297 lines
42 KiB
XML
297 lines
42 KiB
XML
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<mods:title id="A0BAF523702CAE9238D50EF7FD050EB3">New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="D4ACD0C6BF2D228169AA35EDDF303AF0" type="personal">
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<mods:namePart id="16E45B6F0D100DB67C44F8F87F1A48D5">Assing, Volker</mods:namePart>
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<mods:title id="249B35E09D353B0275DF2C19C68F9A2C">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="52F4481353F9682EF7F795BF7EACD76B">
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<mods:date id="24818839C25583A8935745B17E2EC260">2015</mods:date>
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<mods:number id="03EAB7292F0E4A99E32C670C878AC518">2015-12-21</mods:number>
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<mods:number id="D8EAE57A228477D4587CCAF770E99D1C">65</mods:number>
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<treatment id="03E487E3CA63C512FF1F51A6CA06C10C" ID-DOI="http://doi.org/10.5281/zenodo.5878977" ID-GBIF-Taxon="192433646" ID-Zenodo-Dep="5878977" LSID="urn:lsid:plazi:treatment:03E487E3CA63C512FF1F51A6CA06C10C" httpUri="http://treatment.plazi.org/id/03E487E3CA63C512FF1F51A6CA06C10C" lastPageId="11" lastPageNumber="254" pageId="9" pageNumber="252">
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<subSubSection id="C357657ECA63C510FF1F51A6CC76C2A3" box="[189,620,467,494]" pageId="9" pageNumber="252" type="nomenclature">
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<paragraph id="8BF236F5CA63C510FF1F51A6CC76C2A3" blockId="9.[189,620,467,527]" box="[189,620,467,494]" pageId="9" pageNumber="252">
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<heading id="D0BA8199CA63C510FF1F51A6CC76C2A3" box="[189,620,467,494]" centered="true" fontSize="12" level="2" pageId="9" pageNumber="252" reason="2">
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<taxonomicName id="4C4D4D76CA63C510FF1F51A6CC76C2A3" ID-CoL="37TD7" authority="PACE, 1993" authorityName="PACE" authorityYear="1993" box="[189,620,467,494]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="252" phylum="Arthropoda" rank="species" species="yunnanensis" subGenus="Drusilla">
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<emphasis id="B939EAE7CA63C510FF1F51A6CF10C2A3" box="[189,266,467,494]" italics="true" pageId="9" pageNumber="252">Drusilla</emphasis>
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(
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<emphasis id="B939EAE7CA63C510FEBA51A6CF7FC2A3" box="[280,357,467,494]" italics="true" pageId="9" pageNumber="252">Drusilla</emphasis>
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)
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<emphasis id="B939EAE7CA63C510FED051A6CFEEC2A3" box="[370,500,467,494]" italics="true" pageId="9" pageNumber="252">yunnanensis</emphasis>
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<bibRefCitation id="EFDC4B04CA63C510FE5851A6CC76C2A3" author="PACE, R." box="[506,620,467,494]" pageId="9" pageNumber="252" pagination="69 - 126" refId="ref14851" refString="PACE, R. 1993: Aleocharinae della Cina (Coleoptera, Staphylinidae). - Bolletino del Museo Civico di Storia Naturale di Verona 17 (1990): 69 - 126." type="journal article" year="1993">PACE, 1993</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C357657ECA63C510FF1F518CCF3CC142" box="[189,294,505,527]" pageId="9" pageNumber="252" type="description">
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<paragraph id="8BF236F5CA63C510FF1F518CCF3CC142" blockId="9.[189,620,467,527]" box="[189,294,505,527]" pageId="9" pageNumber="252">
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(
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<figureCitation id="13762A70CA63C510FF63518CCF3BC142" box="[193,289,505,527]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="9" pageNumber="252">Figs 34–45</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C357657ECA63C510FF35525CCCF8C1EC" pageId="9" pageNumber="252" type="materials_examined">
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<paragraph id="8BF236F5CA63C510FF35525CCCF8C1EC" blockId="9.[150,756,553,673]" pageId="9" pageNumber="252">
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<emphasis id="B939EAE7CA63C510FF35525CCF6EC10C" bold="true" box="[151,372,553,577]" pageId="9" pageNumber="252">Material examined</emphasis>
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:
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<materialsCitation id="3B253CA8CA63C510FE2D525CCCC4C1EC" ID-GBIF-Occurrence="3447238342" collectingDate="2014-08-22" collectorName=" Assing & Schulke" country="China" elevation="1810" latitude="23.416666" location="Yunnan" longLatPrecision="1256" longitude="102.85" pageId="9" pageNumber="252" specimenCount="1" stateProvince="Yunnan">
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<emphasis id="B939EAE7CA63C510FE2D525CCFC8C10C" bold="true" box="[399,466,553,577]" pageId="9" pageNumber="252">
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<collectingCountry id="F35A7665CA63C510FE2D525CCFC8C10C" box="[399,466,553,577]" name="China" pageId="9" pageNumber="252">China</collectingCountry>
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</emphasis>
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:
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<emphasis id="B939EAE7CA63C510FE4C525CCC5EC10C" bold="true" box="[494,580,553,577]" pageId="9" pageNumber="252">
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<collectingRegion id="4989F817CA63C510FE4C525CCC5EC10C" box="[494,580,553,577]" country="China" name="Yunnan" pageId="9" pageNumber="252">Yunnan</collectingRegion>
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</emphasis>
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: 3 ♂♂, 2 ♀♀,
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<location id="8E92602ECA63C510FF35523FCEF7C12C" LSID="urn:lsid:plazi:treatment:03E487E3CA63C512FF1F51A6CA06C10C:8E92602ECA63C510FF35523FCEF7C12C" box="[151,237,586,609]" country="China" latitude="23.416666" longLatPrecision="1256" longitude="102.85" name="Yunnan" pageId="9" pageNumber="252" stateProvince="Yunnan">Yunnan</location>
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, mountains
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<collectorName id="26B85323CA63C510FE34523CCC1AC12C" box="[406,512,585,609]" pageId="9" pageNumber="252">S Jianshui</collectorName>
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,
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<geoCoordinate id="EE795032CA63C510FDBF523CCC6EC12C" box="[541,628,585,609]" degrees="23" direction="north" minutes="25" orientation="latitude" pageId="9" pageNumber="252" precision="925" value="23.416666">23°25'N</geoCoordinate>
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,
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<geoCoordinate id="EE795032CA63C510FD33523CCCF5C12C" box="[657,751,585,609]" degrees="102" direction="east" minutes="51" orientation="longitude" pageId="9" pageNumber="252" precision="925" value="102.85">102°51'E</geoCoordinate>
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,
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<quantity id="4CB59B10CA63C510FF35521CCEF2C1CC" box="[151,232,617,641]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="9" pageNumber="252" unit="m" value="1810.0">
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<elevation id="0060D1C6CA63C510FF35521CCEF2C1CC" box="[151,232,617,641]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="9" pageNumber="252" unit="m" value="1810.0">1810 m</elevation>
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</quantity>
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, secondary forest margin, litter and various debris sifted,
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<date id="FFF31035CA63C510FF7E52FCCF7CC1EC" box="[220,358,649,673]" pageId="9" pageNumber="252" value="2014-08-22">
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<collectingDate id="EFB7E9DDCA63C510FF7E52FCCF7CC1EC" box="[220,358,649,673]" pageId="9" pageNumber="252" value="2014-08-22">22.VIII.2014</collectingDate>
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</date>
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, leg.
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<collectorName id="26B85323CA63C510FE3852FCCFFBC1EC" box="[410,481,649,673]" pageId="9" pageNumber="252">Assing</collectorName>
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&
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<collectorName id="26B85323CA63C510FDA352FCCC4EC1EC" box="[513,596,649,673]" pageId="9" pageNumber="252">Schülke</collectorName>
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(cSch, cAss)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C357657ECA63C513FCB8509CCC8EC22C" lastPageId="10" lastPageNumber="253" pageId="9" pageNumber="252" type="discussion">
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<paragraph id="8BF236F5CA63C513FCB8509CCC8EC22C" blockId="9.[794,1399,233,674]" lastBlockId="10.[189,794,233,353]" lastPageId="10" lastPageNumber="253" pageId="9" pageNumber="252">
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<emphasis id="B939EAE7CA63C510FCB8509CCD93C24C" bold="true" box="[794,905,233,257]" pageId="9" pageNumber="252">Comment</emphasis>
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: In the original description, which is based on a male
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<typeStatus id="54F68857CA63C510FC31517CCDEBC26C" box="[915,1009,265,289]" pageId="9" pageNumber="252" type="holotype">holotype</typeStatus>
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from “
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<collectingRegion id="4989F817CA63C510FBE0517CCA83C26C" box="[1090,1177,265,289]" country="China" name="Yunnan" pageId="9" pageNumber="252">Yunnan</collectingRegion>
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, Dali” and a female
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<typeStatus id="54F68857CA63C510FCB8515FCD6DC20F" box="[794,887,298,322]" pageId="9" pageNumber="252" type="paratype">paratype</typeStatus>
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from “
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<collectingRegion id="4989F817CA63C510FC67515CCA06C20C" box="[965,1052,297,321]" country="China" name="Yunnan" pageId="9" pageNumber="252">Yunnan</collectingRegion>
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, Kunming”,
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<bibRefCitation id="EFDC4B04CA63C510FB01515CCB3FC20C" author="PACE, R." box="[1187,1317,297,321]" pageId="9" pageNumber="252" pagination="69 - 126" refId="ref14851" refString="PACE, R. 1993: Aleocharinae della Cina (Coleoptera, Staphylinidae). - Bolletino del Museo Civico di Storia Naturale di Verona 17 (1990): 69 - 126." type="journal article" year="1993">PACE (1993)</bibRefCitation>
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describes the coloration as “...; uriti liberi primo e secondo giallo-rossicci; antenne brune con i due articoli basali di un rossiccio scuro; zampe gialle con ginocchia medie e posteriori di un rossiccio scuro”. The other characters specified in the very short original description are too vague to be of taxonomic use. Despite the differences in the coloration (see the redescription below), the examined material is undoubtedly conspecific with the
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<typeStatus id="54F68857CA63C510FCB8523CCD60C12C" box="[794,890,585,609]" pageId="9" pageNumber="252" type="holotype">holotype</typeStatus>
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, as can be inferred from the identical shape of the median lobe of the aedeagus (compare
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<figureCitation id="13762A70CA63C510FAA6521CCB6CC1CC" box="[1284,1398,617,641]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="9" pageNumber="252">Figs 40–41</figureCitation>
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and figures
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<quantity id="4CB59B10CA63C510FC3B52FCCA08C1EC" box="[921,1042,649,673]" metricMagnitude="0" metricUnit="m" metricValue="3.6703" metricValueMax="3.683" metricValueMin="3.6576" pageId="9" pageNumber="252" unit="in" value="144.5" valueMax="145.0" valueMin="144.0">144–145 in</quantity>
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<bibRefCitation id="EFDC4B04CA63C510FBB852FCCA97C1EC" author="PACE, R." box="[1050,1165,649,673]" pageId="9" pageNumber="252" pagination="69 - 126" refId="ref14851" refString="PACE, R. 1993: Aleocharinae della Cina (Coleoptera, Staphylinidae). - Bolletino del Museo Civico di Storia Naturale di Verona 17 (1990): 69 - 126." type="journal article" year="1993">PACE 1993</bibRefCitation>
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). The type specimens may be teneral. The illustration of the spermatheca (
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<bibRefCitation id="EFDC4B04CA60C513FD46509CCEE9C26C" author="PACE, R." pageId="10" pageNumber="253" pagination="69 - 126" refId="ref14851" refString="PACE, R. 1993: Aleocharinae della Cina (Coleoptera, Staphylinidae). - Bolletino del Museo Civico di Storia Naturale di Verona 17 (1990): 69 - 126." type="journal article" year="1993">PACE 1993</bibRefCitation>
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||
: figure 141) of the
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<typeStatus id="54F68857CA60C513FE63517FCC04C26F" box="[449,542,266,290]" pageId="10" pageNumber="253" type="paratype">paratype</typeStatus>
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is misleading. In order to allow a reliable identification of this species, a redescription and new illustrations are provided.
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</paragraph>
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</subSubSection>
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<caption id="DF32667DCA63C510FF3B5799CAE0CB52" ID-DOI="http://doi.org/10.5281/zenodo.5887127" ID-Zenodo-Dep="5887127" httpUri="https://zenodo.org/record/5887127/files/figure.png" pageId="9" pageNumber="252" startId="9.[153,192,2028,2050]" targetBox="[151,1379,695,2010]" targetPageId="9">
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<paragraph id="8BF236F5CA63C510FF3B5799CAE0CB52" blockId="9.[151,1399,2028,2079]" pageId="9" pageNumber="252">
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<emphasis id="B939EAE7CA63C510FF3B5799CEE7CB4C" bold="true" box="[153,253,2028,2050]" pageId="9" pageNumber="252">Figs 26–33</emphasis>
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:
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<emphasis id="B939EAE7CA63C510FEA75799CF93CB4C" box="[261,393,2028,2049]" italics="true" pageId="9" pageNumber="252">Drusillapalata</emphasis>
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<taxonomicNameLabel id="A20A579CCA63C510FE2B5798CFF3CB4F" box="[393,489,2029,2050]" pageId="9" pageNumber="252" rank="species">spec. nov.</taxonomicNameLabel>
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||
:maleforebody (
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||
<emphasis id="B939EAE7CA63C510FDDE5799CC89CB4C" bold="true" box="[636,659,2028,2049]" pageId="9" pageNumber="252">26</emphasis>
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||
);antenna (
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||
<emphasis id="B939EAE7CA63C510FD595799CD08CB4C" bold="true" box="[763,786,2028,2049]" pageId="9" pageNumber="252">27</emphasis>
|
||
);maleabdomen (
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<emphasis id="B939EAE7CA63C510FC175799CDD6CB4C" bold="true" box="[949,972,2028,2049]" pageId="9" pageNumber="252">28</emphasis>
|
||
);maletergiteVIII (
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||
<emphasis id="B939EAE7CA63C510FBDF5799CA8ECB4C" bold="true" box="[1149,1172,2028,2049]" pageId="9" pageNumber="252">29</emphasis>
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||
);malesternite VIII (
|
||
<emphasis id="B939EAE7CA63C510FAF05799CB73CB4C" bold="true" box="[1362,1385,2028,2049]" pageId="9" pageNumber="252">30</emphasis>
|
||
); median lobe of aedeagus in lateral and in ventral view (
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<emphasis id="B939EAE7CA63C510FD36587CCCD4CB53" bold="true" box="[660,718,2057,2078]" pageId="9" pageNumber="252">31–32</emphasis>
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||
); paramere (
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<emphasis id="B939EAE7CA63C510FCE4587CCD47CB53" bold="true" box="[838,861,2057,2078]" pageId="9" pageNumber="252">33</emphasis>
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||
). Scale bars: 26–28: 1.0 mm; 29–33: 0.5 mm.
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</paragraph>
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</caption>
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<subSubSection id="C357657ECA60C512FF1F51FCCAEAC26C" lastPageId="11" lastPageNumber="254" pageId="10" pageNumber="253" type="description">
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<paragraph id="8BF236F5CA60C513FF1F51FCCA90C22C" blockId="10.[189,795,393,481]" lastBlockId="10.[831,1437,233,482]" pageId="10" pageNumber="253">
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<emphasis id="B939EAE7CA60C513FF1F51FCCF4CC2EC" bold="true" box="[189,342,393,417]" pageId="10" pageNumber="253">Redescription</emphasis>
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: Body length
|
||
<quantity id="4CB59B10CA60C513FE5251FCCC68C2EC" box="[496,626,393,417]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.3" metricValueMin="4.8" pageId="10" pageNumber="253" unit="mm" value="5.55" valueMax="6.3" valueMin="4.8">4.8–6.3 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CB59B10CA60C513FF5851DCCF61C28C" box="[250,379,425,449]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="10" pageNumber="253" unit="mm" value="2.45" valueMax="2.6" valueMin="2.3">2.3–2.6 mm</quantity>
|
||
. Coloration: body black, elytra with or without faint bronze hue; legs uniformly yellowish, rarely with the femoral apices indistinctly darker; antennae black, with antennomeres I–III slightly paler blackishbrown; maxillary palpi pale-brown to dark-brown, with the terminal palpomere yellow.
|
||
</paragraph>
|
||
<paragraph id="8BF236F5CA60C512FC9D511CCF10C22C" blockId="10.[831,1437,233,482]" lastBlockId="11.[150,757,233,1857]" lastPageId="11" lastPageNumber="254" pageId="10" pageNumber="253">
|
||
Head (
|
||
<figureCitation id="13762A70CA60C513FC28511CCA1BC2CC" box="[906,1025,361,385]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="10" pageNumber="253">Figs 34–35</figureCitation>
|
||
) approximately 1.1 times as broad as long; punctation of dorsal surface subject to sexual dimorphism; interstices without microsculpture and glossy. Eyes large and bulging, much longer than distance from posterior margin of eye to posterior constriction of head. Antenna (
|
||
<figureCitation id="13762A70CA61C512FEA0517CCF57C26C" box="[258,333,265,289]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 36</figureCitation>
|
||
) 2.0–
|
||
<quantity id="4CB59B10CA61C512FE28517CCFC0C26C" box="[394,474,265,289]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="254" unit="mm" value="2.1">2.1 mm</quantity>
|
||
long; antennomeres III–X gradually increasing in width; antennomeres VI–X transverse.
|
||
</paragraph>
|
||
<caption id="DF32667DCA60C513FF1C57A5CB87CB6D" ID-DOI="http://doi.org/10.5281/zenodo.5887129" ID-Zenodo-Dep="5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="10" pageNumber="253" startId="10.[190,230,2000,2022]" targetBox="[189,1436,500,1981]" targetPageId="10">
|
||
<paragraph id="8BF236F5CA60C513FF1C57A5CB87CB6D" blockId="10.[189,1437,2000,2081]" pageId="10" pageNumber="253">
|
||
<emphasis id="B939EAE7CA60C513FF1C57A5CF3CC4A8" bold="true" box="[190,294,2000,2022]" pageId="10" pageNumber="253">Figs 34–45</emphasis>
|
||
:
|
||
<taxonomicName id="4C4D4D76CA60C513FE9F57A5CC0AC4AB" authorityName="PACE" authorityYear="1993" box="[317,528,2000,2022]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="253" phylum="Arthropoda" rank="species" species="yunnanensis">
|
||
<emphasis id="B939EAE7CA60C513FE9F57A5CC0AC4AB" box="[317,528,2000,2022]" italics="true" pageId="10" pageNumber="253">Drusilla yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
: male forebody (
|
||
<emphasis id="B939EAE7CA60C513FD6957A5CCF9C4A8" bold="true" box="[715,739,2000,2021]" pageId="10" pageNumber="253">34</emphasis>
|
||
); female forebody (
|
||
<emphasis id="B939EAE7CA60C513FC1457A5CDD4C4A8" bold="true" box="[950,974,2000,2021]" pageId="10" pageNumber="253">35</emphasis>
|
||
); antenna (
|
||
<emphasis id="B939EAE7CA60C513FBE557A5CA45C4A8" bold="true" box="[1095,1119,2000,2021]" pageId="10" pageNumber="253">36</emphasis>
|
||
); female abdomen (
|
||
<emphasis id="B939EAE7CA60C513FA9557A5CB55C4A8" bold="true" box="[1335,1359,2000,2021]" pageId="10" pageNumber="253">37</emphasis>
|
||
); male tergite VIII (
|
||
<emphasis id="B939EAE7CA60C513FE945798CF54CB4F" bold="true" box="[310,334,2029,2050]" pageId="10" pageNumber="253">38</emphasis>
|
||
); male sternite VIII (
|
||
<emphasis id="B939EAE7CA60C513FDBD5798CC2DCB4F" bold="true" box="[543,567,2029,2050]" pageId="10" pageNumber="253">39</emphasis>
|
||
); median lobe of aedeagus in lateral and in ventral view (
|
||
<emphasis id="B939EAE7CA60C513FBD75798CAAACB4F" bold="true" box="[1141,1200,2029,2050]" pageId="10" pageNumber="253">40–41</emphasis>
|
||
); paramere (
|
||
<emphasis id="B939EAE7CA60C513FA8D5798CB5DCB4F" bold="true" box="[1327,1351,2029,2050]" pageId="10" pageNumber="253">42</emphasis>
|
||
); female tergite VIII (
|
||
<emphasis id="B939EAE7CA60C513FE94587ECF54CB6D" bold="true" box="[310,334,2059,2080]" pageId="10" pageNumber="253">43</emphasis>
|
||
); female sternite VIII (
|
||
<emphasis id="B939EAE7CA60C513FD88587ECC58CB6D" bold="true" box="[554,578,2059,2080]" pageId="10" pageNumber="253">44</emphasis>
|
||
); spermatheca (
|
||
<emphasis id="B939EAE7CA60C513FD7E587ECCEECB6D" bold="true" box="[732,756,2059,2080]" pageId="10" pageNumber="253">45</emphasis>
|
||
). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF236F5CA61C512FF35511FCC14C12C" blockId="11.[150,757,233,1857]" pageId="11" pageNumber="254">
|
||
Pronotum (
|
||
<figureCitation id="13762A70CA61C512FE82511CCF8CC2CC" box="[288,406,361,385]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Figs 34–35</figureCitation>
|
||
) approximately 1.06 times as broad as long and 1.10–1.13 times as broad as head, broadest at anterior angles; posterior angles obtusely marked; posteriorly with small transverse impression; midline with very fine line extending cephalad from posterior impression, but not reaching anterior margin of pronotum; punctation sexually dimorphic; interstices without microsculpture and glossy.
|
||
</paragraph>
|
||
<paragraph id="8BF236F5CA61C512FF35521CCC70C04C" blockId="11.[150,757,233,1857]" pageId="11" pageNumber="254">
|
||
Elytra (
|
||
<figureCitation id="13762A70CA61C512FF48521CCF7BC1CC" box="[234,353,617,641]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Figs 34–35</figureCitation>
|
||
) approximately 0.9 times as long as pronotum; humeral angles marked; punctation very dense, not very coarse, but much more distinct than that of pronotum. Hind wings fully developed. Metatarsomere I shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BF236F5CA61C512FF35537CCFD9C76C" blockId="11.[150,757,233,1857]" pageId="11" pageNumber="254">
|
||
Abdomen (
|
||
<figureCitation id="13762A70CA61C512FEB5537CCF78C06C" box="[279,354,777,801]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 37</figureCitation>
|
||
) approximately as broad as elytra; tergites III–VII without sexual dimorphism; anterior impressions of tergites III–V and anterior portion of tergite VI impunctate; tergite III with moderately sparse to very sparse fine punctation; tergites IV–VI with setiferous punctures at posterior margin, remainder of tergal surfaces with very sparse and minute punctures; tergite VII with sparse and fine punctation, posterior margin with palisade fringe.
|
||
</paragraph>
|
||
<paragraph id="8BF236F5CA61C512FF35545FCCD4C6AC" blockId="11.[150,757,233,1857]" pageId="11" pageNumber="254">
|
||
♂: postero-median portion of head with moderately sparse, somewhat coarser punctures, remainder of dorsal surface with fine and sparse punctation; pronotum (
|
||
<figureCitation id="13762A70CA61C512FF7154FCCF04C7EC" box="[211,286,1161,1185]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 34</figureCitation>
|
||
) with extensive, but not very deep median impression in posterior half; punctation of pronotum fine and moderately dense, postero-lateral edges of postero-median impression with coarser granulose sculpture; tergite VIII (
|
||
<figureCitation id="13762A70CA61C512FE34557CCFFAC66C" box="[406,480,1289,1313]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 38</figureCitation>
|
||
) strongly transverse and with distinctly serrate posterior margin, postero-lateral angles with more pronounced tooth on either side; sternite VIII (
|
||
<figureCitation id="13762A70CA61C512FEAA551CCF48C6CC" box="[264,338,1385,1409]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 39</figureCitation>
|
||
) weakly transverse, posterior margin weakly convex in the middle; median lobe of aedeagus (
|
||
<figureCitation id="13762A70CA61C512FF3C55DCCF0FC68C" box="[158,277,1449,1473]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Figs 40–41</figureCitation>
|
||
) relatively small, approximately
|
||
<quantity id="4CB59B10CA61C512FD3455DCCCEEC68C" box="[662,756,1449,1473]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="254" unit="mm" value="0.65">0.65 mm</quantity>
|
||
long; paramere
|
||
<quantity id="4CB59B10CA61C512FE9F55BCCF95C6AC" box="[317,399,1481,1505]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="254" unit="mm" value="0.6">0.6 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="13762A70CA61C512FD2255BCCCD0C6AC" box="[640,714,1481,1505]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 42</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF236F5CA61C512FF35559FCC9EC40C" blockId="11.[150,757,233,1857]" pageId="11" pageNumber="254">
|
||
♀: head (
|
||
<figureCitation id="13762A70CA61C512FEAF559CCF4CC54C" box="[269,342,1513,1537]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 35</figureCitation>
|
||
) with extemely fine, barely noticeable sparse punctation; pronotum (
|
||
<figureCitation id="13762A70CA61C512FD88567CCC69C56C" box="[554,627,1545,1569]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 35</figureCitation>
|
||
) somewhat depressed in posterior half, but without distinct impression (aside from the small posterior transverse impression); punctation of pronotum rather dense and fine, but more distinct than that of head; tergite VIII (
|
||
<figureCitation id="13762A70CA61C512FDE756FCCC94C5EC" box="[581,654,1673,1697]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 43</figureCitation>
|
||
) strongly transverse, posterior margin weakly concave in the middle; sternite VIII (
|
||
<figureCitation id="13762A70CA61C512FE8B56BCCF68C5AC" box="[297,370,1737,1761]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 44</figureCitation>
|
||
) distinctly shorter and more transverse than that of male, posterior margin weakly convex; spermatheca
|
||
<quantity id="4CB59B10CA61C512FE85577CCF9BC46C" box="[295,385,1801,1825]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="254" unit="mm" value="0.33">0.33 mm</quantity>
|
||
long, with large, apically truncate, and strongly sclerotized distal portion (
|
||
<figureCitation id="13762A70CA61C512FD8C575CCC63C40C" box="[558,633,1833,1857]" captionStart="Figs 34–45" captionStartId="10.[190,230,2000,2022]" captionTargetBox="[189,1436,500,1981]" captionTargetId="figure-247@10.[189,1436,499,1993]" captionTargetPageId="10" captionText="Figs 34–45: Drusilla yunnanensis: male forebody (34); female forebody (35); antenna (36); female abdomen (37); male tergite VIII (38); male sternite VIII (39); median lobe of aedeagus in lateral and in ventral view (40–41); paramere (42); female tergite VIII (43); female sternite VIII (44); spermatheca (45). Scale bars: 34–37: 1.0 mm; 38–39, 43–44: 0.5 mm; 40–42, 45: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5887129" httpUri="https://zenodo.org/record/5887129/files/figure.png" pageId="11" pageNumber="254">Fig. 45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF236F5CA61C512FF35571CCAEAC26C" blockId="11.[150,758,1897,2081]" lastBlockId="11.[794,1399,233,289]" pageId="11" pageNumber="254">
|
||
<emphasis id="B939EAE7CA61C512FF35571CCF7CC4CC" bold="true" box="[151,358,1897,1921]" pageId="11" pageNumber="254">Comparative notes</emphasis>
|
||
: Among the
|
||
<taxonomicName id="4C4D4D76CA61C512FE48571CCC8BC4CC" box="[490,657,1897,1921]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="254" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B939EAE7CA61C512FE48571CCC25C4CC" box="[490,575,1897,1921]" italics="true" pageId="11" pageNumber="254">Drusilla</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
recorded from
|
||
<collectingCountry id="F35A7665CA61C512FF7057FCCF0CC4EC" box="[210,278,1929,1953]" name="China" pageId="11" pageNumber="254">China</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C4D4D76CA61C512FE8157FFCFDEC4EC" authorityName="PACE" authorityYear="1993" box="[291,452,1930,1953]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="254" phylum="Arthropoda" rank="species" species="yunnanensis">
|
||
<emphasis id="B939EAE7CA61C512FE8157FFCFDEC4EC" box="[291,452,1930,1953]" italics="true" pageId="11" pageNumber="254">D. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
is characterized particularly by the dense punctation of the elytra, the fine punctation of the head and the pronotum, the impunctate anterior impressions of the abdominal tergites III–V, the male secondary sexual characters, the shape of the female tergite VIII, the morphology of the median lobe of the aedeagus, and the shape of the spermatheca.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C357657ECA61C512FCB8513CCA06C10C" pageId="11" pageNumber="254" type="distribution">
|
||
<paragraph id="8BF236F5CA61C512FCB8513CCA06C10C" blockId="11.[794,1399,329,577]" pageId="11" pageNumber="254">
|
||
<emphasis id="B939EAE7CA61C512FCB8513CCA98C22C" bold="true" box="[794,1154,329,353]" pageId="11" pageNumber="254">Distribution and natural history</emphasis>
|
||
: The currently known distribution is confined to three localities in
|
||
<collectingRegion id="4989F817CA61C512FABF511FCB6EC2CC" box="[1309,1396,362,385]" country="China" name="Yunnan" pageId="11" pageNumber="254">Yunnan</collectingRegion>
|
||
.
|
||
<bibRefCitation id="EFDC4B04CA61C512FCB851FCCA89C2EC" author="MARUYAMA, M. & KISHIMOTO, T." box="[794,1171,393,417]" pageId="11" pageNumber="254" pagination="227 - 232" refId="ref14664" refString="MARUYAMA, M. & KISHIMOTO, T. 2002 a: Myrmecophilous species of Drusilla (Coleoptera, Staphylinidae, Aleocharinae) associated with Lasius (Dendrolasius) spp. (Hymenoptera, Formicidae, Formicinae) from China. Part 1. - Special Bulletin of the Japanese Society of Coleopterology 5: 227 - 232." type="journal article" year="2002">MARUYAMA & KISHIMOTO (2002a)</bibRefCitation>
|
||
collected the species from trails of
|
||
<taxonomicName id="4C4D4D76CA61C512FC0D51DFCD57C2AC" authority="WHEELER, 1910" authorityName="WHEELER" authorityYear="1910" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="254" phylum="Arthropoda" rank="species" species="spathepus" subGenus="Dendrolasius">
|
||
<emphasis id="B939EAE7CA61C512FC0D51DFCDEBC28C" box="[943,1009,426,449]" italics="true" pageId="11" pageNumber="254">Lasius</emphasis>
|
||
(
|
||
<emphasis id="B939EAE7CA61C512FBA351DCCA90C28C" box="[1025,1162,425,449]" italics="true" pageId="11" pageNumber="254">Dendrolasius</emphasis>
|
||
)
|
||
<emphasis id="B939EAE7CA61C512FB3851DCCB1AC28C" box="[1178,1280,425,449]" italics="true" pageId="11" pageNumber="254">spathepus</emphasis>
|
||
WHEELER, 1910
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D76CA61C512FC2A51BFCB69C2AC" authority="KUZNETSOV-UGAMSKY, 1927" authorityName="KUZNETSOV-UGAMSKY" authorityYear="1927" box="[904,1395,457,481]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="254" phylum="Arthropoda" rank="species" species="capitatus" subGenus="Dendrolasius">
|
||
<emphasis id="B939EAE7CA61C512FC2A51BFCD87C2AC" box="[904,925,458,481]" italics="true" pageId="11" pageNumber="254">L.</emphasis>
|
||
(
|
||
<emphasis id="B939EAE7CA61C512FC0E51BFCDDFC2AC" box="[940,965,458,481]" italics="true" pageId="11" pageNumber="254">D.</emphasis>
|
||
)
|
||
<emphasis id="B939EAE7CA61C512FC7651BFCA28C2AC" box="[980,1074,458,481]" italics="true" pageId="11" pageNumber="254">capitatus</emphasis>
|
||
KUZNETSOV-UGAMSKY, 1927
|
||
</taxonomicName>
|
||
. The examined specimens were sifted from litter and various debris at the margin of a secondary mixed forest at an altitude of
|
||
<quantity id="4CB59B10CA61C512FC64525CCA0DC10C" box="[966,1047,553,577]" metricMagnitude="3" metricUnit="m" metricValue="1.81" pageId="11" pageNumber="254" unit="m" value="1810.0">1810 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |