treatments-xml/data/03/E4/87/03E487E3CA69C51FFC9D53C6CDF5C2AC.xml

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<mods:title id="4E299A511D18C4938D1373562E31B37F">New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<treatment id="03E487E3CA69C51FFC9D53C6CDF5C2AC" ID-DOI="http://doi.org/10.5281/zenodo.5878975" ID-GBIF-Taxon="192433636" ID-Zenodo-Dep="5878975" LSID="urn:lsid:plazi:treatment:03E487E3CA69C51FFC9D53C6CDF5C2AC" httpUri="http://treatment.plazi.org/id/03E487E3CA69C51FFC9D53C6CDF5C2AC" lastPageId="6" lastPageNumber="249" pageId="3" pageNumber="246">
<subSubSection id="C357657ECA69C51AFC9D53C6CD68C0A3" pageId="3" pageNumber="246" type="nomenclature">
<paragraph id="8BF236F5CA69C51AFC9D53C6CD68C0A3" blockId="3.[831,1393,947,1039]" pageId="3" pageNumber="246">
<taxonomicName id="4C4D4D76CA69C51AFC9D53C6CD68C0A3" ID-CoL="37TD6" authority="MARUYAMA &amp; KISHIMOTO, 2002" authorityName="MARUYAMA &amp; KISHIMOTO" authorityYear="2002" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="246" phylum="Arthropoda" rank="species" species="watanabei" subGenus="Drusilla">
<emphasis id="B939EAE7CA69C51AFC9D53C6CD96C083" box="[831,908,947,974]" italics="true" pageId="3" pageNumber="246">Drusilla</emphasis>
(
<emphasis id="B939EAE7CA69C51AFC3853C6CDFDC083" box="[922,999,947,974]" italics="true" pageId="3" pageNumber="246">Drusilla</emphasis>
) 
<emphasis id="B939EAE7CA69C51AFC5753C6CA79C083" box="[1013,1123,947,974]" italics="true" pageId="3" pageNumber="246">watanabei</emphasis>
MARUYAMA &amp; KISHIMOTO, 2002
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C357657ECA69C51AFC9D538CCD84C742" box="[831,926,1017,1039]" pageId="3" pageNumber="246" type="discussion">
<paragraph id="8BF236F5CA69C51AFC9D538CCD84C742" blockId="3.[831,1393,947,1039]" box="[831,926,1017,1039]" pageId="3" pageNumber="246">
(
<figureCitation id="13762A70CA69C51AFCE6538CCD82C742" box="[836,920,1017,1039]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="3" pageNumber="246">Figs 1–13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C357657ECA69C51AFCB8545CCDABC78C" pageId="3" pageNumber="246" type="materials_examined">
<paragraph id="8BF236F5CA69C51AFCB8545CCDABC78C" blockId="3.[793,1399,1065,1217]" pageId="3" pageNumber="246">
<emphasis id="B939EAE7CA69C51AFCB8545CCDF0C70C" bold="true" box="[794,1002,1065,1089]" pageId="3" pageNumber="246">Material examined</emphasis>
: 
<materialsCitation id="3B253CA8CA69C51AFC5B545CCDB4C78C" ID-GBIF-Occurrence="3447238316" collectingDate="2014-08-14" collectorName="W Dongchuan &amp; Assing" country="China" county="Sedan Snow Mountain Scenic Resort" elevation="2620" latitude="26.1" location="Sedan Snow Mountain Scenic Resort" longLatPrecision="1243" longitude="102.916664" municipality="Sedan Snow Mountain" pageId="3" pageNumber="246" specimenCount="1" stateProvince="Yunnan">
<emphasis id="B939EAE7CA69C51AFC5B545CCA26C70C" bold="true" box="[1017,1084,1065,1089]" pageId="3" pageNumber="246">
<collectingCountry id="F35A7665CA69C51AFC5B545CCA26C70C" box="[1017,1084,1065,1089]" name="China" pageId="3" pageNumber="246">China</collectingCountry>
</emphasis>
: 
<emphasis id="B939EAE7CA69C51AFBEE545CCAB8C70C" bold="true" box="[1100,1186,1065,1089]" pageId="3" pageNumber="246">
<collectingRegion id="4989F817CA69C51AFBEE545CCAB8C70C" box="[1100,1186,1065,1089]" country="China" name="Yunnan" pageId="3" pageNumber="246">Yunnan</collectingRegion>
</emphasis>
: 3 ♂♂, 1 ♀, mountains 
<collectorName id="26B85323CA69C51AFCFF543FCA1DC72C" box="[861,1031,1097,1121]" pageId="3" pageNumber="246">W Dongchuan</collectorName>
, 
<collectingCounty id="62934E79CA69C51AFBB9543CCD78C7CC" pageId="3" pageNumber="246">Sedan Snow Mountain Scenic Resort</collectingCounty>
, 
<geoCoordinate id="EE795032CA69C51AFCC9541CCDDBC7CC" box="[875,961,1129,1153]" degrees="26" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="246" precision="925" value="26.1">26°06'N</geoCoordinate>
, 
<geoCoordinate id="EE795032CA69C51AFC69541CCA30C7CC" box="[971,1066,1129,1153]" degrees="102" direction="east" minutes="55" orientation="longitude" pageId="3" pageNumber="246" precision="925" value="102.916664">102°55'E</geoCoordinate>
, 
<quantity id="4CB59B10CA69C51AFB91541CCA9FC7CC" box="[1075,1157,1129,1153]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="3" pageNumber="246" unit="m" value="2620.0">
<elevation id="0060D1C6CA69C51AFB91541CCA9FC7CC" box="[1075,1157,1129,1153]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="3" pageNumber="246" unit="m" value="2620.0">2620 m</elevation>
</quantity>
, secondary pine forest, litter, moss, and roots of herbs sifted, 
<date id="FFF31035CA69C51AFB1954FCCB5EC7EC" box="[1211,1348,1161,1185]" pageId="3" pageNumber="246" value="2014-08-14">
<collectingDate id="EFB7E9DDCA69C51AFB1954FCCB5EC7EC" box="[1211,1348,1161,1185]" pageId="3" pageNumber="246" value="2014-08-14">14.VIII.2014</collectingDate>
</date>
, leg. 
<collectorName id="26B85323CA69C51AFCB854DCCD7BC78C" box="[794,865,1193,1217]" pageId="3" pageNumber="246">Assing</collectorName>
(cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357657ECA69C51AFCB8549CCDDEC6EC" pageId="3" pageNumber="246" type="discussion">
<paragraph id="8BF236F5CA69C51AFCB8549CCDDEC6EC" blockId="3.[794,1400,1257,1441]" pageId="3" pageNumber="246">
<emphasis id="B939EAE7CA69C51AFCB8549CCD93C64C" bold="true" box="[794,905,1257,1281]" pageId="3" pageNumber="246">Comment</emphasis>
: The original description is based on 
<specimenCount id="9D4BFD7CCA69C51AFA8B549CCB6CC64F" box="[1321,1398,1257,1282]" pageId="3" pageNumber="246" type="generic" typeStatus="type">16 type</specimenCount>
specimens from two localities in 
<collectingRegion id="4989F817CA69C51AFBD5557CCB2CC66C" box="[1143,1334,1289,1313]" isEnumeration="true" pageId="3" pageNumber="246">Hubei and Hunan</collectingRegion>
provinces (
<bibRefCitation id="EFDC4B04CA69C51AFCC7555CCAC1C60C" author="MARUYAMA, M. &amp; KISHIMOTO, T." box="[869,1243,1321,1345]" pageId="3" pageNumber="246" pagination="227 - 232" refId="ref14664" refString="MARUYAMA, M. &amp; KISHIMOTO, T. 2002 a: Myrmecophilous species of Drusilla (Coleoptera, Staphylinidae, Aleocharinae) associated with Lasius (Dendrolasius) spp. (Hymenoptera, Formicidae, Formicinae) from China. Part 1. - Special Bulletin of the Japanese Society of Coleopterology 5: 227 - 232." type="journal article" year="2002">MARUYAMA &amp; KISHIMOTO 2002a</bibRefCitation>
). The authors did not find (and illustrate) a spermatheca, although the 
<typeStatus id="54F68857CA69C51AFCB9551FCD53C6CF" box="[795,841,1386,1410]" pageId="3" pageNumber="246">type</typeStatus>
series includes 
<specimenCount id="9D4BFD7CCA69C51AFC52551FCA9FC6CC" box="[1008,1157,1385,1409]" pageId="3" pageNumber="246" type="female">seven females</specimenCount>
. A full redescription is provided below.
</paragraph>
</subSubSection>
<subSubSection id="C357657ECA69C51FFCB855BCCCA9CB6C" lastPageId="6" lastPageNumber="249" pageId="3" pageNumber="246" type="description">
<paragraph id="8BF236F5CA69C51AFCB855BCCA8CC44C" blockId="3.[793,1400,1481,2081]" pageId="3" pageNumber="246">
<emphasis id="B939EAE7CA69C51AFCB855BCCD86C6AC" bold="true" box="[794,924,1481,1505]" pageId="3" pageNumber="246">Description</emphasis>
: Relatively large species; body length 7.0– 
<quantity id="4CB59B10CA69C51AFCE5559CCD81C54C" box="[839,923,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="246" unit="mm" value="7.8">7.8 mm</quantity>
; length of forebody 2.6–3.0 mm. Coloration: body black, with the posterior margins of the abdominal segments paler; legs with the tibiae and tarsi yellowish to dark-yellowish, the profemora blackish, the mesofemora blackish with the basal third yellowish, and the metafemora yellowish in basal half and infuscate in apical half; antennae reddish-yellow, with antennomere I weakly to distinctly infuscate; maxillary palpi dark-brown, with the terminal palpomere reddish-yellow.
</paragraph>
<paragraph id="8BF236F5CA69C51CFCB8577CCC56C26C" blockId="3.[793,1400,1481,2081]" lastBlockId="5.[151,756,233,385]" lastPageId="5" lastPageNumber="248" pageId="3" pageNumber="246">
Head (
<figureCitation id="13762A70CA69C51AFCC3577CCDA0C46C" box="[865,954,1801,1825]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="3" pageNumber="246">Figs 1–2</figureCitation>
) with sexually dimorphic punctation and with fine and shallow microreticulation. Eyes distinctly shorter than distance from posterior margin of eye to posterior constriction of head. Antenna (
<figureCitation id="13762A70CA69C51AFB08571CCAFFC4CC" box="[1194,1253,1897,1921]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="3" pageNumber="246">Fig. 3</figureCitation>
) 
<quantity id="4CB59B10CA69C51AFB58571CCB6DC4CC" box="[1274,1399,1897,1921]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="3" pageNumber="246" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6–2.9 mm</quantity>
long and slender; all antennomeres at least weakly oblong. Pronotum (
<figureCitation id="13762A70CA69C51AFC3857DCCDECC48C" box="[922,1014,1961,1985]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="3" pageNumber="246">Figs 1–2</figureCitation>
) 1.00–1.05 times as long as broad and 1.17–1.18 times as broad as head, broadest in anterior half; posterior angles weakly marked; near posterior margin with pronounced transverse impression; other impressions and punctation sexually dimorphic; interstices without microsculpture and glossy.
</paragraph>
<caption id="DF32667DCA6EC51DFF1F5454CD57C73F" ID-DOI="http://doi.org/10.5281/zenodo.5887109" ID-Zenodo-Dep="5887109" httpUri="https://zenodo.org/record/5887109/files/figure.png" pageId="4" pageNumber="247" startId="4.[189,234,1057,1078]" targetBox="[189,1436,232,1050]" targetPageId="4">
<paragraph id="8BF236F5CA6EC51DFF1F5454CD57C73F" blockId="4.[189,1437,1057,1138]" pageId="4" pageNumber="247">
<emphasis id="B939EAE7CA6EC51DFF1F5454CEE6C77B" bold="true" box="[189,252,1057,1078]" pageId="4" pageNumber="247">Map 1</emphasis>
: Distributions of 
<taxonomicName id="4C4D4D76CA6EC51DFE0B5454CC25C77A" box="[425,575,1057,1079]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B939EAE7CA6EC51DFE0B5454CFECC77B" box="[425,502,1057,1078]" italics="true" pageId="4" pageNumber="247">Drusilla</emphasis>
species
</taxonomicName>
recorded from China: 
<taxonomicName id="4C4D4D76CA6EC51DFC825457CD8BC77B" authorityName="Assing" authorityYear="2015" box="[800,913,1057,1079]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="flagellata">
<emphasis id="B939EAE7CA6EC51DFC825457CD8BC77B" box="[800,913,1057,1079]" italics="true" pageId="4" pageNumber="247">D. flagellata</emphasis>
</taxonomicName>
(black stars); 
<taxonomicName id="4C4D4D76CA6EC51DFBB95457CA78C77B" authorityName="Assing" authorityYear="2015" box="[1051,1122,1057,1079]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="gilva">
<emphasis id="B939EAE7CA6EC51DFBB95457CA78C77B" box="[1051,1122,1057,1079]" italics="true" pageId="4" pageNumber="247">D. gilva</emphasis>
</taxonomicName>
(white square); 
<taxonomicName id="4C4D4D76CA6EC51DFAA35457CB40C77A" authorityName="MARUYAMA &amp; KISHIMOTO" authorityYear="2002" box="[1281,1370,1058,1079]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="imurai">
<emphasis id="B939EAE7CA6EC51DFAA35457CB40C77A" box="[1281,1370,1058,1079]" italics="true" pageId="4" pageNumber="247">D. imurai</emphasis>
</taxonomicName>
(black triangle); 
<taxonomicName id="4C4D4D76CA6EC51DFEBE544ACF85C71E" authorityName="MARUYAMA &amp; KISHIMOTO" authorityYear="2002" box="[284,415,1086,1108]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="dendrolasii">
<emphasis id="B939EAE7CA6EC51DFEBE544ACF85C71E" box="[284,415,1086,1108]" italics="true" pageId="4" pageNumber="247">D. dendrolasii</emphasis>
</taxonomicName>
(white diamond); 
<taxonomicName id="4C4D4D76CA6EC51DFDF5544ACCF4C71E" authorityName="PACE" authorityYear="1998" box="[599,750,1086,1108]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="zhejiangensis">
<emphasis id="B939EAE7CA6EC51DFDF5544ACCF4C71E" box="[599,750,1086,1108]" italics="true" pageId="4" pageNumber="247">D. zhejiangensis</emphasis>
</taxonomicName>
(white star); 
<taxonomicName id="4C4D4D76CA6EC51DFCD3544ACA22C71E" authorityName="PACE" authorityYear="2012" box="[881,1080,1086,1108]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="cenwanglaomontis">
<emphasis id="B939EAE7CA6EC51DFCD3544ACA22C71E" box="[881,1080,1086,1108]" italics="true" pageId="4" pageNumber="247">D. cenwanglaomontis</emphasis>
</taxonomicName>
(white circle); 
<taxonomicName id="4C4D4D76CA6EC51DFB6F544ACB46C71E" authorityName="PACE" authorityYear="2012" box="[1229,1372,1086,1108]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="damingensis">
<emphasis id="B939EAE7CA6EC51DFB6F544ACB46C71E" box="[1229,1372,1086,1108]" italics="true" pageId="4" pageNumber="247">D. damingensis</emphasis>
</taxonomicName>
(black circle); 
<taxonomicName id="4C4D4D76CA6EC51DFEA65429CF63C73C" authorityName="PACE" authorityYear="1998" box="[260,377,1116,1137]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="gibberella">
<emphasis id="B939EAE7CA6EC51DFEA65429CF63C73C" box="[260,377,1116,1137]" italics="true" pageId="4" pageNumber="247">D. gibberella</emphasis>
</taxonomicName>
(black diamonds); 
<taxonomicName id="4C4D4D76CA6EC51DFD905429CCB6C73C" baseAuthorityName="BERNHAUER" baseAuthorityYear="1915" box="[562,684,1116,1137]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="philippina">
<emphasis id="B939EAE7CA6EC51DFD905429CCB6C73C" box="[562,684,1116,1137]" italics="true" pageId="4" pageNumber="247">D. philippina</emphasis>
</taxonomicName>
(white triangle).
</paragraph>
</caption>
<caption id="DF32667DCA6EC51DFF1F5798CD1ECB52" ID-DOI="http://doi.org/10.5281/zenodo.5887113" ID-Zenodo-Dep="5887113" httpUri="https://zenodo.org/record/5887113/files/figure.png" pageId="4" pageNumber="247" startId="4.[189,234,2029,2050]" targetBox="[189,1436,1203,2021]" targetPageId="4">
<paragraph id="8BF236F5CA6EC51DFF1F5798CD1ECB52" blockId="4.[189,1437,2028,2080]" pageId="4" pageNumber="247">
<emphasis id="B939EAE7CA6EC51DFF1F5798CEE4CB4C" bold="true" box="[189,254,2028,2050]" pageId="4" pageNumber="247">Map 2</emphasis>
: Distributions of 
<taxonomicName id="4C4D4D76CA6EC51DFE0D5799CC52CB4F" box="[431,584,2028,2050]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B939EAE7CA6EC51DFE0D5799CFE6CB4C" box="[431,508,2028,2049]" italics="true" pageId="4" pageNumber="247">Drusilla</emphasis>
species
</taxonomicName>
recorded from China: 
<taxonomicName id="4C4D4D76CA6EC51DFC925798CD9CCB4C" authorityName="Assing" authorityYear="2015" box="[816,902,2028,2050]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="palata">
<emphasis id="B939EAE7CA6EC51DFC925798CD9CCB4C" box="[816,902,2028,2050]" italics="true" pageId="4" pageNumber="247">D. palata</emphasis>
</taxonomicName>
(white stars); 
<taxonomicName id="4C4D4D76CA6EC51DFBB55798CABDCB4F" authorityName="PACE" authorityYear="1993" box="[1047,1191,2029,2050]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B939EAE7CA6EC51DFBB55798CABDCB4F" box="[1047,1191,2029,2050]" italics="true" pageId="4" pageNumber="247">D. yunnanensis</emphasis>
</taxonomicName>
(black and grey circles); 
<taxonomicName id="4C4D4D76CA6EC51DFF1F587ECF23CB52" authorityName="MARUYAMA &amp; KISHIMOTO" authorityYear="2002" box="[189,313,2058,2080]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="watanabei">
<emphasis id="B939EAE7CA6EC51DFF1F587ECF23CB52" box="[189,313,2058,2080]" italics="true" pageId="4" pageNumber="247">D. watanabei</emphasis>
</taxonomicName>
(white and grey circles); 
<taxonomicName id="4C4D4D76CA6EC51DFD88587ECC96CB52" authorityName="PACE" authorityYear="2012" box="[554,652,2058,2080]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="247" phylum="Arthropoda" rank="species" species="sinicola">
<emphasis id="B939EAE7CA6EC51DFD88587ECC96CB52" box="[554,652,2058,2080]" italics="true" pageId="4" pageNumber="247">D. sinicola</emphasis>
</taxonomicName>
(black star).
</paragraph>
</caption>
<paragraph id="8BF236F5CA6FC51CFF35515CCA34C22C" blockId="5.[151,756,233,385]" lastBlockId="5.[794,1399,233,353]" pageId="5" pageNumber="248">
Elytra (
<figureCitation id="13762A70CA6FC51CFF52515CCF57C20C" box="[240,333,297,321]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="5" pageNumber="248">Figs 1–2</figureCitation>
) short, approximately 0.65 times as long as pronotum; humeral angles moderately marked; punctation sexually dimorphic; interstices without microsculpture. Hind wings reduced to short stubs extending slightly beyond posterior margin of elytra. Metatarsomere I elongate, approximately as long as the combined length of II–IV.
</paragraph>
<caption id="DF32667DCA6FC51CFF3557A5CA80CB6D" ID-DOI="http://doi.org/10.5281/zenodo.5887117" ID-Zenodo-Dep="5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="5" pageNumber="248" startId="5.[151,191,2000,2022]" targetBox="[151,1398,416,1977]" targetPageId="5">
<paragraph id="8BF236F5CA6FC51CFF3557A5CA80CB6D" blockId="5.[151,1399,2000,2081]" pageId="5" pageNumber="248">
<emphasis id="B939EAE7CA6FC51CFF3557A5CEEFC4A8" bold="true" box="[151,245,2000,2022]" pageId="5" pageNumber="248">Figs 1–13</emphasis>
: 
<taxonomicName id="4C4D4D76CA6FC51CFEA257A5CFAFC4A8" authorityName="MARUYAMA &amp; KISHIMOTO" authorityYear="2002" box="[256,437,2000,2021]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="248" phylum="Arthropoda" rank="species" species="watanabei">
<emphasis id="B939EAE7CA6FC51CFEA257A5CFAFC4A8" box="[256,437,2000,2021]" italics="true" pageId="5" pageNumber="248">Drusilla watanabei</emphasis>
</taxonomicName>
: male forebody (
<emphasis id="B939EAE7CA6FC51CFDFB57A5CC7FC4A8" bold="true" box="[601,613,2000,2021]" pageId="5" pageNumber="248">1</emphasis>
); female forebody (
<emphasis id="B939EAE7CA6FC51CFC8057A5CD34C4A8" bold="true" box="[802,814,2000,2021]" pageId="5" pageNumber="248">2</emphasis>
); antenna (
<emphasis id="B939EAE7CA6FC51CFC3F57A5CDB3C4A8" bold="true" box="[925,937,2000,2021]" pageId="5" pageNumber="248">3</emphasis>
); male abdomen (
<emphasis id="B939EAE7CA6FC51CFBF857A5CA7CC4A8" bold="true" box="[1114,1126,2000,2021]" pageId="5" pageNumber="248">4</emphasis>
); male tergite VIII (
<emphasis id="B939EAE7CA6FC51CFA8A57A5CB2EC4A8" bold="true" box="[1320,1332,2000,2021]" pageId="5" pageNumber="248">5</emphasis>
); male sternite VIII (
<emphasis id="B939EAE7CA6FC51CFEB95798CF3DCB4F" bold="true" box="[283,295,2029,2050]" pageId="5" pageNumber="248">6</emphasis>
); median lobe of aedeagus in lateral and in ventral view (
<emphasis id="B939EAE7CA6FC51CFCE75798CD72CB4F" bold="true" box="[837,872,2029,2050]" pageId="5" pageNumber="248">7–8</emphasis>
); paramere (
<emphasis id="B939EAE7CA6FC51CFC465798CDEACB4F" bold="true" box="[996,1008,2029,2050]" pageId="5" pageNumber="248">9</emphasis>
); female tergite VIII (
<emphasis id="B939EAE7CA6FC51CFB625798CAC2CB4F" bold="true" box="[1216,1240,2029,2050]" pageId="5" pageNumber="248">10</emphasis>
); female sternite VIII (
<emphasis id="B939EAE7CA6FC51CFF6C587ECEFCCB6D" bold="true" box="[206,230,2059,2080]" pageId="5" pageNumber="248">11</emphasis>
); spermatheca in different aspects (
<emphasis id="B939EAE7CA6FC51CFD9E587ECC6DCB6D" bold="true" box="[572,631,2059,2080]" pageId="5" pageNumber="248">12–13</emphasis>
). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF236F5CA6CC51FFF1F509CCF8EC2EC" blockId="6.[187,796,233,1857]" pageId="6" pageNumber="249">
Abdomen (
<figureCitation id="13762A70CA6CC51FFEEA509CCF9FC24C" box="[328,389,233,257]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 4</figureCitation>
) distinctly broader than elytra; tergites III–VIII with sexual dimorphism; punctation sexually dimorphic; anterior impressions of tergites III– VI impunctate; tergites with very weak, nearly obsolete transverse microsculpture; posterior margin of tergite VII with palisade fringe.
</paragraph>
<paragraph id="8BF236F5CA6CC51FFF1F51DFCCBAC64C" blockId="6.[187,796,233,1857]" pageId="6" pageNumber="249">
♂: lateral portions of dorsal surface of head with dense and rather coarse punctation, with the interstices narrower than diameter of punctures; frons, posterior portion, and the narrow midline of the head impunctate or nearly so (
<figureCitation id="13762A70CA6CC51FFF66525CCEE7C10C" box="[196,253,553,577]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 1</figureCitation>
); pronotum in posterior half with pronounced longitudinal elevation on either side, between these elevations with broadly flattened impression; median portion of pronotum somewhat shiny and with moderately dense and moderately coarse punctation, with the interstices as broad as, or broader than, diameter of punctures; lateral portions of pronotum practically matt, with very dense and coarsely granulose punctation, with the interstices narrower than diameter of punctures (
<figureCitation id="13762A70CA6CC51FFDF5535CCC8AC00C" box="[599,656,809,833]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 1</figureCitation>
); elytra with dense and coarsely granulose punctation (
<figureCitation id="13762A70CA6CC51FFDD1533CCCB0C02C" box="[627,682,841,865]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 1</figureCitation>
); abdominal tergite III (
<figureCitation id="13762A70CA6CC51FFEF7531CCF88C0CC" box="[341,402,873,897]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 4</figureCitation>
) with a pronounced median tubercle of somewhat variable shape and with scattered fine punctures; tergites IV–VI each with a median pair of distinct granules near posterior margin, impunctate (or nearly so), except for some punctures at posterior margins; tergite VII with a pronounced median granule and numerous additional smaller granules in posterior half (
<figureCitation id="13762A70CA6CC51FFD72545CCD17C70C" box="[720,781,1065,1089]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 4</figureCitation>
); tergite VIII (
<figureCitation id="13762A70CA6CC51FFE9D543CCF60C72C" box="[319,378,1097,1121]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 5</figureCitation>
) with densely granulose punctation and with distinct microreticulation, posterior margin broadly concavely excavate; sternite VIII (
<figureCitation id="13762A70CA6CC51FFDBA54FCCC4FC7EC" box="[536,597,1161,1185]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 6</figureCitation>
) weakly transverse and with nearly truncate posterior margin; median lobe of aedeagus large, 1.0– 
<quantity id="4CB59B10CA6CC51FFE3754BCCFF9C7AC" box="[405,483,1225,1249]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="249" unit="mm" value="1.1">1.1 mm</quantity>
long, shaped as in 
<figureCitation id="13762A70CA6CC51FFD1F54BCCD0CC7AC" box="[701,790,1225,1249]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Figs 7–8</figureCitation>
; paramere 
<quantity id="4CB59B10CA6CC51FFE87549CCF6FC64C" box="[293,373,1257,1281]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="249" unit="mm" value="0.9">0.9 mm</quantity>
long and shaped as in 
<figureCitation id="13762A70CA6CC51FFDC2549CCC86C64C" box="[608,668,1257,1281]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 9</figureCitation>
.
</paragraph>
<paragraph id="8BF236F5CA6CC51FFF1F557FCFDEC40C" blockId="6.[187,796,233,1857]" pageId="6" pageNumber="249">
♀: head (
<figureCitation id="13762A70CA6CC51FFE85557CCF7CC66C" box="[295,358,1289,1313]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 2</figureCitation>
) with sparse and very fine punctation, interstices much broader than diameter of punctures; pronotum (
<figureCitation id="13762A70CA6CC51FFE9F553CCF66C62C" box="[317,380,1353,1377]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 2</figureCitation>
) with weakly pronounced elevations in posterior half; punctation moderately fine, moderately dense, and non-granulose; elytra (
<figureCitation id="13762A70CA6CC51FFD3155FCCCC8C6EC" box="[659,722,1417,1441]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 2</figureCitation>
) with weakly granulose puncation; abdominal tergites III–V with fine and sparse punctation; tergite VI impunctate except for a median pair of setiferous punctures bearing long dark setae and additional punctures at posterior margin; tergite VII with a median pair of setiferous punctures bearing long dark setae and with additional fine and sparse punctation in posterior half; posterior margin of tergite VIII somewhat truncate, only indistinctly concave in the middle (
<figureCitation id="13762A70CA6CC51FFD8856DCCC62C58C" box="[554,632,1705,1729]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 10</figureCitation>
); sternite VIII (
<figureCitation id="13762A70CA6CC51FFF6756BCCF0BC5AC" box="[197,273,1737,1761]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Fig. 11</figureCitation>
) distinctly shorter and more transverse than that of male, posterior margin truncate; spermatheca very small in relation to body size, 
<quantity id="4CB59B10CA6CC51FFDEB577CCCB3C46C" box="[585,681,1801,1825]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="249" unit="mm" value="0.25">0.25 mm</quantity>
long, and shaped as in 
<figureCitation id="13762A70CA6CC51FFEEA575CCFDAC40C" box="[328,448,1833,1857]" captionStart="Figs 1–13" captionStartId="5.[151,191,2000,2022]" captionTargetBox="[151,1398,416,1977]" captionTargetId="figure-189@5.[151,1398,416,1993]" captionTargetPageId="5" captionText="Figs 1–13: Drusilla watanabei: male forebody (1); female forebody (2); antenna (3); male abdomen (4); male tergite VIII (5); male sternite VIII (6); median lobe of aedeagus in lateral and in ventral view (7–8); paramere (9); female tergite VIII (10); female sternite VIII (11); spermatheca in different aspects (12–13). Scale bars: 1–4: 1.0 mm; 5–11: 0.5 mm; 12–13: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887117" httpUri="https://zenodo.org/record/5887117/files/figure.png" pageId="6" pageNumber="249">Figs 12–13</figureCitation>
.
</paragraph>
<paragraph id="8BF236F5CA6CC51FFF1F571CCCA9CB6C" blockId="6.[189,796,1897,2081]" pageId="6" pageNumber="249">
<emphasis id="B939EAE7CA6CC51FFF1F571CCF80C4CC" bold="true" box="[189,410,1897,1921]" pageId="6" pageNumber="249">Comparative notes</emphasis>
: This highly distinctive species differs from other 
<taxonomicName id="4C4D4D76CA6CC51FFE2357FCCC32C4EC" box="[385,552,1929,1953]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="249" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B939EAE7CA6CC51FFE2357FCCFCCC4EC" box="[385,470,1929,1953]" italics="true" pageId="6" pageNumber="249">Drusilla</emphasis>
species
</taxonomicName>
known from 
<collectingCountry id="F35A7665CA6CC51FFD1B57FCCCE0C4EC" box="[697,762,1929,1953]" name="China" pageId="6" pageNumber="249">China</collectingCountry>
by numerous characters, particularly its large body size, the long and slender antennae, the short elytra, the reduced hind wings, the male secondary sexual characters, and the male and female primary sexual characters.
</paragraph>
</subSubSection>
<subSubSection id="C357657ECA6CC51FFC9D509CCDF5C2AC" pageId="6" pageNumber="249" type="distribution">
<paragraph id="8BF236F5CA6CC51FFC9D509CCDF5C2AC" blockId="6.[831,1437,233,481]" pageId="6" pageNumber="249">
<emphasis id="B939EAE7CA6CC51FFC9D509CCAA2C24C" bold="true" box="[831,1208,233,257]" pageId="6" pageNumber="249">Distribution and natural history</emphasis>
: The known distribution now includes a total of three localities in 
<collectingRegion id="4989F817CA6CC51FFAF7517CCB83C26C" box="[1365,1433,265,289]" country="China" name="Hubei" pageId="6" pageNumber="249">Hubei</collectingRegion>
, 
<collectingRegion id="4989F817CA6CC51FFC9D515FCD91C20C" box="[831,907,298,321]" country="China" name="Hunan" pageId="6" pageNumber="249">Hunan</collectingRegion>
, and 
<collectingRegion id="4989F817CA6CC51FFC61515FCA0DC20C" box="[963,1047,298,321]" country="China" name="Yunnan" pageId="6" pageNumber="249">Yunnan</collectingRegion>
provinces. The examined specimens were sifted from litter, moss, and the roots of herbs in a terraced secondary pine forest at an altitude of 
<quantity id="4CB59B10CA6CC51FFAEB511CCB82C2CC" box="[1353,1432,361,385]" metricMagnitude="3" metricUnit="m" metricValue="2.62" pageId="6" pageNumber="249" unit="m" value="2620.0">2620 m</quantity>
. 
<bibRefCitation id="EFDC4B04CA6CC51FFC9D51FCCAA4C2EC" author="MARUYAMA, M. &amp; KISHIMOTO, T." box="[831,1214,393,417]" pageId="6" pageNumber="249" pagination="227 - 232" refId="ref14664" refString="MARUYAMA, M. &amp; KISHIMOTO, T. 2002 a: Myrmecophilous species of Drusilla (Coleoptera, Staphylinidae, Aleocharinae) associated with Lasius (Dendrolasius) spp. (Hymenoptera, Formicidae, Formicinae) from China. Part 1. - Special Bulletin of the Japanese Society of Coleopterology 5: 227 - 232." type="journal article" year="2002">MARUYAMA &amp; KISHIMOTO (2002a)</bibRefCitation>
found 
<taxonomicName id="4C4D4D76CA6CC51FFAB051FFCB86C2EC" authorityName="MARUYAMA &amp; KISHIMOTO" authorityYear="2002" box="[1298,1436,393,417]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="249" phylum="Arthropoda" rank="species" species="watanabei">
<emphasis id="B939EAE7CA6CC51FFAB051FFCB86C2EC" box="[1298,1436,393,417]" italics="true" pageId="6" pageNumber="249">D. watanabei</emphasis>
</taxonomicName>
associated with the ant 
<taxonomicName id="4C4D4D76CA6CC51FFBE751DFCDF1C2AC" authority="WHEELER, 1910" authorityName="WHEELER" authorityYear="1910" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="249" phylum="Arthropoda" rank="species" species="spathepus" subGenus="Dendrolasius">
<emphasis id="B939EAE7CA6CC51FFBE751DFCA9DC28C" box="[1093,1159,426,449]" italics="true" pageId="6" pageNumber="249">Lasius</emphasis>
(
<emphasis id="B939EAE7CA6CC51FFB3851DCCB39C28C" box="[1178,1315,425,449]" italics="true" pageId="6" pageNumber="249">Dendrolasius</emphasis>
) 
<emphasis id="B939EAE7CA6CC51FFA9451DCCB86C28C" box="[1334,1436,425,449]" italics="true" pageId="6" pageNumber="249">spathepus</emphasis>
WHEELER, 1910
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>