416 lines
56 KiB
XML
416 lines
56 KiB
XML
<document id="590B2BB5B6E2DC22FBF4DE4B17B9D522" ID-DOI="10.5281/zenodo.186599" ID-GBIF-Dataset="36da1480-9be8-4f00-aba5-23883071cf46" ID-ISSN="1175-5326" ID-Zenodo-Dep="186599" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461053771789" checkinUser="plazi" docAuthor="Staples, David A." docDate="2009" docId="03E48F65D64E5165FF7EF8A4FAD9FA4E" docLanguage="en" docName="zt02051p068.pdf" docOrigin="Zootaxa 2051" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Rhopalorhynchus magdalena Staples, 2009, sp. nov." docType="treatment" docVersion="8" lastPageNumber="67" masterDocId="FFDDF71DD64E5160FFE9FFF8FFD8FFB1" masterDocTitle="A new species of Rhopalorhynchus (Pycnogonida: Colossendeidae) from Lizard Island, North Queensland, Australia" masterLastPageNumber="68" masterPageNumber="62" pageNumber="62" updateTime="1698580492714" updateUser="plazi">
|
||
<mods:mods id="7AEAF7ADFEA0D38BB670CFDA94A9DFEF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0280CF98F2E62A455042703F23E3EC89">
|
||
<mods:title id="6E58C6E4A793B63CA7220F9A2DCB8C00">A new species of Rhopalorhynchus (Pycnogonida: Colossendeidae) from Lizard Island, North Queensland, Australia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="22912EDD4B62BE2BD281AF88EAD4BDCD" type="personal">
|
||
<mods:role id="B488313BABC6E51CB49DDDD46AE0A41B">
|
||
<mods:roleTerm id="4C20C744EA872804DFD9575645E1449C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D8D8BB960810C5B3EEDC4E6792680546">Staples, David A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3A59EE01E3339C557FA46062D47BB14E">text</mods:typeOfResource>
|
||
<mods:relatedItem id="075BDB05EC5BFBEB013273FDC8504AF0" type="host">
|
||
<mods:titleInfo id="51203AE5CF5FA9782A83707805BC2EDF">
|
||
<mods:title id="9CD76661C47BD2D340EFFBE17A7D165C">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="4FA811880E1FFF414B3534FAA2B25560">
|
||
<mods:date id="F53CD73E66969D67EDB6AB75381E4A51">2009</mods:date>
|
||
<mods:detail id="1C010251C5BAAB23F3B2AF91CA085529" type="volume">
|
||
<mods:number id="FD85FB4EF42C185C788D705750E0D412">2051</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="CC98C468D6D6F1E33B6845FA59F793D2" unit="page">
|
||
<mods:start id="4E1BF71E3AA315EEE697F41DD8E9DA94">62</mods:start>
|
||
<mods:end id="54AEC9842DB02FC7FBBCA5FC0592B3E8">68</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="D29EBB3EA071FCB5A491E4C0EEAD6CFA">journal article</mods:classification>
|
||
<mods:identifier id="78C1ACAA1E756849DFB5CE7E9268406F" type="DOI">10.5281/zenodo.186599</mods:identifier>
|
||
<mods:identifier id="442C4CC7EBDBE27C5FF362A15F9C0958" type="GBIF-Dataset">36da1480-9be8-4f00-aba5-23883071cf46</mods:identifier>
|
||
<mods:identifier id="EFE8FCC2B9A06E58FBE1CEC4FB737528" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="FB16824B70414C1CBAD0453A089A94B4" type="Zenodo-Dep">186599</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E48F65D64E5165FF7EF8A4FAD9FA4E" ID-DOI="http://doi.org/10.5281/zenodo.5694033" ID-GBIF-Taxon="119612557" ID-Zenodo-Dep="5694033" LSID="urn:lsid:plazi:treatment:03E48F65D64E5165FF7EF8A4FAD9FA4E" httpUri="http://treatment.plazi.org/id/03E48F65D64E5165FF7EF8A4FAD9FA4E" lastPageId="5" lastPageNumber="67" pageId="0" pageNumber="62">
|
||
<subSubSection id="C3576DF8D64E5160FF7EF8A4FEF1F828" pageId="0" pageNumber="62" type="nomenclature">
|
||
<paragraph id="8BF23E73D64E5160FF7EF8A4FDB9F8C7" blockId="0.[151,609,1884,1945]" box="[151,609,1884,1910]" pageId="0" pageNumber="62">
|
||
<heading id="D0BA891FD64E5160FF7EF8A4FDB9F8C7" bold="true" box="[151,609,1884,1910]" fontSize="11" level="1" pageId="0" pageNumber="62" reason="1">
|
||
<emphasis id="B939E261D64E5160FF7EF8A4FDB9F8C7" bold="true" box="[151,609,1884,1910]" pageId="0" pageNumber="62">
|
||
<taxonomicName id="4C4D45F0D64E5160FF7EF8A4FE24F8C7" ID-CoL="4SLP6" box="[151,508,1884,1910]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="0" pageNumber="62" phylum="Arthropoda" rank="species" species="magdalena" status="sp. nov.">
|
||
<emphasis id="B939E261D64E5160FF7EF8A4FE24F8C7" bold="true" box="[151,508,1884,1910]" italics="true" pageId="0" pageNumber="62">Rhopalorhynchus magdalena</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A5F1AD64E5160FDEDF8A4FDB9F8C7" box="[516,609,1884,1910]" pageId="0" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64E5160FF7EF887FEF1F828" blockId="0.[151,609,1884,1945]" box="[151,297,1919,1945]" pageId="0" pageNumber="62">
|
||
<figureCitation id="137622F6D64E5160FF7EF887FEDDF828" box="[151,261,1919,1945]" captionStart="FIGURE 1" captionStartId="2.[151,255,1942,1966]" captionTargetBox="[184,1411,186,1920]" captionTargetId="figure@2.[168,1422,173,1928]" captionTargetPageId="2" captionText="FIGURE 1. Rhopalorhynchus magdalena sp nov. female holotype (QM S 83740): A, trunk, lateral; B, trunk, dorsal; C, cephalon lateral; D, cephalon, dorsal; E, cephalon, ventral; F, leg, 3; G, leg 4, tarsus, propodus, claw; H, coxae, genital pore; I, male paratype (J 58389) abdomen, ventral." httpUri="https://zenodo.org/record/186600/files/figure.png" pageId="0" pageNumber="62">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="137622F6D64E5160FEFAF887FEFDF828" box="[275,293,1919,1945]" captionStart="FIGURE 2" captionStartId="3.[151,259,1906,1930]" captionTargetBox="[216,1392,181,1880]" captionTargetId="figure@3.[189,1404,180,1892]" captionTargetPageId="3" captionText="FIGURE 2. Rhopalorhynchus magdalena sp. nov. female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth. Rhopalorhynchus gracillimus (Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis." httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="0" pageNumber="62">2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576DF8D64E5161FF7EF834FE71FEFD" lastPageId="1" lastPageNumber="63" pageId="0" pageNumber="62" type="materials_examined">
|
||
<paragraph id="8BF23E73D64E5161FF7EF834FEDAFF69" blockId="0.[151,1436,1996,2023]" lastBlockId="1.[151,1437,152,2034]" lastPageId="1" lastPageNumber="63" pageId="0" pageNumber="62">
|
||
<emphasis id="B939E261D64E5160FF7EF834FDD6F857" bold="true" box="[151,526,1996,2022]" pageId="0" pageNumber="62">
|
||
Material examined.
|
||
<typeStatus id="54F680D1D64E5160FE73F834FDD6F857" box="[410,526,1996,2022]" pageId="0" pageNumber="62" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
: North Qld., Lizard I. (14˚ 6₁.061˝S, 145˚ ₁5.365˝E),
|
||
<quantity id="4CB59396D64E5160FB4DF834FB3DF857" box="[1188,1253,1996,2022]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="0" pageNumber="62" unit="m" value="15.0">15 m</quantity>
|
||
, amongst coral rubble, sand, collected from crevices, hand dredge, M. Blazewicz-Paszkowycz,
|
||
<date id="FFF318B3D64F5161FBB6FF60FB23FF03" box="[1119,1275,152,178]" pageId="1" pageNumber="63" value="2008-04-18">18 Apr 2008</date>
|
||
, QM
|
||
<accessionNumber id="941EA390D64F5161FAA8FF60FA44FF03" box="[1345,1436,152,178]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83741" pageId="1" pageNumber="63" type="EnaNcbi">S83741</accessionNumber>
|
||
(female).
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FFF1DFE71FEFD" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
<typeStatus id="54F680D1D64F5161FF2FFF1DFE9BFF4E" box="[198,323,229,255]" pageId="1" pageNumber="63" type="paratype">
|
||
<emphasis id="B939E261D64F5161FF2FFF1DFE9BFF4E" bold="true" box="[198,323,229,255]" pageId="1" pageNumber="63">Paratypes</emphasis>
|
||
</typeStatus>
|
||
: Lizard I. (14˚ 6₁.567˝S, 145˚15•325˝E),
|
||
<quantity id="4CB59396D64F5161FCABFF1DFC5DFF4E" box="[834,901,229,255]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="63" unit="m" value="10.0">10 m</quantity>
|
||
, amongst coral rubble, sand, collected from crevices, hand dredge, M. Blazewicz-Paszkowycz,
|
||
<date id="FFF318B3D64F5161FCE3FEF4FC89FE97" box="[778,849,268,294]" pageId="1" pageNumber="63">4 Apr</date>
|
||
0 8, QM
|
||
<accessionNumber id="941EA390D64F5161FC28FEF4FBC5FE97" box="[961,1053,268,294]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83740" pageId="1" pageNumber="63" type="EnaNcbi">S83740</accessionNumber>
|
||
(
|
||
<specimenCount id="9D4BF5FAD64F5161FBDBFEF4FB50FE97" box="[1074,1160,268,294]" pageId="1" pageNumber="63" type="male">1 male</specimenCount>
|
||
). Same collection data
|
||
<collectionCode id="ED5CA6B6D64F5161FF7EFECAFF04FEFD" LSID="urn:lsid:biocol.org:col:34978" box="[151,220,306,332]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="1" pageNumber="63">NMV</collectionCode>
|
||
<accessionNumber id="941EA390D64F5161FF0AFECAFEE0FEFD" box="[227,312,306,332]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J58389" pageId="1" pageNumber="63" type="EnaNcbi">J58389</accessionNumber>
|
||
(
|
||
<specimenCount id="9D4BF5FAD64F5161FEA3FECAFE44FEFD" box="[330,412,306,332]" pageId="1" pageNumber="63" type="male">1 male</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576DF8D64F5161FF2FFEA0FEA4FDEA" pageId="1" pageNumber="63" type="diagnosis">
|
||
<paragraph id="8BF23E73D64F5161FF2FFEA0FEA4FDEA" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
<emphasis id="B939E261D64F5161FF2FFEA0FE9EFEC3" bold="true" box="[198,326,344,370]" pageId="1" pageNumber="63">Diagnosis.</emphasis>
|
||
A small species compared to its congeners, leg span about
|
||
<quantity id="4CB59396D64F5161FBE5FEA1FBBCFEC2" box="[1036,1124,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="63" unit="mm" value="20.0">20 mm</quantity>
|
||
. Lateral processes 2 and 3 separated by about 6 times their basal diameter, ocular tubercle bearing tall, finger-like process about threequarters of total height. Inflated part of proboscis about 3 times as long as maximum width, tooth prominent, positioned at about 30% along the inflated part, height almost one-fifth proboscis width, basal stalk about 40% of total length of proboscis. Palp segment 10, 3.9 times as long as wide. Legs brevitarsal, tarsal ratio about 55%; tarsus 34–37 % of propodus length; propodus marginally wider distally, claw about one-third length of propodus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576DF8D64F5164FF2CFD9FFF0AFE97" lastPageId="4" lastPageNumber="66" pageId="1" pageNumber="63" type="description">
|
||
<paragraph id="8BF23E73D64F5161FF2CFD9FFCECFC21" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
<emphasis id="B939E261D64F5161FF2CFD9FFE3FFD30" bold="true" box="[197,487,615,641]" pageId="1" pageNumber="63">
|
||
Description of
|
||
<typeStatus id="54F680D1D64F5161FE95FD9FFE3FFD30" box="[380,487,615,641]" pageId="1" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Leg span about
|
||
<quantity id="4CB59396D64F5161FD46FD90FD26FD33" box="[687,766,616,642]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="63" unit="mm" value="20.0">20mm</quantity>
|
||
. Trunk segmented, segment 1 about 0.53 times as long as segment 2, segment 3 about 0.82 times as long as segment 2, segment 4 half length segment of 3; lateral processes short, constricted at bases, processes 1 and 2 separated by almost 6 times their own basal diameter, processes 2 and 3 separated by about 5 times their basal width., processes 3 and 4 separated by about twice their own width. Ocular tubercle with tall, finger-like apical process (about three-quarters height of basal part), bluntly rounded at tip, leaning slightly forward, 4 eyes, anterior eyes marginally larger than posterior pair, pigmented, lateral sensory organs present. Abdomen small, oval, widely cleft at tip, articulated at base, placed between the fourth pair of legs, directed ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FFC65FC24FBE3" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">Proboscis length a little shorter than trunk, basal stalk 42% of total length; inflated part 3.1 times as long as maximum width, angled down at about 40º to the stalk, tapering distally to rounded tip, tip not bent downwards, dorsal tooth placed at about 32% of inflated part, prominent, height almost one-fifth of maximum proboscis width, proboscis otherwise smooth. Arthrodial membrane broad in ventral view, indicating ability to extend proboscis through wide ventral arc, possibly to vertical position.</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FFBA6FBD2FB5D" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
Palp (
|
||
<figureCitation id="137622F6D64F5161FEE2FBA6FEA9FBC9" box="[267,369,1118,1144]" captionStart="FIGURE 2" captionStartId="3.[151,259,1906,1930]" captionTargetBox="[216,1392,181,1880]" captionTargetId="figure@3.[189,1404,180,1892]" captionTargetPageId="3" captionText="FIGURE 2. Rhopalorhynchus magdalena sp. nov. female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth. Rhopalorhynchus gracillimus (Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis." httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="1" pageNumber="63">Figure 2</figureCitation>
|
||
C) 10–segmented, attached to anterior margin of cephalon, first segment inflated dorsally; segment 2 very short, less than half segment 1, bearing single ventral seta; segment 3, 1.6 times length of segment 5; segment 6 about two-thirds length of segment of 7; segments 7–9 subequal; segment 10 slightly shorter, 3.9 times as long as wide; segments 6 to 10 with long ventral setae.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FFB01FBB5FA1F" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
Oviger (
|
||
<figureCitation id="137622F6D64F5161FEC1FB01FE57FAA2" box="[296,399,1273,1299]" captionStart="FIGURE 2" captionStartId="3.[151,259,1906,1930]" captionTargetBox="[216,1392,181,1880]" captionTargetId="figure@3.[189,1404,180,1892]" captionTargetPageId="3" captionText="FIGURE 2. Rhopalorhynchus magdalena sp. nov. female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth. Rhopalorhynchus gracillimus (Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis." httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="1" pageNumber="63">Figure 2</figureCitation>
|
||
A) attached to ventrolateral surface near anterior margin of cephalon; 3 basal segments short; segment 4 longest, about 1.1 times as long as segment 6, strigilis tightly curled, laying over segment 7, segments 7 to 10 bearing long, fine spines in several irregular rows, distal spine on segment 10 robust, much larger than preceding spines but not forming obvious subchelate structure with the terminal claw, crenulations on spine margins not evident, claw gently curved, stout, width about 75% of length.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FFA42FC2BF8BB" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
Legs (
|
||
<figureCitation id="137622F6D64F5161FEE7FA42FEABFA65" box="[270,371,1466,1492]" captionStart="FIGURE 1" captionStartId="2.[151,255,1942,1966]" captionTargetBox="[184,1411,186,1920]" captionTargetId="figure@2.[168,1422,173,1928]" captionTargetPageId="2" captionText="FIGURE 1. Rhopalorhynchus magdalena sp nov. female holotype (QM S 83740): A, trunk, lateral; B, trunk, dorsal; C, cephalon lateral; D, cephalon, dorsal; E, cephalon, ventral; F, leg, 3; G, leg 4, tarsus, propodus, claw; H, coxae, genital pore; I, male paratype (J 58389) abdomen, ventral." httpUri="https://zenodo.org/record/186600/files/figure.png" pageId="1" pageNumber="63">Figure 1</figureCitation>
|
||
F, G) brevitarsal (tarsal ratio of third leg 56%). Coxae short, coxa 2 longer than coxae 1 or 3; femur clearly the longest segment, gently curved throughout, conspicuously swollen distally, club-like with few, very fine filamentous setae dorsally, one or more longer setae on distodorsal surface; tibia 1 swollen distally but much less so than femur, size and arrangement of setae similar to femur, longest seta dorsodistal, equal to segment width; tibia 2 slightly longer than tibia 1, gently curved, few dorsodistal setae longer than twice segment width; length of tarsus slightly variable between legs, 34–37% of propodus length, proportionally longest on leg 1; propodus gently curved, marginally wider distally; soles of tarsus and propodus densely setose. Terminal claw, 31–36% propodus length. Auxiliary claws absent. Gonopores conspicuous on ventral surface of coxa 2 of all legs, not evident in males.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FF8EEFC2FF8CF" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">
|
||
<emphasis id="B939E261D64F5161FF2FF8EEFEA5F881" bold="true" box="[198,381,1814,1840]" pageId="1" pageNumber="63">Measurements</emphasis>
|
||
(female
|
||
<typeStatus id="54F680D1D64F5161FE0EF8EEFD88F881" box="[487,592,1814,1840]" pageId="1" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
, mm). Trunk length (frontal margin of cephalic segment to tip of fourth lateral process), 2.35; trunk segment 1, 0.45; trunk segment 2, 0.85; trunk segment 3, 0.70; trunk segment 4 (to tip of lateral process) 0.35; width across second lateral processes 0.63.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FF872FA44F815" blockId="1.[151,1437,152,2034]" box="[198,1436,1930,1956]" pageId="1" pageNumber="63">
|
||
Proboscis (using notation of
|
||
<bibRefCitation id="EFDC4382D64F5161FDFDF872FD45F815" author="Stock" box="[532,669,1930,1956]" pageId="1" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock 1958</bibRefCitation>
|
||
): α 1.60; β.88; γ 1.40; ε 0.44; ξ 0.15; δ 0.98; height of tooth, 0.09.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64F5161FF2FF849FE17F843" blockId="1.[151,1437,152,2034]" pageId="1" pageNumber="63">Third leg: coxa 1, 0.13; coxa 2, 0.20; coxa 3, 0.15; femur, 2.78; tibia 1, 2.50; tibia 2, 2.40; tarsus, 0.25; propodus, 0.83; claw 0.28.</paragraph>
|
||
<caption id="DF326EFBD64C5162FF7EF86EFD75F842" httpUri="https://zenodo.org/record/186600/files/figure.png" pageId="2" pageNumber="64" targetBox="[184,1411,186,1920]" targetPageId="2">
|
||
<paragraph id="8BF23E73D64C5162FF7EF86EFD75F842" blockId="2.[151,1436,1942,2035]" pageId="2" pageNumber="64">
|
||
<emphasis id="B939E261D64C5162FF7EF86EFECBF81F" bold="true" box="[151,275,1942,1966]" pageId="2" pageNumber="64">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C4D45F0D64C5162FEC8F86FFD84F81F" box="[289,604,1943,1966]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="magdalena">
|
||
<emphasis id="B939E261D64C5162FEC8F86FFD84F81F" box="[289,604,1943,1966]" italics="true" pageId="2" pageNumber="64">Rhopalorhynchus magdalena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B939E261D64C5162FD8AF86EFD6BF81F" bold="true" box="[611,691,1942,1966]" pageId="2" pageNumber="64">sp nov.</emphasis>
|
||
female holotype (QM
|
||
<accessionNumber id="941EA390D64C5162FC43F86FFBD9F81F" box="[938,1025,1943,1966]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S83740" pageId="2" pageNumber="64" type="EnaNcbi">S83740</accessionNumber>
|
||
): A, trunk, lateral; B, trunk, dorsal; C, cephalon lateral; D, cephalon, dorsal; E, cephalon, ventral; F, leg, 3; G, leg 4, tarsus, propodus, claw; H, coxae, genital pore; I, male paratype (
|
||
<accessionNumber id="941EA390D64C5162FE66F824FE07F842" box="[399,479,2012,2035]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J58389" pageId="2" pageNumber="64" type="EnaNcbi">J58389</accessionNumber>
|
||
) abdomen, ventral.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF326EFBD64D5163FF7EF88AFE5DF840" httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="3" pageNumber="65" targetBox="[216,1392,181,1880]" targetPageId="3">
|
||
<paragraph id="8BF23E73D64D5163FF7EF88AFE5DF840" blockId="3.[151,1437,1906,2033]" pageId="3" pageNumber="65">
|
||
<emphasis id="B939E261D64D5163FF7EF88AFEC1F83B" bold="true" box="[151,281,1906,1930]" pageId="3" pageNumber="65">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C4D45F0D64D5163FEC1F88AFDAAF838" box="[296,626,1906,1929]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="magdalena" status="sp. nov.">
|
||
<emphasis id="B939E261D64D5163FEC1F88AFDAAF838" box="[296,626,1906,1929]" italics="true" pageId="3" pageNumber="65">Rhopalorhynchus magdalena</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A5F1AD64D5163FD92F88AFD16F83B" box="[635,718,1906,1930]" pageId="3" pageNumber="65" rank="species">
|
||
<emphasis id="B939E261D64D5163FD92F88AFD16F83B" bold="true" box="[635,718,1906,1930]" pageId="3" pageNumber="65">sp.nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth.
|
||
<taxonomicName id="4C4D45F0D64D5163FD85F86DFC6AF81D" box="[620,946,1941,1964]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64D5163FD85F86DFC6AF81D" box="[620,946,1941,1964]" italics="true" pageId="3" pageNumber="65">Rhopalorhynchus gracillimus</emphasis>
|
||
</taxonomicName>
|
||
(Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23E73D64A5164FF2FFF60FE25FF69" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">Oviger: seg.2, 0.14; seg. 3, 0.12; seg. 4, 1.29; seg. 5, 0.14; seg. 6, 1.18; seg. 7, 0.24; seg. 8, 0.20; seg. 9, 0.20; seg. 10, 0.14; claw, 0.04.</paragraph>
|
||
<paragraph id="8BF23E73D64A5164FF2FFF1DFF0AFE97" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">Palp: seg. 3, 1.62; seg. 4, 0.05; seg. 5, 0.98; seg. 6, 0.12; seg. 7, 0.18; seg. 8, 0.18; seg. 9, 0.18; seg. 10, 0.15.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576DF8D64A5164FF2FFECAFDE1FEC2" pageId="4" pageNumber="66" type="etymology">
|
||
<paragraph id="8BF23E73D64A5164FF2FFECAFDE1FEC2" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">
|
||
<emphasis id="B939E261D64A5164FF2FFECAFE8BFEFD" bold="true" box="[198,339,306,332]" pageId="4" pageNumber="66">Etymology.</emphasis>
|
||
This species is named for its collector, Dr.
|
||
<taxonomicName id="4C4D45F0D64A5164FCBBFECAFC0CFEFD" box="[850,980,306,332]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="magdalena">Magdalena</taxonomicName>
|
||
Blazewicz-Paszkowycz, University of Łódź,
|
||
<collectingCountry id="F35A7EE3D64A5164FF0BFEA1FEEFFEC2" box="[226,311,345,371]" name="Poland" pageId="4" pageNumber="66">Poland</collectingCountry>
|
||
; a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576DF8D64A5165FF2FFE87FAD9FA4E" lastPageId="5" lastPageNumber="67" pageId="4" pageNumber="66" type="discussion">
|
||
<paragraph id="8BF23E73D64A5164FF2FFE87FC77FE71" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">
|
||
<emphasis id="B939E261D64A5164FF2FFE87FE98FE28" bold="true" box="[198,320,383,409]" pageId="4" pageNumber="66">Remarks.</emphasis>
|
||
The new species is another delicate, almost transparent form similar to
|
||
<taxonomicName id="4C4D45F0D64A5164FB04FE78FE43FE71" authority="Carpenter 1907: 100" authorityName="Carpenter" authorityPageNumber="100" authorityYear="1907" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64A5164FB04FE78FA43FE28" box="[1261,1435,384,409]" italics="true" pageId="4" pageNumber="66">R. gracillimus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDC4382D64A5164FF77FE5EFE4AFE71" author="Carpenter" box="[158,402,422,448]" pageId="4" pageNumber="68" refString="Carpenter, G. H. (1907) Pycnogonida. The Percy Sladen Trust Expedition to the Indian Ocean. Transactions of the Linnean Society London, series 2. (Zoology), 12, 95 - 101, pls. 12 - 13." type="journal article" year="1907">Carpenter 1907: 100</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and the smallest known species in the genus.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64A5164FF2FFE35FA44FD7E" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">
|
||
One of the third pair of legs of the
|
||
<typeStatus id="54F680D1D64A5164FD87FE35FD0CFE56" box="[622,724,461,487]" pageId="4" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
is missing, otherwise the specimen is complete. The dorsal process on the ocular tubercle is angled slightly more forward than those of the
|
||
<typeStatus id="54F680D1D64A5164FBDCFE0CFB7DFDBF" box="[1077,1189,500,526]" pageId="4" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
suggesting it may be bent. Most long setae on the legs and palps of all specimens are slightly reflexed distally. Based on the shape of the femora and the absence of genital pores, the smaller
|
||
<typeStatus id="54F680D1D64A5164FCBBFDB9FC1AFDEA" box="[850,962,577,603]" pageId="4" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
(about half the size of the
|
||
<typeStatus id="54F680D1D64A5164FAE9FDB9FAB1FDEA" box="[1280,1385,577,603]" pageId="4" pageNumber="66" type="holotype">holotype</typeStatus>
|
||
) are judged to be subadult males. The
|
||
<typeStatus id="54F680D1D64A5164FDDCFD90FD71FD33" box="[565,681,616,642]" pageId="4" pageNumber="66" type="paratype">paratypes</typeStatus>
|
||
exhibit the characteristic male femora which are less inflated distally than those of the female but otherwise conform to the species diagnosis generally. In a family where allometry is well documented, any variation in ratios and proportions of those specimens was not investigated.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64A5164FF2FFD24FB77FA8B" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">
|
||
Compared to the new species,
|
||
<taxonomicName id="4C4D45F0D64A5164FDC4FD24FC71FD44" box="[557,937,732,757]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="pedunculatum">
|
||
<emphasis id="B939E261D64A5164FDC4FD24FC71FD44" box="[557,937,732,757]" italics="true" pageId="4" pageNumber="66">Rhopalorhynchus pedunculatum</emphasis>
|
||
</taxonomicName>
|
||
has a proboscis stalk that is clearly longer than the inflated part (ca 1.5 times), the more widely spaced lateral processes (about 8 times) and a very short first trunk segment relative to segment 2 (25%). Geographically, the closest brevitarsal relatives are
|
||
<taxonomicName id="4C4D45F0D64A5164FA6BFCD1FF2FFCD8" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="clavipes">
|
||
<emphasis id="B939E261D64A5164FA6BFCD1FF2FFCD8" italics="true" pageId="4" pageNumber="66">R. clavipes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D45F0D64A5164FED8FCA8FE0CFCD8" box="[305,468,848,873]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="mortenseni">
|
||
<emphasis id="B939E261D64A5164FED8FCA8FE0CFCD8" box="[305,468,848,873]" italics="true" pageId="4" pageNumber="66">R. mortenseni</emphasis>
|
||
</taxonomicName>
|
||
, which are sympatric in the Coral Sea.
|
||
<taxonomicName id="4C4D45F0D64A5164FC4FFCA8FB05FCD8" box="[934,1245,848,873]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="clavipes">
|
||
<emphasis id="B939E261D64A5164FC4FFCA8FB05FCD8" box="[934,1245,848,873]" italics="true" pageId="4" pageNumber="66">Rhopalorhynchus clavipes</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by a longer tarsus and claw, a lower conical ocular tubercle, possession of a ‘minute’ or a ‘not very prominent’ dorsal tooth on the proboscis (
|
||
<bibRefCitation id="EFDC4382D64A5164FE16FC65FD1CFC06" author="Stock" box="[511,708,925,951]" pageId="4" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock, 1958:127</bibRefCitation>
|
||
) and a much shorter first trunk segment (41% of segment 2).
|
||
<bibRefCitation id="EFDC4382D64A5164FF7EFC3CFEE9FC6F" author="Clark" box="[151,305,964,990]" pageId="4" pageNumber="68" refString="Clark, W. C (1970) New Pycnogonida from Queensland. Transactions of the Royal Society of New Zealand, Biological Sciences, 12, 13 - 20." type="journal article" year="1970">Clark (1970)</bibRefCitation>
|
||
provisionally assigned a specimen collected from east of Jumpin Pin Bar, Qld, to
|
||
<taxonomicName id="4C4D45F0D64A5164FAF8FC3CFA4DFC6C" box="[1297,1429,964,990]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="clavipes">
|
||
<emphasis id="B939E261D64A5164FAF8FC3CFAFBFC6C" box="[1297,1315,964,989]" italics="true" pageId="4" pageNumber="66">R</emphasis>
|
||
.
|
||
<emphasis id="B939E261D64A5164FADAFC3CFA4DFC6C" box="[1331,1429,964,989]" italics="true" pageId="4" pageNumber="66">clavipes</emphasis>
|
||
</taxonomicName>
|
||
. Compared to the new species, Clark’s specimen is very much larger and differs in proportions of the proboscis and leg segments.
|
||
<taxonomicName id="4C4D45F0D64A5164FE9CFBE9FD0BFB9B" box="[373,723,1041,1066]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="mortenseni">
|
||
<emphasis id="B939E261D64A5164FE9CFBE9FD0BFB9B" box="[373,723,1041,1066]" italics="true" pageId="4" pageNumber="66">Rhopalorhynchus mortenseni</emphasis>
|
||
</taxonomicName>
|
||
was only briefly and incompletely described, but differs in the lower terminal process on the ocular tubercle, in the longer tarsus (longer than propodus), longer terminal claw (about 50% of propodus) and the more slender distal palp segments.
|
||
<taxonomicName id="4C4D45F0D64A5164FBEAFBA7FAF9FBC9" box="[1027,1313,1119,1144]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="filipes">
|
||
<emphasis id="B939E261D64A5164FBEAFBA7FAF9FBC9" box="[1027,1313,1119,1144]" italics="true" pageId="4" pageNumber="66">Rhopalorhynchus filipes</emphasis>
|
||
</taxonomicName>
|
||
is another much larger species (trunk length more than 2.5 times that of
|
||
<taxonomicName id="4C4D45F0D64A5164FC52FB7DFBB5FB2F" box="[955,1133,1157,1182]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="magdalena" status="sp. nov.">
|
||
<emphasis id="B939E261D64A5164FC52FB7DFBB5FB2F" box="[955,1133,1157,1182]" italics="true" pageId="4" pageNumber="66">R. magdalena</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A5F1AD64A5164FB9FFB7DFB04FB2E" box="[1142,1244,1157,1183]" pageId="4" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and is further distinguished by the very thin, almost filiform distal palp segments; the pointed ocular tubercle; a longer tarsus (subequal to the propodus), and longer claw (about half the propodus length).
|
||
<taxonomicName id="4C4D45F0D64A5164FB85FB2BFA44FB5D" box="[1132,1436,1235,1260]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="claudus">
|
||
<emphasis id="B939E261D64A5164FB85FB2BFA44FB5D" box="[1132,1436,1235,1260]" italics="true" pageId="4" pageNumber="66">Rhopalorhynchus claudus</emphasis>
|
||
</taxonomicName>
|
||
(based on a damaged specimen) can also be distinguished by the longer tarsus and terminal claw (about half the length of the propodus) and a longer neck region of the cephalon (66% of segment 2).
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64A5164FF2FFABEFB1DF90D" blockId="4.[151,1437,152,2034]" pageId="4" pageNumber="66">
|
||
The new species is morphologically closest to
|
||
<taxonomicName id="4C4D45F0D64A5164FD1BFABFFC41FAD1" box="[754,921,1351,1376]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64A5164FD1BFABFFC41FAD1" box="[754,921,1351,1376]" italics="true" pageId="4" pageNumber="66">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
in as much as this species is defined by its
|
||
<typeStatus id="54F680D1D64A5164FF7EFA95FEDEFA36" box="[151,262,1389,1415]" pageId="4" pageNumber="66" type="lectotype">lectotype</typeStatus>
|
||
. Both share the finger-shaped apical point on the ocular tubercle and similar proportions of the distal palp segments which were recorded by
|
||
<bibRefCitation id="EFDC4382D64A5164FD80FA6CFCEAFA1F" author="Carpenter" box="[617,818,1428,1454]" pageId="4" pageNumber="68" refString="Carpenter, G. H. (1907) Pycnogonida. The Percy Sladen Trust Expedition to the Indian Ocean. Transactions of the Linnean Society London, series 2. (Zoology), 12, 95 - 101, pls. 12 - 13." type="journal article" year="1907">Carpenter (1907)</bibRefCitation>
|
||
as 5:8:8:8:7.
|
||
<bibRefCitation id="EFDC4382D64A5164FC3BFA6CFBB3FA1F" author="Stock" box="[978,1131,1428,1454]" pageId="4" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock (1958)</bibRefCitation>
|
||
described palp segment 6 as being relatively long, measuring slightly shorter to slightly longer than segment 7 and did not comment on Carpenter’s description. Palp segment
|
||
<quantity id="4CB59396D64A5164FD8CFA19FD4BFA4A" box="[613,659,1505,1531]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="66" unit="in" value="6.0">6 in</quantity>
|
||
the new species is clearly shorter (by one-third) than segment 7. The new species otherwise differs in several respects: its overall size is only two-thirds that of
|
||
<taxonomicName id="4C4D45F0D64A5164FB1CF9F0FA44F990" box="[1269,1436,1544,1569]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64A5164FB1CF9F0FA44F990" box="[1269,1436,1544,1569]" italics="true" pageId="4" pageNumber="66">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
(trunk length
|
||
<quantity id="4CB59396D64A5164FEA9F9D6FE76F9F9" box="[320,430,1582,1608]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="4" pageNumber="66" unit="mm" value="2.35">2.35 mm</quantity>
|
||
v
|
||
<quantity id="4CB59396D64A5164FE26F9D6FD9AF9F9" box="[463,578,1582,1608]" metricMagnitude="-3" metricUnit="m" metricValue="3.71" pageId="4" pageNumber="66" unit="mm" value="3.71">3.71 mm</quantity>
|
||
); the tarsus and terminal claw are one-third or less than the propodus compared to the tarsus of
|
||
<taxonomicName id="4C4D45F0D64A5164FE26F9ADFDAFF9DF" box="[463,631,1621,1646]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64A5164FE26F9ADFDAFF9DF" box="[463,631,1621,1646]" italics="true" pageId="4" pageNumber="66">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
being half the length of the propodus and the terminal claw a little less than half, and in the shape and proportions of the proboscis (
|
||
<tableCitation id="C6CF0BC8D64A5164FC70F984FC29F927" box="[921,1009,1660,1686]" captionStart="TABLE 1" captionStartId="5.[151,239,460,484]" captionTargetBox="[151,1435,592,855]" captionTargetPageId="5" captionText="TABLE 1. Comparison of female specimens assigned to R. gracillimus (after Stock, 1958). Rhopalorhynchus magdalena holotype data is included for comparison." httpUri="http://table.plazi.org/id/DF326EFBD64B5165FF7EFE34FD8AFDB6" pageId="4" pageNumber="66" tableUuid="DF326EFBD64B5165FF7EFE34FD8AFDB6">Table 1</tableCitation>
|
||
). A distally expanded propodus has not been described or figured previously and may be an additional distinguishing character.
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64A5165FF2FF931FA5BFE2B" blockId="4.[151,1437,152,2034]" lastBlockId="5.[151,1437,152,410]" lastPageId="5" lastPageNumber="67" pageId="4" pageNumber="66">
|
||
Carpenter’s material of
|
||
<taxonomicName id="4C4D45F0D64A5164FE08F931FD51F953" box="[481,649,1737,1762]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64A5164FE08F931FD51F953" box="[481,649,1737,1762]" italics="true" pageId="4" pageNumber="66">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
consisted of eight specimens collected from three locations in the
|
||
<collectingCountry id="F35A7EE3D64A5164FF7EF908FE8BF8BB" box="[151,339,1776,1802]" name="Maldives" pageId="4" pageNumber="66">Maldive Islands</collectingCountry>
|
||
and a single male from the Saya de Malha Bank which he noted differed in several respects.
|
||
<bibRefCitation id="EFDC4382D64A5164FF7EF8EEFEE8F881" author="Stock" box="[151,304,1814,1840]" pageId="4" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock (1958)</bibRefCitation>
|
||
reviewed Carpenter’s material and assigned the Saya de Malha Bank specimen to
|
||
<taxonomicName id="4C4D45F0D64A5164FAEDF8EFFAAFF881" box="[1284,1399,1815,1840]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="lomani">
|
||
<emphasis id="B939E261D64A5164FAEDF8EFFAAFF881" box="[1284,1399,1815,1840]" italics="true" pageId="4" pageNumber="66">R. lomani</emphasis>
|
||
</taxonomicName>
|
||
, at the same time designating one female from South Nilandu as the
|
||
<typeStatus id="54F680D1D64A5164FC46F8C5FBC5F8E6" box="[943,1053,1853,1879]" pageId="4" pageNumber="66" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4D45F0D64A5164FBAEF8C5FB2DF8E7" box="[1095,1269,1853,1879]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64A5164FBAEF8C5FB81F8E7" box="[1095,1113,1853,1878]" italics="true" pageId="4" pageNumber="66">R</emphasis>
|
||
.
|
||
<emphasis id="B939E261D64A5164FB80F8C5FB2DF8E7" box="[1129,1269,1853,1878]" italics="true" pageId="4" pageNumber="66">gracillimus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFDC4382D64A5164FAEBF8C5FED4F8CF" author="Stock" pageId="4" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock (1958: figure 44)</bibRefCitation>
|
||
illustrated a “female
|
||
<typeStatus id="54F680D1D64A5164FDE4F89CFDB3F8CF" box="[525,619,1892,1918]" pageId="4" pageNumber="66" type="syntype">syntype</typeStatus>
|
||
” but his illustration of a leg (un-numbered) with moderately swollen distal part, is consistent with male morphology and similar to the male specimen figured by
|
||
<bibRefCitation id="EFDC4382D64A5164FB3DF872FEC1F87A" author="Carpenter" pageId="4" pageNumber="68" refString="Carpenter, G. H. (1907) Pycnogonida. The Percy Sladen Trust Expedition to the Indian Ocean. Transactions of the Linnean Society London, series 2. (Zoology), 12, 95 - 101, pls. 12 - 13." type="journal article" year="1907">Carpenter (1907: Figure 26)</bibRefCitation>
|
||
. It is not certain therefore, whether the illustrated
|
||
<typeStatus id="54F680D1D64A5164FC92F849FC00F87A" box="[891,984,1969,1995]" pageId="4" pageNumber="66" type="syntype">syntype</typeStatus>
|
||
is the male of the ‘one male and one female’ recorded by Carpenter from S. Nilandu or whether it is the female
|
||
<typeStatus id="54F680D1D64A5164FBC4F820FB44F843" box="[1069,1180,2008,2034]" pageId="4" pageNumber="66" type="lectotype">lectotype</typeStatus>
|
||
or another specimen altogether. Stock also assigned one female from Sapeh Bay, Sumbawa I. (Siboga stn 310) and
|
||
<specimenCount id="9D4BF5FAD64B5165FAECFF60FABDFF03" box="[1285,1381,152,178]" pageId="5" pageNumber="67" type="male">2 males</specimenCount>
|
||
, one female, from off Durban,
|
||
<collectingCountry id="F35A7EE3D64B5165FE22FF46FDBAFF69" box="[459,610,190,216]" name="South Africa" pageId="5" pageNumber="67">South Africa</collectingCountry>
|
||
to this species. In his key,
|
||
<bibRefCitation id="EFDC4382D64B5165FC4FFF46FB29FF69" author="Stock" box="[934,1265,190,216]" pageId="5" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958" yearSuffix="b">Stock (1958, couplet B b.2)</bibRefCitation>
|
||
, described the distal part of the proboscis as “narrowly produced, bent downward”, and used this character to distinguish
|
||
<taxonomicName id="4C4D45F0D64B5165FA6AFF1DFEC7FE94" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64B5165FA6AFF1DFEC7FE94" italics="true" pageId="5" pageNumber="67">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
from its congeners. However, Stock’s (1958: Figure 42) illustration of a “female
|
||
<typeStatus id="54F680D1D64B5165FB1AFEF4FA8BFE97" box="[1267,1363,268,294]" pageId="5" pageNumber="67" type="syntype">syntype</typeStatus>
|
||
” does not show a distal downturn, nor does he refer to a downturn in his species diagnosis (
|
||
<bibRefCitation id="EFDC4382D64B5165FB4DFECAFA56FEFD" author="Stock" box="[1188,1422,306,332]" pageId="5" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958" yearSuffix="p">Stock, 1958: p 129</bibRefCitation>
|
||
). Although not discussed by Carpenter, a distal downturn of the proboscis is evident in his Figure 26, so whether Stock relied solely on Carpenter’s figure or observed the downturn in other specimens, is not clear.
|
||
</paragraph>
|
||
<caption id="DF326EFBD64B5165FF7EFE34FD8AFDB6" ID-Table-UUID="DF326EFBD64B5165FF7EFE34FD8AFDB6" httpUri="http://table.plazi.org/id/DF326EFBD64B5165FF7EFE34FD8AFDB6" pageId="5" pageNumber="67" targetBox="[151,1435,592,855]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="8BF23E73D64B5165FF7EFE34FD8AFDB6" blockId="5.[151,1435,460,572]" pageId="5" pageNumber="67">
|
||
<emphasis id="B939E261D64B5165FF7EFE34FED0FE55" bold="true" box="[151,264,460,484]" pageId="5" pageNumber="67">TABLE 1.</emphasis>
|
||
Comparison of female specimens assigned to
|
||
<taxonomicName id="4C4D45F0D64B5165FD1DFE35FC53FE55" box="[756,907,461,484]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64B5165FD1DFE35FC53FE55" box="[756,907,461,484]" italics="true" pageId="5" pageNumber="67">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
(after Stock, 1958).
|
||
<taxonomicName id="4C4D45F0D64B5165FB8DFE35FA43FE55" box="[1124,1435,461,484]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="magdalena">
|
||
<emphasis id="B939E261D64B5165FB8DFE35FA43FE55" box="[1124,1435,461,484]" italics="true" pageId="5" pageNumber="67">Rhopalorhynchus magdalena</emphasis>
|
||
</taxonomicName>
|
||
holotype data is included for comparison.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF23E73D64B5165FDE6FDDAFB95FD8D" blockId="5.[151,1435,460,572]" box="[527,1101,546,572]" pageId="5" pageNumber="67">PROBOSCIS LEGS</paragraph>
|
||
<paragraph id="8BF23E73D64B5165FF76FDA8FAF4FC85" pageId="5" pageNumber="67">
|
||
<table id="F94DCCD3D64BAE9FFF7EFDA8FA43FCE6" box="[151,1435,592,855]" gridcols="5" gridrows="5" pageId="5" pageNumber="67">
|
||
<tr id="357D3C31D64BAE9FFF7EFDA8FA43FD38" box="[151,1435,592,649]" gridrow="0" pageId="5" pageNumber="67">
|
||
<th id="76AC554DD64BAE9FFF7EFDA8FEF3FD38" box="[151,299,592,649]" gridcol="0" gridrow="0" pageId="5" pageNumber="67">Location</th>
|
||
<th id="76AC554DD64BAE9FFEB4FDA8FD4EFD38" box="[349,662,592,649]" gridcol="1" gridrow="0" pageId="5" pageNumber="67">Length: Width Stalk length: Total length</th>
|
||
<th id="76AC554DD64BAE9FFD24FDA8FCD1FD38" box="[717,777,592,649]" gridcol="2" gridrow="0" pageId="5" pageNumber="67">Tooth</th>
|
||
<th id="76AC554DD64BAE9FFCBFFDA8FC16FD38" box="[854,974,592,649]" gridcol="3" gridrow="0" pageId="5" pageNumber="67">Tarsal ratio</th>
|
||
<th id="76AC554DD64BAE9FFBEDFDA8FA43FD38" box="[1028,1435,592,649]" gridcol="4" gridrow="0" pageId="5" pageNumber="67">Propodus: Tarsus Propodus: Claw</th>
|
||
</tr>
|
||
<tr id="357D3C31D64BAE9FFF7EFD65FA43FD05" box="[151,1435,669,692]" gridrow="1" pageId="5" pageNumber="67">
|
||
<th id="76AC554DD64BAE9FFF7EFD65FEF3FD05" box="[151,299,669,692]" gridcol="0" gridrow="1" pageId="5" pageNumber="67">Maldive Is.</th>
|
||
<td id="76AC554DD64BAE9FFEB4FD65FD4EFD05" box="[349,662,669,692]" gridcol="1" gridrow="1" pageId="5" pageNumber="67">3.9 36%</td>
|
||
<td id="76AC554DD64BAE9FFD24FD65FCD1FD05" box="[717,777,669,692]" gridcol="2" gridrow="1" pageId="5" pageNumber="67">27%</td>
|
||
<td id="76AC554DD64BAE9FFCBFFD65FC16FD05" box="[854,974,669,692]" gridcol="3" gridrow="1" pageId="5" pageNumber="67">66%</td>
|
||
<td id="76AC554DD64BAE9FFBEDFD65FA43FD05" box="[1028,1435,669,692]" gridcol="4" gridrow="1" pageId="5" pageNumber="67">63% 44%</td>
|
||
</tr>
|
||
<tr id="357D3C31D64BAE9FFF7EFD30FA43FD6E" box="[151,1435,712,735]" gridrow="2" pageId="5" pageNumber="67">
|
||
<th id="76AC554DD64BAE9FFF7EFD30FEF3FD6E" box="[151,299,712,735]" gridcol="0" gridrow="2" pageId="5" pageNumber="67">Sumbawa I.</th>
|
||
<td id="76AC554DD64BAE9FFEB4FD30FD4EFD6E" box="[349,662,712,735]" gridcol="1" gridrow="2" pageId="5" pageNumber="67">4.1 39%</td>
|
||
<td id="76AC554DD64BAE9FFD24FD30FCD1FD6E" box="[717,777,712,735]" gridcol="2" gridrow="2" pageId="5" pageNumber="67">24%</td>
|
||
<td id="76AC554DD64BAE9FFCBFFD30FC16FD6E" box="[854,974,712,735]" gridcol="3" gridrow="2" pageId="5" pageNumber="67">55%</td>
|
||
<td id="76AC554DD64BAE9FFBEDFD30FA43FD6E" box="[1028,1435,712,735]" gridcol="4" gridrow="2" pageId="5" pageNumber="67">83% 40%</td>
|
||
</tr>
|
||
<tr id="357D3C31D64BAE9FFF7EFD0AFA43FCB8" box="[151,1435,754,777]" gridrow="3" pageId="5" pageNumber="67">
|
||
<th id="76AC554DD64BAE9FFF7EFD0AFEF3FCB8" box="[151,299,754,777]" gridcol="0" gridrow="3" pageId="5" pageNumber="67">Durban</th>
|
||
<td id="76AC554DD64BAE9FFEB4FD0AFD4EFCB8" box="[349,662,754,777]" gridcol="1" gridrow="3" pageId="5" pageNumber="67">3.8 18%</td>
|
||
<td id="76AC554DD64BAE9FFD24FD0AFCD1FCB8" box="[717,777,754,777]" gridcol="2" gridrow="3" pageId="5" pageNumber="67">18%</td>
|
||
<td id="76AC554DD64BAE9FFCBFFD0AFC16FCB8" box="[854,974,754,777]" gridcol="3" gridrow="3" pageId="5" pageNumber="67">51%</td>
|
||
<td id="76AC554DD64BAE9FFBEDFD0AFA43FCB8" box="[1028,1435,754,777]" gridcol="4" gridrow="3" pageId="5" pageNumber="67">48% 40%</td>
|
||
</tr>
|
||
<tr id="357D3C31D64BAE9FFF7EFCE5FA43FCE6" box="[151,1435,797,855]" gridrow="4" pageId="5" pageNumber="67">
|
||
<th id="76AC554DD64BAE9FFF7EFCE5FEF3FCE6" box="[151,299,797,855]" gridcol="0" gridrow="4" pageId="5" pageNumber="67">R.magdelena Holotype</th>
|
||
<td id="76AC554DD64BAE9FFEB4FCE5FD4EFCE6" box="[349,662,797,855]" gridcol="1" gridrow="4" pageId="5" pageNumber="67">3.2 42%</td>
|
||
<td id="76AC554DD64BAE9FFD24FCE5FCD1FCE6" box="[717,777,797,855]" gridcol="2" gridrow="4" pageId="5" pageNumber="67">32%</td>
|
||
<td id="76AC554DD64BAE9FFCBFFCE5FC16FCE6" box="[854,974,797,855]" gridcol="3" gridrow="4" pageId="5" pageNumber="67">56%</td>
|
||
<td id="76AC554DD64BAE9FFBEDFCE5FA43FCE6" box="[1028,1435,797,855]" gridcol="4" gridrow="4" pageId="5" pageNumber="67">34–37% 25–32%</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF23E73D64B5165FF2FFC59FAD9FA4E" blockId="5.[151,1437,929,1535]" pageId="5" pageNumber="67">
|
||
The proboscis of the Durban specimen illustrated by
|
||
<bibRefCitation id="EFDC4382D64B5165FCD3FC59FB8CFC0A" author="Stock" box="[826,1108,929,955]" pageId="5" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock (1958: Figure 56)</bibRefCitation>
|
||
is closest to that figured by
|
||
<bibRefCitation id="EFDC4382D64B5165FF7EFC30FE35FC53" author="Carpenter" box="[151,493,968,994]" pageId="5" pageNumber="68" refString="Carpenter, G. H. (1907) Pycnogonida. The Percy Sladen Trust Expedition to the Indian Ocean. Transactions of the Linnean Society London, series 2. (Zoology), 12, 95 - 101, pls. 12 - 13." type="journal article" year="1907">Carpenter (1907: Figure 26)</bibRefCitation>
|
||
, but the specimen is otherwise distinguished by the significantly shorter basal stalk and the downward orientation of the entire inflated part. The inflated part of the proboscis in the Durban specimen is orientated at about 40° to the basal stalk (
|
||
<figureCitation id="137622F6D64B5165FCC9FBEDFC5FFB9E" box="[800,903,1045,1071]" captionStart="FIGURE 2" captionStartId="3.[151,259,1906,1930]" captionTargetBox="[216,1392,181,1880]" captionTargetId="figure@3.[189,1404,180,1892]" captionTargetPageId="3" captionText="FIGURE 2. Rhopalorhynchus magdalena sp. nov. female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth. Rhopalorhynchus gracillimus (Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis." httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="5" pageNumber="67">Figure 2</figureCitation>
|
||
H) compared to about 30° in the
|
||
<collectingCountry id="F35A7EE3D64B5165FAFDFBEDFAAFFB9E" box="[1300,1399,1045,1071]" name="Maldives" pageId="5" pageNumber="67">Maldive</collectingCountry>
|
||
Is.
|
||
<typeStatus id="54F680D1D64B5165FF7EFBC4FF22FBE7" box="[151,250,1084,1110]" pageId="5" pageNumber="67" type="syntype">syntype</typeStatus>
|
||
(
|
||
<figureCitation id="137622F6D64B5165FEE2FBC4FEA2FBE7" box="[267,378,1084,1110]" captionStart="FIGURE 2" captionStartId="3.[151,259,1906,1930]" captionTargetBox="[216,1392,181,1880]" captionTargetId="figure@3.[189,1404,180,1892]" captionTargetPageId="3" captionText="FIGURE 2. Rhopalorhynchus magdalena sp. nov. female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth. Rhopalorhynchus gracillimus (Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis." httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="5" pageNumber="67">Figure 2</figureCitation>
|
||
J) and 50° for the Sumbawa I. specimen (
|
||
<figureCitation id="137622F6D64B5165FC67FBC4FC25FBE7" box="[910,1021,1084,1110]" captionStart="FIGURE 2" captionStartId="3.[151,259,1906,1930]" captionTargetBox="[216,1392,181,1880]" captionTargetId="figure@3.[189,1404,180,1892]" captionTargetPageId="3" captionText="FIGURE 2. Rhopalorhynchus magdalena sp. nov. female holotype, A, oviger; B, oviger claw; C, palp; D, ocular tubercle; E, proboscis; F, proboscis tooth. Rhopalorhynchus gracillimus (Figures not to scale). Durban specimen, G, ocular tubercle, H, proboscis; Maldive lectotype, I, ocular tubercle, J, proboscis; Sumbawa I. specimen, K, ocular tubercle, L, proboscis." httpUri="https://zenodo.org/record/186601/files/figure.png" pageId="5" pageNumber="67">Figure 2</figureCitation>
|
||
L). When comparing the Durban specimens, Stock cited the length of the tarsus as a distinguishing character and regarded the long seventh palp joint and finger-shaped tip of eye tubercle as “somewhat aberrant”. The Sumbawa I. specimen also differs in the proportions of the distal leg segments and in the shape and proportions of the proboscis.
|
||
<bibRefCitation id="EFDC4382D64B5165FAADFB48FE89FB41" author="Arnaud" pageId="5" pageNumber="68" refString="Arnaud, F. & Child, A. C. (1988) The South African Museum's Meiring Naude cruises. Part 17. Pycnogonida. Annals of the South African Museum 98: 121 - 187." type="journal article" year="1988">Arnaud & Child (1988)</bibRefCitation>
|
||
contributed to the confusion by assigning a longitarsal specimen from East London (
|
||
<collectingCountry id="F35A7EE3D64B5165FABCFB2EFF3EFAA6" name="South Africa" pageId="5" pageNumber="67">South Africa</collectingCountry>
|
||
) to this species. Their observation that the specimen had abnormally short distal palp segments is a further indication that it represented a distinct species. It is puzzling that Stock attributed the diverse and sometimes aberrant morphology evident in his
|
||
<taxonomicName id="4C4D45F0D64B5165FD18FAB3FC17FAD5" box="[753,975,1355,1380]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B939E261D64B5165FD18FAB3FC17FAD5" box="[753,975,1355,1380]" italics="true" pageId="5" pageNumber="67">Rhopalorhynchus</emphasis>
|
||
</taxonomicName>
|
||
material to intraspecific variability, particularly in light of his observation that “speciation through geographic isolation is of outstanding importance in marine organisms such as the Pycnogonida” (
|
||
<bibRefCitation id="EFDC4382D64B5165FC79FA60FBB5FA03" author="Stock" box="[912,1133,1432,1458]" pageId="5" pageNumber="68" refString="Stock, J. H. (1958) The pycnogonid genus Rhopalorhynchus. Tijdschrift voor Entomologie, 101, 113 - 137." type="journal article" year="1958">Stock, 1958: 113</bibRefCitation>
|
||
). Differences in general morphology and variation in ratios (
|
||
<tableCitation id="C6CF0BC8D64B5165FDD3FA46FD48FA69" box="[570,656,1470,1496]" captionStart="TABLE 1" captionStartId="5.[151,239,460,484]" captionTargetBox="[151,1435,592,855]" captionTargetPageId="5" captionText="TABLE 1. Comparison of female specimens assigned to R. gracillimus (after Stock, 1958). Rhopalorhynchus magdalena holotype data is included for comparison." httpUri="http://table.plazi.org/id/DF326EFBD64B5165FF7EFE34FD8AFDB6" pageId="5" pageNumber="67" tableUuid="DF326EFBD64B5165FF7EFE34FD8AFDB6">Table 1</tableCitation>
|
||
) suggest that the wide variability attributed to
|
||
<taxonomicName id="4C4D45F0D64B5165FB59FA47FA8EFA69" box="[1200,1366,1471,1496]" class="Pycnogonida" family="Colossendeidae" genus="Rhopalorhynchus" higherTaxonomySource="CoL" kingdom="Animalia" order="Pantopoda" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="gracillimus">
|
||
<emphasis id="B939E261D64B5165FB59FA47FA8EFA69" box="[1200,1366,1471,1496]" italics="true" pageId="5" pageNumber="67">R. gracillimus</emphasis>
|
||
</taxonomicName>
|
||
is not justified and that the specimens from Sumbawa I. and
|
||
<collectingCountry id="F35A7EE3D64B5165FCFCFA1DFC73FA4E" box="[789,939,1509,1535]" name="South Africa" pageId="5" pageNumber="67">South Africa</collectingCountry>
|
||
may belong to other species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |