8D063C2B01C2FA6C" box="[623,654,1591,1616]" italics="true" pageId="38" pageNumber="39">slc</emphasis>
|
||
) extending from posterior pronotal corners and as long as 1/5–1/4 of PL; base with one pair of small internal pits (
|
||
<figureCitation id="13642A02FFF1182E8DF43C40004CFA48" box="[669,768,1628,1653]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8C643C41006FFA48" box="[781,803,1629,1652]" italics="true" pageId="38" pageNumber="39">ip</emphasis>
|
||
) in middle separated by narrow longitudinal wrinkle or carina (
|
||
<figureCitation id="13642A02FFF1182E8F843C63021CFAA4" box="[237,336,1663,1688]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E343C9D0232FAA4" box="[349,382,1665,1688]" italics="true" pageId="38" pageNumber="39">mc</emphasis>
|
||
), and with variously distinct lateral impressions (
|
||
<figureCitation id="13642A02FFF1182E8CDE3C630756FAA4" box="[951,1050,1663,1688]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 103</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8B4E3C63077AFAA4" box="[1063,1078,1663,1688]" italics="true" pageId="38" pageNumber="39">li</emphasis>
|
||
) adjacent to sublateral carinae. Sides of pronotum with thick and sparse bristles (
|
||
<figureCitation id="13642A02FFF1182E8DA83CB8006FFA80" box="[705,803,1700,1725]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 103</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE3CD401B0FB4C" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Prosternum with long basisternal part (
|
||
<figureCitation id="13642A02FFF1182E8DED3CDB01ABFADC" box="[644,743,1735,1760]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8D9C3CDB0059FADC" box="[757,789,1735,1760]" italics="true" pageId="38" pageNumber="39">bst</emphasis>
|
||
) distinctly demarcated from procoxal cavities (
|
||
<figureCitation id="13642A02FFF1182E8A5D3CDB06DBFADC" box="[1332,1431,1735,1760]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8FFE3CF103F2FB38" box="[151,190,1773,1796]" italics="true" pageId="38" pageNumber="39">pcc</emphasis>
|
||
); median part of sternum slightly raised and forming narrow and indistinct prosternal carina; procoxal sockets (
|
||
<figureCitation id="13642A02FFF1182E8FF63D13024EFB14" box="[159,258,1807,1832]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E663D0D0278FB14" box="[271,308,1809,1832]" italics="true" pageId="38" pageNumber="39">pcs</emphasis>
|
||
) closed by posterolateral lobes of prosternum; hypomera (
|
||
<figureCitation id="13642A02FFF1182E8CA73D13077DFB14" box="[974,1073,1807,1832]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8B573D130714FB14" box="[1086,1112,1807,1832]" italics="true" pageId="38" pageNumber="39">hy</emphasis>
|
||
) elongate, each divided into broad lateral part confluent with pronotum and narrower internal (adcoxal) part; hypomeral ridges (
|
||
<figureCitation id="13642A02FFF1182E8B973D28062DFB70" box="[1278,1377,1844,1869]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8A043D2806DEFB71" box="[1389,1426,1844,1869]" italics="true" pageId="38" pageNumber="39">hyr</emphasis>
|
||
) complete; pronotosternal sutures (
|
||
<figureCitation id="13642A02FFF1182E8D7D3D4B0139FB4C" box="[532,629,1879,1904]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 102</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8DE83D4501E9FB4C" box="[641,677,1881,1904]" italics="true" pageId="38" pageNumber="39">nss</emphasis>
|
||
) entire.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFF1182E8FAE3D60075EFB84" blockId="38.[151,1437,151,1977]" pageId="38" pageNumber="39">
|
||
Mesocutellum (
|
||
<figureCitation id="13642A02FFF1182E8E153D6002AEFBA8" box="[380,482,1916,1941]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 104</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8E9D3D60016DFBA9" box="[500,545,1916,1941]" italics="true" pageId="38" pageNumber="39">scl2</emphasis>
|
||
) subtriangular, in intact specimen hidden by posterior margin of pronotum overlapping with elytral base; mesoscutoscutellar suture (
|
||
<figureCitation id="13642A02FFF1182E8C723DBC0030FB84" box="[795,892,1952,1977]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="38" pageNumber="39">Fig. 104</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF1182E8CE13DBD00E5FB84" box="[904,937,1953,1976]" italics="true" pageId="38" pageNumber="39">sss</emphasis>
|
||
) present.
|
||
</paragraph>
|
||
<caption id="DF20660FFFF0182F8FFE3C3401D2FADB" httpUri="https://zenodo.org/record/286533/files/figure.png" pageId="39" pageNumber="40" targetBox="[166,1429,193,1556]" targetPageId="39">
|
||
<paragraph id="8BE03687FFF0182F8FFE3C3401D2FADB" blockId="39.[151,1437,1576,1767]" pageId="39" pageNumber="40">
|
||
<emphasis id="B92BEA95FFF0182F8FFE3C34022DFA01" bold="true" box="[151,353,1576,1599]" pageId="39" pageNumber="40">FIGURES 97–101.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C5F4D04FFF0182F8D633C340049FA02" box="[522,773,1576,1598]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="acheron" status="sp. nov.">
|
||
<emphasis id="B92BEA95FFF0182F8D633C340049FA02" box="[522,773,1576,1598]" italics="true" pageId="39" pageNumber="40">Spinosciacharis acheron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92BEA95FFF0182F8C643C35000BFA02" bold="true" box="[781,839,1577,1598]" pageId="39" pageNumber="40">
|
||
<taxonomicNameLabel id="A21857EEFFF0182F8C643C35000BFA02" box="[781,839,1577,1598]" pageId="39" pageNumber="40" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Head in ventral view (97), distal part of labium in ventral view (98), labrum in dorsal view (99), left mandible in dorsal view (100), and left maxilla in ventral view (101). Abbreviations: bst, basistipes; cd, cardo; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; lac, lacinia; lg, ligula; lhph, lateral lobe of hypopharynx; lp1–3, labial palpomere 1–3; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palpus; mxp1–4, maxillary palpomere 1–4; occ, occipital constriction; ppf, palpifer; pst, prostheca; ptp, posterior tentorial pit; sat, subapical tooth; smn, submentum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE03687FFF018208FAE3D0D0152FCE9" blockId="39.[151,1437,1809,2014]" lastBlockId="40.[151,1437,151,680]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
|
||
Mesoventrite (
|
||
<figureCitation id="13642A02FFF0182F8E063D0D029FFB16" box="[367,467,1809,1834]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="39" pageNumber="40">Fig. 105</figureCitation>
|
||
) with narrow and distinctly demarcated anterior ridge (
|
||
<figureCitation id="13642A02FFF0182F8B0F3D0D0780FB16" box="[1126,1228,1809,1834]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="39" pageNumber="40">Fig. 105</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF0182F8BB43D0F07BAFB16" box="[1245,1270,1811,1834]" italics="true" pageId="39" pageNumber="40">ar</emphasis>
|
||
); mesoventral process (
|
||
<figureCitation id="13642A02FFF0182F8F933D290217FB72" box="[250,347,1845,1870]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="39" pageNumber="40">Fig. 105</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF0182F8E0E3D2B02EDFB72" box="[359,417,1847,1870]" italics="true" pageId="39" pageNumber="40">msvp</emphasis>
|
||
) carinate and moderately expanding ventrally, anteriorly connected with anterior ridge and terminated near middle of mesocoxal cavities, indistinctly separating mesocoxae; sides of mesoventrite with shallow procoxal rests (
|
||
<figureCitation id="13642A02FFF0182F8ECC3D610144FBAA" box="[421,520,1917,1942]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="39" pageNumber="40">Fig. 105</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF0182F8D7C3D630176FBAA" box="[533,570,1919,1942]" italics="true" pageId="39" pageNumber="40">pcr</emphasis>
|
||
) filled with bristles (= setose impressions) but without asetose impressions; mesanepisternum with moderately long prepectus (
|
||
<figureCitation id="13642A02FFF0182F8DBB3DBD0078FB86" box="[722,820,1953,1978]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="39" pageNumber="40">Fig. 105</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF0182F8C563DBF002FFB86" box="[831,867,1955,1978]" italics="true" pageId="39" pageNumber="40">pre</emphasis>
|
||
), sides of mesoventrite with moderately broad and deep setose ventrolateral foveae (
|
||
<figureCitation id="13642A02FFF0182F8D653DD9013BFBE2" box="[524,631,1989,2014]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="39" pageNumber="40">Figs. 108</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFF0182F8DEA3DDA01E3FBE2" box="[643,687,1990,2014]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="39" pageNumber="40">110</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFF0182F8DD23DD9019BFBE2" box="[699,727,1989,2014]" italics="true" pageId="39" pageNumber="40">vlf</emphasis>
|
||
); dorsolateral foveae absent; mesocoxal projections (
|
||
<figureCitation id="13642A02FFF018208A453DD9038FFC8C" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">Figs. 105, 107</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFFF18208FB83A84027EFC8C" box="[209,306,152,176]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="40" pageNumber="41">109–110</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFFF18208E253A850237FC8C" box="[332,379,153,176]" italics="true" pageId="40" pageNumber="41">mcp</emphasis>
|
||
) prominent, with mesocoxal sockets (
|
||
<figureCitation id="13642A02FFFF18208C463A8B009BFC8C" box="[815,983,151,176]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="40" pageNumber="41">Figs. 105, 107</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFFF18208C8C3A84070BFC8C" box="[997,1095,152,176]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="40" pageNumber="41">109–110</figureCitation>
|
||
) located on their mesoventral surface and visible in ventral view.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE3AC30641FD03" blockId="40.[151,1437,151,680]" pageId="40" pageNumber="41">
|
||
Metaventrite (
|
||
<figureCitation id="13642A02FFFF18208E033AC3015FFCC4" box="[362,531,223,248]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="40" pageNumber="41">Figs. 109–110</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFFF18208D4B3AFC0171FCC4" box="[546,573,224,248]" italics="true" pageId="40" pageNumber="41">
|
||
<date id="FFE11047FFFF18208D4B3AFC0171FCC4" box="[546,573,224,248]" pageId="40" pageNumber="41">v3</date>
|
||
</emphasis>
|
||
) subrectangular, anteriorly fused with mesoventrite, posteriorly shallowly bisinuate and with broadly subtriangular metaventral intercoxal process (
|
||
<figureCitation id="13642A02FFFF18208C8F3B180705FD20" box="[998,1097,260,285]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="40" pageNumber="41">Fig. 111</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFFF18208B303B1907C3FD20" box="[1113,1167,261,284]" italics="true" pageId="40" pageNumber="41">mtvp</emphasis>
|
||
) bearing small median notch; anterior metaventral process present (
|
||
<figureCitation id="13642A02FFFF18208DE13B3B01A6FD7C" box="[648,746,295,320]" captionStart="FIGURES 105 – 108" captionStartId="41.[151,264,1463,1486]" captionTargetBox="[151,1436,195,1442]" captionTargetId="figure@41.[151,1436,195,1442]" captionTargetPageId="41" captionText="FIGURES 105 – 108. Morphology of Spinosciacharis acheron sp. n. Mesothorax and anterior part of metathorax in ventral view (105), intermesocoxal part of pterothorax in ventral view (106), left mesocoxal socket in ventral view (107), opening of right ventrolateral fovea in ventral view (108). Abbreviations: amvp, anterior metaventral process; ar, anterior ridge; mscc, mesocoxal cavity; mcp, mesocoxal projection; mscs, mesocoxal socket; msvp, mesoventral process; pcr, procoxal rest; pre, prepectus; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286535/files/figure.png" pageId="40" pageNumber="41">Fig. 106</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFFF18208D9C3B35007EFD7C" box="[757,818,297,320]" italics="true" pageId="40" pageNumber="41">amvp</emphasis>
|
||
). Metanepisterna and metepimera narrow.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE3B50075BFD58" blockId="40.[151,1437,151,680]" box="[199,1047,332,357]" pageId="40" pageNumber="41">
|
||
Metafurca with long stalk and divergent lateral furcal arms (
|
||
<figureCitation id="13642A02FFFF18208C003B500085FD58" box="[873,969,332,357]" captionStart="FIGURES 109 – 111" captionStartId="42.[151,264,1720,1743]" captionTargetBox="[151,1436,663,1698]" captionTargetId="figure@42.[151,1436,663,1698]" captionTargetPageId="42" captionText="FIGURES 109 – 111. Morphology of Spinosciacharis acheron sp. n. Pterothorax in ventral view (109 – 110), metaventral intercoxal process in ventral view (111) Abbreviations: aest 3, metanepisternum; amvp, anterior metaventral process; ar, anterior ridge; cxs 3, metacoxa; lmfa, lateral metafurcal arm; mcp, mesocoxal projection; mscc, mesocoxal cavity; mscs, mesocoxal socket; msf, mesofurca; msvp, mesoventral process; mtvp, metaventral intercoxal process; pcr, procoxal rest; pre, prepectus; tr 3, metatrochanter; v 3, metaventrite; vlf, ventrolateral fovea." httpUri="https://zenodo.org/record/286536/files/figure.png" pageId="40" pageNumber="41">Fig. 110</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFFF18208CBC3B50074BFD59" box="[981,1031,332,357]" italics="true" pageId="40" pageNumber="41">lmfa</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE3B73070CFD91" blockId="40.[151,1437,151,680]" pageId="40" pageNumber="41">
|
||
Elytra (
|
||
<figureCitation id="13642A02FFFF18208E743B7302C0FDB4" box="[285,396,367,392]" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="40" pageNumber="41">Figs. 104</figureCitation>
|
||
,
|
||
<figureCitation id="13642A02FFFF18208EF33B6C0179FDB4" box="[410,565,367,392]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="40" pageNumber="41">112, 114–116</figureCitation>
|
||
) oval, each with two moderately large, deep and asetose basal foveae (
|
||
<figureCitation id="13642A02FFFF18208A063B730388FD90" captionStart="FIGURES 102 – 104" captionStartId="40.[151,264,1499,1522]" captionTargetBox="[151,1436,723,1478]" captionTargetId="figure@40.[151,1436,723,1479]" captionTargetPageId="40" captionText="FIGURES 102 – 104. Morphology of Spinosciacharis acheron sp. n. Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc 2, mesoscutum; scl 2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture." httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="40" pageNumber="41">Fig. 104</figureCitation>
|
||
;
|
||
<emphasis id="B92BEA95FFFF18208FA63B8803BDFD91" box="[207,241,404,429]" italics="true" pageId="40" pageNumber="41">bef</emphasis>
|
||
) not connected by groove; humeral denticles and sub-humeral lines absent.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE3BAB001CFDEC" blockId="40.[151,1437,151,680]" box="[199,848,439,464]" pageId="40" pageNumber="41">
|
||
<collectingCountry id="F3487617FFFF18208FAE3BAB03B3FDEC" box="[199,255,439,464]" name="India" pageId="40" pageNumber="41">Hind</collectingCountry>
|
||
wings well-developed, about twice as long as elytra.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE3BC1078BFE00" blockId="40.[151,1437,151,680]" pageId="40" pageNumber="41">
|
||
Legs (
|
||
<figureCitation id="13642A02FFFF18208E7A3BC002BBFDC9" box="[275,503,476,501]" captionStart="FIGURES 112 – 116" captionStartId="44.[151,262,1924,1947]" captionTargetBox="[211,1378,208,1879]" captionTargetId="figure@44.[198,1388,195,1896]" captionTargetPageId="44" captionText="FIGURES 112 – 116. Spinosciacharis spp. Dorsal habitus of males (112, 114 – 116), and original set of labels of holotype (113). Spinosciacharis puncticeps (Franz) comb. n. (112 – 113); Spinosciacharis sassy sp. n. (114), Spinosciacharis clydensis sp. n. (115), and Spinosciacharis acheron sp. n. (116)." httpUri="https://zenodo.org/record/286537/files/figure.png" pageId="40" pageNumber="41">Figs. 112, 114–116</figureCitation>
|
||
) long and slender; procoxae subglobose, mesocoxae oval, metacoxae strongly transverse; all trochanters short; all femora moderately strongly clavate; tibiae and tarsi long and slender. Males of all species have variously shaped projections on ventral margin of metafemora (
|
||
<figureCitation id="13642A02FFFF18208B7F383807F7FE00" box="[1046,1211,548,573]" captionStart="FIGURES 117 – 120" captionStartId="45.[151,264,696,719]" captionTargetBox="[161,1431,193,663]" captionTargetId="figure@45.[151,1436,193,675]" captionTargetPageId="45" captionText="FIGURES 117 – 120. Spinosciacharis spp. distal part of left metafemur in posterior view. Spinosciacharis puncticeps (Franz) comb. n. (117); Spinosciacharis sassy sp. n. (118), Spinosciacharis clydensis sp. n. (119), and Spinosciacharis acheron sp. n. (120)." httpUri="https://zenodo.org/record/286538/files/figure.png" pageId="40" pageNumber="41">Figs. 117–120</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE385B02FBFE9B" blockId="40.[151,1437,151,680]" pageId="40" pageNumber="41">
|
||
Aedeagus (
|
||
<figureCitation id="13642A02FFFF18208E3B385B014EFE5C" box="[338,514,583,608]" captionStart="FIGURES 121 – 128" captionStartId="46.[151,264,1877,1900]" captionTargetBox="[168,1425,220,1825]" captionTargetId="figure@46.[151,1436,193,1856]" captionTargetPageId="46" captionText="FIGURES 121 – 128. Aedeagi of Spinosciacharis spp. in ventral (121, 123, 125, 127) and lateral (122, 124, 126, 128) views. Spinosciacharis puncticeps (Franz) comb. n. (121 – 122); Spinosciacharis sassy sp. n. (123 – 124), Spinosciacharis clydensis sp. n. (125 – 126), and Spinosciacharis acheron sp. n. (127 – 128)." httpUri="https://zenodo.org/record/286539/files/figure.png" pageId="40" pageNumber="41">Figs. 121–128</figureCitation>
|
||
) elongate and moderately darkly sclerotized, relatively thin-walled, with symmetrical median lobe and symmetrical, complicated assemblage of internal sclerites; parameres free and slender, with apical setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF20660FFFFF18208FFE3FC700C8FA50" httpUri="https://zenodo.org/record/286534/files/figure.png" pageId="40" pageNumber="41" targetBox="[151,1436,723,1478]" targetPageId="40">
|
||
<paragraph id="8BE03687FFFF18208FFE3FC700C8FA50" blockId="40.[151,1436,1499,1644]" pageId="40" pageNumber="41">
|
||
<emphasis id="B92BEA95FFFF18208FFE3FC7023CF9CD" bold="true" box="[151,368,1499,1522]" pageId="40" pageNumber="41">FIGURES 102–104.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C5F4D04FFFF18208D493FC00051F9CE" box="[544,797,1500,1522]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="acheron" status="sp. nov.">
|
||
<emphasis id="B92BEA95FFFF18208D493FC00051F9CE" box="[544,797,1500,1522]" italics="true" pageId="40" pageNumber="41">Spinosciacharis acheron</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B92BEA95FFFF18208C4F3FC0002EF9CD" bold="true" box="[806,866,1500,1521]" pageId="40" pageNumber="41">
|
||
<taxonomicNameLabel id="A21857EEFFFF18208C4F3FC0002EF9CD" box="[806,866,1500,1521]" pageId="40" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Prothorax in ventral view (102), base of pronotum in dorsal view (103), mesonotum and elytral base (104). Abbreviations: bef, basal elytral fovea; bst, basisternal part of prosternum; eal, elytral articulating lobe; hyr, hypomeral ridge; ip, internal antebasal pit; li, lateral antebasal impression; mc, median carina; nss, pronotosternal suture; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; sc2, mesoscutum; scl2, mesoscutellum; slc, sublateral carina; sss, mesoscutoscutellar suture.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C345650CFFFF18208FAE3C8807FFFB24" pageId="40" pageNumber="41" type="etymology">
|
||
<paragraph id="8BE03687FFFF18208FAE3C88074FFAEC" blockId="40.[151,1436,1684,2032]" pageId="40" pageNumber="41">
|
||
<emphasis id="B92BEA95FFFF18208FAE3C880202FA91" bold="true" box="[199,334,1684,1709]" pageId="40" pageNumber="41">Etymology.</emphasis>
|
||
<taxonomicName id="4C5F4D04FFFF18208E3E3C880146FA91" box="[343,522,1684,1709]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFF18208E3E3C880146FA91" box="[343,522,1684,1709]" italics="true" pageId="40" pageNumber="41">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
is a combination of the stem
|
||
<taxonomicName id="4C5F4D04FFFF18208C013C880092FA91" box="[872,990,1684,1709]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFF18208C013C880092FA91" box="[872,990,1684,1709]" italics="true" pageId="40" pageNumber="41">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
(a genus name) with the prefix
|
||
<emphasis id="B92BEA95FFFF18208A3F3C8906DFFA90" box="[1366,1427,1685,1708]" italics="true" pageId="40" pageNumber="41">spino</emphasis>
|
||
- referring to the femoral spines found in males of all species. Gender feminine.
|
||
</paragraph>
|
||
<paragraph id="8BE03687FFFF18208FAE3CC007FFFB24" blockId="40.[151,1436,1684,2032]" pageId="40" pageNumber="41">
|
||
<emphasis id="B92BEA95FFFF18208FAE3CC0017EFAC9" bold="true" box="[199,562,1756,1781]" pageId="40" pageNumber="41">Composition and distribution</emphasis>
|
||
(
|
||
<figureCitation id="13642A02FFFF18208D2E3CC001E2FAC9" box="[583,686,1756,1781]" captionStart="FIGURES 193 – 198" captionStartId="72.[151,264,1762,1785]" captionTargetBox="[195,1376,193,1707]" captionTargetId="figure@72.[191,1378,193,1739]" captionTargetPageId="72" captionText="FIGURES 193 – 198. Distribution of revised Australian species of Horaeomorphus Schaufuss (193), Leascydmus gen. n. (194), Scydmaenozila gen. n. (195), Spinosciacharis gen. n. (196), Sciacharoides gen. n. (197), and Sciacharis Broun. Horaeomorphus anthicoides (Lea) comb. n. (193 a), Horaeomorphus verus sp. n. (193 b), Leascydmus simplicicornis (Lea) comb. n. (194 a), Leascydmus armstrongi sp. n. (194 b), Scydmaenozila macrosticta (Lea) comb. n. (195 a), Scydmaenozila latipennis (Lea) comb. n. (195 b), Scydmaenozila mallacootana sp. n. (195 c), Scydmaenozila inexpectata sp. n. (195 d), Spinosciacharis puncticeps (Franz) comb. n. (196 a), Spinosciacharis sassy sp. n. (196 b), Spinosciacharis clydensis sp. n. (196 c), Spinosciacharis acheron sp. n. (196 d), Sciacharoides thompsoni (Franz) comb. n. (197 a), Sciacharoides eucalypti (Franz) comb. n. (197 b), Sciacharoides maipotonensis (Franz) comb. n. (197 c), Sciacharoides mantistamborinensis (Franz) comb. n. (197 d), and Sciacharis (Maorinus) tasmaniensis (Franz) comb. n. (198)." httpUri="https://zenodo.org/record/286555/files/figure.png" pageId="40" pageNumber="41">Fig. 196</figureCitation>
|
||
). Three species of
|
||
<taxonomicName id="4C5F4D04FFFF18208CFD3CC0070BFAC9" box="[916,1095,1756,1781]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFF18208CFD3CC0070BFAC9" box="[916,1095,1756,1781]" italics="true" pageId="40" pageNumber="41">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
are distributed in the southeastern part of the Australian continent (New South
|
||
<collectingCountry id="F3487617FFFF18208DB63CE3006FFB24" box="[735,803,1791,1816]" name="United Kingdom" pageId="40" pageNumber="41">Wales</collectingCountry>
|
||
and Victoria) and one in Tasmania.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C345650CFFFF18228FAE3D3801B4FE63" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41" type="discussion">
|
||
<paragraph id="8BE03687FFFF18228FAE3D3801B4FE63" blockId="40.[151,1436,1684,2032]" lastBlockId="42.[151,1436,151,608]" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
|
||
<emphasis id="B92BEA95FFFF18208FAE3D380277FB01" bold="true" box="[199,315,1828,1853]" pageId="40" pageNumber="41">Remarks.</emphasis>
|
||
Among all Australian genera of
|
||
<taxonomicName id="4C5F4D04FFFF18208DC53D380019FB01" box="[684,853,1828,1853]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5F4D04FFFF18208C083D380758FB01" box="[865,1044,1828,1853]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFF18208C083D380758FB01" box="[865,1044,1828,1853]" italics="true" pageId="40" pageNumber="41">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C5F4D04FFFF18208B8D3D3806D7FB01" box="[1252,1435,1828,1853]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFF18208B8D3D3806D7FB01" box="[1252,1435,1828,1853]" italics="true" pageId="40" pageNumber="41">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
in the general body form, but to
|
||
<taxonomicName id="4C5F4D04FFFF18208D6A3D5B0135FB5C" box="[515,633,1863,1888]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFF18208D6A3D5B0135FB5C" box="[515,633,1863,1888]" italics="true" pageId="40" pageNumber="41">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFF18208DE93D5401F7FB63" box="[640,699,1864,1887]" pageId="40" pageNumber="41" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
in ventral structures. The flattened and slender body with weakly rounded sides of elytra, the shape of pronotum, and the femoral projections in males may be mistaken with similar features of some
|
||
<taxonomicName id="4C5F4D04FFFF18208E003D93016DFB94" box="[361,545,1935,1960]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFF18208E003D93016DFB94" box="[361,545,1935,1960]" italics="true" pageId="40" pageNumber="41">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="4C5F4D04FFFF18208DFA3D93000AFB94" box="[659,838,1935,1960]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFF18208DFA3D93000AFB94" box="[659,838,1935,1960]" italics="true" pageId="40" pageNumber="41">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from
|
||
<taxonomicName id="4C5F4D04FFFF18208B2F3D9307B2FB94" box="[1094,1278,1935,1960]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFF18208B2F3D9307B2FB94" box="[1094,1278,1935,1960]" italics="true" pageId="40" pageNumber="41">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
in numerous characters: the absence of pits on posteromesal margins of weakly marked supraantennal tubercles (in
|
||
<taxonomicName id="4C5F4D04FFFF18208FFE3DCB0203FBCC" box="[151,335,2007,2032]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFF18208FFE3DCB0203FBCC" box="[151,335,2007,2032]" italics="true" pageId="40" pageNumber="41">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
pits present and tubercles prominent); posterior tentorial pits located far in front of the anterior margin of 'neck region' (in
|
||
<taxonomicName id="4C5F4D04FFFE18218E8A3C7301D6FAB4" box="[483,666,1647,1672]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218E8A3C7301D6FAB4" box="[483,666,1647,1672]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
pits are adjacent to the anterior margin of 'neck region'); much narrower occipital constriction; absence of median antebasal pronotal pit (present in
|
||
<taxonomicName id="4C5F4D04FFFE18218B253C8F064FFA90" box="[1100,1283,1683,1708]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218B253C8F064FFA90" box="[1100,1283,1683,1708]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); presence of sublateral pronotal carinae (absent in
|
||
<taxonomicName id="4C5F4D04FFFE18218D3F3CAB0041FAEC" box="[598,781,1719,1744]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218D3F3CAB0041FAEC" box="[598,781,1719,1744]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); narrow, ridge-like prosternal intercoxal carina (carina narrow but subrectangular with distinct lateral margins in
|
||
<taxonomicName id="4C5F4D04FFFE18218C4E3CC70093FAC8" box="[807,991,1755,1780]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218C4E3CC70093FAC8" box="[807,991,1755,1780]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); basal elytral foveae not connected by inversely U-shaped groove (in
|
||
<taxonomicName id="4C5F4D04FFFE18218E923CE301FFFB24" box="[507,691,1791,1816]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218E923CE301FFFB24" box="[507,691,1791,1816]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
groove present); narrow anterior ridge of mesoventrite (broad in
|
||
<taxonomicName id="4C5F4D04FFFE18218FFE3D3F0203FB00" box="[151,335,1827,1852]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218FFE3D3F0203FB00" box="[151,335,1827,1852]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); presence of ventrolateral mesothoracic foveae (absent in
|
||
<taxonomicName id="4C5F4D04FFFE18218B2D3D3F07B0FB00" box="[1092,1276,1827,1852]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218B2D3D3F07B0FB00" box="[1092,1276,1827,1852]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); mesocoxae indistinctly separated by narrow mesoventral process only in their anterior half (distinctly separated in
|
||
<taxonomicName id="4C5F4D04FFFE18218FFE3D770203FBB8" box="[151,335,1899,1924]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218FFE3D770203FBB8" box="[151,335,1899,1924]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); presence of anterior metaventral process (absent in
|
||
<taxonomicName id="4C5F4D04FFFE18218CCA3D770717FBB8" box="[931,1115,1899,1924]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218CCA3D770717FBB8" box="[931,1115,1899,1924]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
); and metaventral intercoxal process without spines (with a pair of long spines in
|
||
<taxonomicName id="4C5F4D04FFFE18218D983D9300E4FB94" box="[753,936,1935,1960]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFE18218D983D9300E4FB94" box="[753,936,1935,1960]" italics="true" pageId="41" pageNumber="42">Horaeomorphus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C5F4D04FFFE18218CA83D930738FB94" box="[961,1140,1935,1960]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFE18218CA83D930738FB94" box="[961,1140,1935,1960]" italics="true" pageId="41" pageNumber="42">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C5F4D04FFFE18218A653D9306CEFB94" box="[1292,1410,1935,1960]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFE18218A653D9306CEFB94" box="[1292,1410,1935,1960]" italics="true" pageId="41" pageNumber="42">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFE18218AE33D8C03F5FBF7" pageId="41" pageNumber="42" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
in the general body shape (especially the shape of the head), all ventral characters of the head; the presence of pronotal internal antebasal pits and sublateral carinae; feebly developed prosternal intercoxal carina; presence of two distinct basal elytral foveae; presence of only ventrolateral mesothoracic fovea; and the shape of metaventral intercoxal process.
|
||
<taxonomicName id="4C5F4D04FFFD18228E133AA00161FCE9" box="[378,557,188,213]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFD18228E133AA00161FCE9" box="[378,557,188,213]" italics="true" pageId="42" pageNumber="43">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C5F4D04FFFD18228DB83AA0000BFCE9" box="[721,839,188,213]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFD18228DB83AA0000BFCE9" box="[721,839,188,213]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFD18228C3D3AA100D8FCE8" box="[852,916,189,212]" pageId="42" pageNumber="43" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
in robust subtriangular mandibles with two subapical teeth (one small subapical tooth beneath narrow apical tooth and above strongly broadened mandibular base in
|
||
<taxonomicName id="4C5F4D04FFFD18228F9B3B180224FD21" box="[242,360,260,285]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFD18228F9B3B180224FD21" box="[242,360,260,285]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFD18228E1B3B1902FFFD20" box="[370,435,260,284]" pageId="42" pageNumber="43" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
); sublateral pronotal carinae extending much farther anteriorly; prosternum completely demarcated from adcoxal parts of hypomera (in
|
||
<taxonomicName id="4C5F4D04FFFD18228DA53B3B000EFD7C" box="[716,834,295,320]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFD18228DA53B3B000EFD7C" box="[716,834,295,320]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFD18228C253B3400C7FD03" box="[844,907,296,319]" pageId="42" pageNumber="43" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
prosternum partly fused with hypomera); the mesoventral intercoxal process not interrupted (in
|
||
<taxonomicName id="4C5F4D04FFFD18228DA43B50000FFD59" box="[717,835,332,357]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFD18228DA43B50000FFD59" box="[717,835,332,357]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFD18228C223B5100CAFD58" box="[843,902,333,356]" pageId="42" pageNumber="43" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
interrupted near anterior margins of mesocoxal cavities); and the presence of anterior metaventral process (absent in
|
||
<taxonomicName id="4C5F4D04FFFD18228CA23B73070DFDB4" box="[971,1089,367,392]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B92BEA95FFFD18228CA23B73070DFDB4" box="[971,1089,367,392]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21857EEFFFD18228B253B6C07C3FDB4" box="[1100,1167,368,392]" pageId="42" pageNumber="43" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
). The same characters, except for the prosternum completely separated vs. partly fused with hypomera differentiate
|
||
<taxonomicName id="4C5F4D04FFFD18228BC23B880612FD91" box="[1195,1374,404,429]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFD18228BC23B880612FD91" box="[1195,1374,404,429]" italics="true" pageId="42" pageNumber="43">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C5F4D04FFFD18228FFE3BAB02D4FDEC" authority="(Maorinus)" baseAuthorityName="Maorinus" box="[151,408,439,464]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFD18228FFE3BAB0241FDEC" box="[151,269,439,464]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
(
|
||
<emphasis id="B92BEA95FFFD18228E483BA502C2FDEC" box="[289,398,441,464]" italics="true" pageId="42" pageNumber="43">Maorinus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
. The latter taxon has an elongated head, distinctly thickened maxillary palpi, hypomeral ridges not reaching posterior margins of adcoxal parts of hypomera; and long mesoventral intercoxal process extending from just behind anterior ridge of mesoventrite up to posterior margins of mesocoxal cavities (all these characters different in
|
||
<taxonomicName id="4C5F4D04FFFD18228EF43838011CFE01" box="[413,592,548,573]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFD18228EF43838011CFE01" box="[413,592,548,573]" italics="true" pageId="42" pageNumber="43">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
). The similarities between
|
||
<taxonomicName id="4C5F4D04FFFD18228CE73838070DFE01" box="[910,1089,548,573]" class="Insecta" family="Staphylinidae" genus="Spinosciacharis" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="73" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B92BEA95FFFD18228CE73838070DFE01" box="[910,1089,548,573]" italics="true" pageId="42" pageNumber="43">Spinosciacharis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5F4D04FFFD18228B17383807B8FE01" box="[1150,1268,548,573]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFD18228B17383807B8FE01" box="[1150,1268,548,573]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
may in future justify placing the former as a subgenus of
|
||
<taxonomicName id="4C5F4D04FFFD18228D13385B01BCFE5C" box="[634,752,583,608]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92BEA95FFFD18228D13385B01BCFE5C" box="[634,752,583,608]" italics="true" pageId="42" pageNumber="43">Sciacharis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |