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<document id="5D7C697FA4DBE2E49C042D2CB1FDFD67" ID-DOI="10.11646/zootaxa.5446.2.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="11101702" ID-ZooBank="2107786C-661B-42F7-8D0C-AFC1D1444C35" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714634805000" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="03E487DEFFFFF84BFF65FAB401FAFCAA" docLanguage="en" docName="zootaxa.5446.2.5.pdf" docOrigin="Zootaxa 5446 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5446.2.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paratrichodromeus angustilobatus Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="245" masterDocId="FFDDFFA6FFF8F842FFF2FFD90549FFF7" masterDocTitle="New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="246" masterPageNumber="236" pageNumber="243" updateTime="1714635388609" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="930DD7FE6AF02109502BA7D58FD1D968">New species and records of Hygrodromicus Tronquet, 1981 and Paratrichodromeus Zerche, 1992 from the Middle Asia, the Himalayan Region and China (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="4538E222FF8AFB9E3971C09C87BC472B">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="A2EC91E6620CEEF3EBFFB8B984090AEF">2024</mods:date>
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<treatment id="03E487DEFFFFF84BFF65FAB401FAFCAA" ID-DOI="http://doi.org/10.5281/zenodo.11101737" ID-Zenodo-Dep="11101737" LSID="urn:lsid:plazi:treatment:03E487DEFFFFF84BFF65FAB401FAFCAA" httpUri="http://treatment.plazi.org/id/03E487DEFFFFF84BFF65FAB401FAFCAA" lastPageId="9" lastPageNumber="245" pageId="7" pageNumber="243">
<subSubSection id="C3576543FFFFF845FF65FAB407CDFA7F" box="[151,644,1389,1416]" pageId="7" pageNumber="243" type="nomenclature">
<paragraph id="8BF236C8FFFFF845FF65FAB407CDFA7F" blockId="7.[151,644,1389,1453]" box="[151,644,1389,1416]" pageId="7" pageNumber="243">
<heading id="D0BA81A4FFFFF845FF65FAB407CDFA7F" bold="true" box="[151,644,1389,1416]" fontSize="11" level="1" pageId="7" pageNumber="243" reason="1">
<emphasis id="B939EADAFFFFF845FF65FAB407CDFA7F" bold="true" box="[151,644,1389,1416]" pageId="7" pageNumber="243">
<taxonomicName id="4C4D4D4BFFFFF845FF65FAB40772FA7F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,571,1389,1416]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="243" phylum="Arthropoda" rank="species" species="angustilobatus" status="sp. nov.">
<emphasis id="B939EADAFFFFF845FF65FAB40772FA7F" bold="true" box="[151,571,1389,1416]" italics="true" pageId="7" pageNumber="243">Paratrichodromeus angustilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57A1FFFFF845FDB0FAB707CDFA7F" box="[578,644,1390,1416]" pageId="7" pageNumber="243" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3576543FFFFF845FF65FA4B0411FA5A" box="[151,344,1426,1453]" pageId="7" pageNumber="243" type="description">
<paragraph id="8BF236C8FFFFF845FF65FA4B0411FA5A" blockId="7.[151,644,1389,1453]" box="[151,344,1426,1453]" pageId="7" pageNumber="243">
(
<figureCitation id="13762A4DFFFFF845FF6DFA4B046EFA5B" box="[159,295,1426,1452]" captionStart-0="FIGURES 611" captionStart-1="FIGURES 1214" captionStartId-0="4.[151,264,1943,1967]" captionStartId-1="8.[151,264,1166,1190]" captionTargetBox-0="[185,1403,184,1904]" captionTargetBox-1="[163,1424,193,1130]" captionTargetId-0="figure-16@4.[181,1406,181,1919]" captionTargetId-1="figure-16@8.[151,1436,181,1142]" captionTargetPageId-0="4" captionTargetPageId-1="8" captionText-0="FIGURES 611. Aedeagus of Hygrodromicus (Figs. 67: H. klapperichi, Figs. 89: H. latilobatus (holotype)) and Paratrichodromeus (Figs. 1011: P. angustilobatus): 6, 8, 10—parameral view, 7, 9, 11—lateral view. Scale bar: 0.1 mm." captionText-1="FIGURES 1214. Habitus of Paratrichodromeus: 12—P. angustilobatus (holotype), 13—P. punctulatus (Tajikistan, Hissar Mts.), 14—P. sukranensis (Kyrgyzstan, Trans-Alay Range). Scale bars: 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.11101706" figureDoi-1="http://doi.org/10.5281/zenodo.11101708" httpUri-0="https://zenodo.org/record/11101706/files/figure.png" httpUri-1="https://zenodo.org/record/11101708/files/figure.png" pageId="7" pageNumber="243">Figs. 1012</figureCitation>
,
<figureCitation id="13762A4DFFFFF845FEC1FA4B0419FA5A" box="[307,336,1426,1453]" captionStart="FIGURE 15" captionStartId="9.[151,250,1610,1634]" captionTargetBox="[309,1278,885,1586]" captionTargetId="figure-308@9.[309,1278,885,1586]" captionTargetPageId="9" captionText="FIGURE 15. Distribution of Paratrichodromeus in Central Asia: P. angustilobatus (triangle), P. punctulatus (circles), P. sukranensis (squares)." figureDoi="http://doi.org/10.5281/zenodo.11101712" httpUri="https://zenodo.org/record/11101712/files/figure.png" pageId="7" pageNumber="243">15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3576543FFFFF845FF65FA030151F9CB" pageId="7" pageNumber="243" type="materials_examined">
<paragraph id="8BF236C8FFFFF845FF65FA030151F9CB" blockId="7.[151,1437,1497,1956]" pageId="7" pageNumber="243">
<emphasis id="B939EADAFFFFF845FF65FA030488FA04" bold="true" box="[151,449,1497,1524]" pageId="7" pageNumber="243">Type material examined.</emphasis>
<typeStatus id="54F6886AFFFFF845FE3BFA030778FA03" box="[457,561,1498,1524]" pageId="7" pageNumber="243" type="holotype">Holotype</typeStatus>
<specimenCount id="9D4BFD41FFFFF845FDCBFA030707FA03" box="[569,590,1498,1524]" count="1" pageId="7" pageNumber="243" type="male"></specimenCount>
(
<figureCitation id="13762A4DFFFFF845FDADFA0307FAFA03" box="[607,691,1498,1524]" captionStart="FIGURES 1214" captionStartId="8.[151,264,1166,1190]" captionTargetBox="[163,1424,193,1130]" captionTargetId="figure-16@8.[151,1436,181,1142]" captionTargetPageId="8" captionText="FIGURES 1214. Habitus of Paratrichodromeus: 12—P. angustilobatus (holotype), 13—P. punctulatus (Tajikistan, Hissar Mts.), 14—P. sukranensis (Kyrgyzstan, Trans-Alay Range). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11101708" httpUri="https://zenodo.org/record/11101708/files/figure.png" pageId="7" pageNumber="243">Fig. 12</figureCitation>
; dissected): NE-Afghanistan, 2008 | Shiwa Lake,
<quantity id="4CB59B2DFFFFF845FB01FA0300C6FA03" box="[1267,1423,1498,1524]" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="7" pageNumber="243" unit="m" value="2850.0" valueMax="2900.0" valueMin="2800.0">28002900 m</quantity>
| 37°14/
<geoCoordinate id="EE79500FFFFFF845FF18FA270400F9EE" box="[234,329,1534,1561]" degrees="70" direction="east" minutes="58" orientation="longitude" pageId="7" pageNumber="243" precision="925" value="70.96667">70°58E</geoCoordinate>
| 19.21. VII., C. Reuter &lt;printed&gt;,
<typeStatus id="54F6886AFFFFF845FD0EFA2706DCF9EF" box="[764,917,1534,1560]" pageId="7" pageNumber="243" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="4C4D4D4BFFFFF845FC5FFA2701CEF9EF" authorityName="Zerche" authorityYear="1992" box="[941,1159,1534,1560]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="243" phylum="Arthropoda" rank="genus">
<emphasis id="B939EADAFFFFF845FC5FFA2701CEF9EF" box="[941,1159,1534,1560]" italics="true" pageId="7" pageNumber="243">Paratrichodromeus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C4D4D4BFFFFF845FB6DFA27000DF9EF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1183,1348,1534,1560]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="243" phylum="Arthropoda" rank="species" species="angustilobatus" status="sp. nov.">
<emphasis id="B939EADAFFFFF845FB6DFA27000DF9EF" box="[1183,1348,1534,1560]" italics="true" pageId="7" pageNumber="243">angustilobatus</emphasis>
</taxonomicName>
<emphasis id="B939EADAFFFFF845FABFFA2700C7F9EF" bold="true" box="[1357,1422,1534,1560]" pageId="7" pageNumber="243">
<taxonomicNameLabel id="A20A57A1FFFFF845FABFFA2700C7F9EF" box="[1357,1422,1534,1560]" pageId="7" pageNumber="243" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2024 &lt;red, printed&gt; (cF, to be eventually deposited in
<collectionCode id="ED5CAE0DFFFFF845FC3BF9FB0145F9CB" box="[969,1036,1570,1596]" country="Germany" name="Sammlung des Naturkundemseum Erfurt" pageId="7" pageNumber="243">NME</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3576543FFFFF84BFF35F99C01D7FD5E" lastPageId="9" lastPageNumber="245" pageId="7" pageNumber="243" type="description">
<paragraph id="8BF236C8FFFFF845FF35F99C040BF95E" blockId="7.[151,1437,1497,1956]" pageId="7" pageNumber="243">
<emphasis id="B939EADAFFFFF845FF35F99C043CF9A8" bold="true" box="[199,373,1605,1631]" pageId="7" pageNumber="243">Redescription.</emphasis>
Measurements: HW: 0.85; HL: 0.65; OL: 0.17; TL: 0.15; AL: 2.64; PL: 0.77; PWmax: 0.97; PWmin: 0.81; ESL: 1.32; EW: 1.52; MTbL: 1.10; MTrL: 0.61 (MTrL 14: 0.31; MTrL 5: 0.30); AW: 1.49; AedL: 1.22; BL: 5.75.
</paragraph>
<paragraph id="8BF236C8FFFFF845FF35F96B0672F853" blockId="7.[151,1437,1497,1956]" pageId="7" pageNumber="243">
Body narrow, elongate and moderately convex (
<figureCitation id="13762A4DFFFFF845FD20F96B066AF93B" box="[722,803,1714,1740]" captionStart="FIGURES 1214" captionStartId="8.[151,264,1166,1190]" captionTargetBox="[163,1424,193,1130]" captionTargetId="figure-16@8.[151,1436,181,1142]" captionTargetPageId="8" captionText="FIGURES 1214. Habitus of Paratrichodromeus: 12—P. angustilobatus (holotype), 13—P. punctulatus (Tajikistan, Hissar Mts.), 14—P. sukranensis (Kyrgyzstan, Trans-Alay Range). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11101708" httpUri="https://zenodo.org/record/11101708/files/figure.png" pageId="7" pageNumber="243">Fig. 12</figureCitation>
). Body reddish-brown, with distinctly paler apical portion of elytra and abdominal tergites IIIV; antennae brown, with paler basal parts of antennomeres 15; mouthparts and legs yellow-brown (tarsi slightly paler). Body glossy; head with fine microsculpture, fine and dense in middle of vertex, very dense and irregular in middle, and dense and isodiametric on infraorbital portions; neck with dense and coarse isodiametric microsculpture; pronotum with fine and dense transverse microreticulation, invisible in middle; scutellum with very fine isodiametric meshes in middle, lateral portions of visible parts of scutellum without any sculpture; abdomen with dense transverse microreticulation.
</paragraph>
<caption id="DF326640FFF0F84AFF65FB570681FB3D" ID-DOI="http://doi.org/10.5281/zenodo.11101708" ID-Zenodo-Dep="11101708" httpUri="https://zenodo.org/record/11101708/files/figure.png" pageId="8" pageNumber="244" startId="8.[151,264,1166,1190]" targetBox="[163,1424,193,1130]" targetPageId="8" targetType="figure">
<paragraph id="8BF236C8FFF0F84AFF65FB570681FB3D" blockId="8.[151,1436,1166,1227]" pageId="8" pageNumber="244">
<emphasis id="B939EADAFFF0F84AFF65FB57041EFB51" bold="true" box="[151,343,1166,1190]" pageId="8" pageNumber="244">FIGURES 1214.</emphasis>
Habitus of
<taxonomicName id="4C4D4D4BFFF0F84AFE24FB5707D2FB51" authorityName="Zerche" authorityYear="1992" box="[470,667,1166,1190]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B939EADAFFF0F84AFE24FB5707D2FB51" box="[470,667,1166,1190]" italics="true" pageId="8" pageNumber="244">Paratrichodromeus</emphasis>
</taxonomicName>
: 12—
<taxonomicName id="4C4D4D4BFFF0F84AFD2DFB5606D9FB51" authorityName="Shavrin" authorityYear="2024" box="[735,912,1166,1190]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="244" phylum="Arthropoda" rank="species" species="angustilobatus">
<emphasis id="B939EADAFFF0F84AFD2DFB5606D9FB51" box="[735,912,1166,1190]" italics="true" pageId="8" pageNumber="244">P. angustilobatus</emphasis>
</taxonomicName>
(holotype), 13—
<taxonomicName id="4C4D4D4BFFF0F84AFBB7FB56019FFB51" baseAuthorityName="Luze" baseAuthorityYear="1904" box="[1093,1238,1166,1190]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="244" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B939EADAFFF0F84AFBB7FB56019FFB51" box="[1093,1238,1166,1190]" italics="true" pageId="8" pageNumber="244">P. punctulatus</emphasis>
</taxonomicName>
(Tajikistan, Hissar Mts.), 14—
<taxonomicName id="4C4D4D4BFFF0F84AFEFFFB6A04E8FB3D" authorityName="Zerche" authorityYear="1992" box="[269,417,1202,1226]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="244" phylum="Arthropoda" rank="species" species="sukranensis">
<emphasis id="B939EADAFFF0F84AFEFFFB6A0455FB3D" box="[269,284,1203,1226]" italics="true" pageId="8" pageNumber="244">P</emphasis>
.
<emphasis id="B939EADAFFF0F84AFEDBFB6B04E8FB3D" box="[297,417,1202,1226]" italics="true" pageId="8" pageNumber="244">sukranensis</emphasis>
</taxonomicName>
(Kyrgyzstan, Trans-Alay Range). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BF236C8FFF0F84AFF35FAC40044F934" blockId="8.[151,1437,1309,2019]" pageId="8" pageNumber="244">Head 1.3 times as broad as long, with distinctly elevated medioapical portions between bases of antennae and infraorbital portions; supra-antennal protuberances moderately wide, strongly elevated; medioapical depression wide and very deep, narrowing posteriad and connected with moderately deep and narrow subrectangular interocellar depression; anteocellar foveae narrow, subdiagonally stretching anteriad toward level of anterior third of eyes; temples convex, long, slightly shorter than longitudinal length of eyes. Eyes moderately small, convex. Ocelli reduced. Punctation irregular, finer and sparser in middle and larger and denser on infraorbital portions; basal portions of vertex without sculpture. Preapical segment of maxillary palpi about as long as preceding segment, from middle gradually narrowed toward acute apex. Antennae reaching about middle length of elytra when reclined; basal antennomere about three times as long as wide, antennomere 2 slightly narrower and distinctly shorter than basal antennomere, 3 slightly shorter and narrower than 2, 4 slightly shorter than 3, 56 slightly longer than 4, 7 slightly longer and indistinctly broader than 6, 810 slightly shorter and broader than 7, apical antennomere slightly shorter than previous two antennomeres, from apical third gradually narrowing toward subacute apex.</paragraph>
<paragraph id="8BF236C8FFF0F84AFF35F91406D2F8A4" blockId="8.[151,1437,1309,2019]" pageId="8" pageNumber="244">Pronotum 1.2 times as broad as long, slightly broader than head, from widest anterior portion gradually narrowed posteriad toward obtuse posterior angles; laterobasal margins indistinctly concave; anterior margin rounded, about as long as slightly concave posterior margin; mediobasal portion with indistinct, small and oval impression. Punctation dense and fine, distinctly sparser in middle and meddiobasal portion.</paragraph>
<paragraph id="8BF236C8FFF0F84AFF35F8840753F880" blockId="8.[151,1437,1309,2019]" box="[199,538,1885,1911]" pageId="8" pageNumber="244">Scutellum without punctation.</paragraph>
<paragraph id="8BF236C8FFF0F84AFF35F8580724F814" blockId="8.[151,1437,1309,2019]" pageId="8" pageNumber="244">Elytra slightly broader than long, 1.7 times as long as pronotum, distinctly broadened posteriad, with very wide apical margins. Punctation about as that on pronotum, denser and deeper in basal and significantly finer and sparser in middle portion. Wings fully developed.</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FF4E0406FF22" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="245">Metatarsus slightly less than twice as long as metatibia; apical metatarsomere slightly shorter the preceding four metatarsomeres.</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FF0600D3FEB6" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="245">Abdomen slightly narrower than elytra, with two wide and transverse tomentose spots in the middle of abdominal tergite IV and indistinct two very small and transverse spots in the middle of tergite V; apical margin of abdominal tergite VII without distinct palisade fringe. Punctation regular, fine and dense, distinctly finer on abdominal tergite IV.</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FE9206D2FE02" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="245">
Male. Apical margin of abdominal tergite VIII rounded. Apical margin of abdominal sternite VIII sinuate. Aedeagus with wide basal portion, gradually narrowed toward median lobe with moderately wide apical part gradually narrowed toward small rounded apex; parameres narrow, distinctly exceeding apex of median lobe, with small two apical and two preapical setae; internal sac with elongate sclerotized structures in middle and moderately long flagellum (
<figureCitation id="13762A4DFFF1F84BFEB9FE0204D7FE02" box="[331,414,475,501]" captionStart="FIGURES 611" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1403,184,1904]" captionTargetId="figure-16@4.[181,1406,181,1919]" captionTargetPageId="4" captionText="FIGURES 611. Aedeagus of Hygrodromicus (Figs. 67: H. klapperichi, Figs. 89: H. latilobatus (holotype)) and Paratrichodromeus (Figs. 1011: P. angustilobatus): 6, 8, 10—parameral view, 7, 9, 11—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11101706" httpUri="https://zenodo.org/record/11101706/files/figure.png" pageId="9" pageNumber="245">Fig. 10</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="13762A4DFFF1F84BFCB4FE0206DEFE02" box="[838,919,475,501]" captionStart="FIGURES 611" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1403,184,1904]" captionTargetId="figure-16@4.[181,1406,181,1919]" captionTargetPageId="4" captionText="FIGURES 611. Aedeagus of Hygrodromicus (Figs. 67: H. klapperichi, Figs. 89: H. latilobatus (holotype)) and Paratrichodromeus (Figs. 1011: P. angustilobatus): 6, 8, 10—parameral view, 7, 9, 11—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11101706" httpUri="https://zenodo.org/record/11101706/files/figure.png" pageId="9" pageNumber="245">Fig. 11</figureCitation>
.
</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FE2604D8FDEE" blockId="9.[151,1437,151,861]" box="[199,401,511,537]" pageId="9" pageNumber="245">Female unknown.</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FDFB077FFD72" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="245">
<emphasis id="B939EADAFFF1F84BFF35FDFB04E6FDCA" bold="true" box="[199,431,546,573]" pageId="9" pageNumber="245">Comparative notes.</emphasis>
<taxonomicName id="4C4D4D4BFFF1F84BFE46FDFA0671FDCA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[436,824,547,573]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="angustilobatus" status="sp. nov.">
<emphasis id="B939EADAFFF1F84BFE46FDFA0671FDCA" box="[436,824,547,573]" italics="true" pageId="9" pageNumber="245">Paratrichodromeus angustilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57A1FFF1F84BFCCFFDFA0632FDCA" box="[829,891,547,573]" pageId="9" pageNumber="245" rank="species">
<emphasis id="B939EADAFFF1F84BFCCFFDFA063DFDCA" bold="true" box="[829,884,547,573]" pageId="9" pageNumber="245">sp. n</emphasis>
.
</taxonomicNameLabel>
can be reliably distinguished from
<taxonomicName id="4C4D4D4BFFF1F84BFB0EFDFA0461FD96" authority="(Luze, 1904)" baseAuthorityName="Luze" baseAuthorityYear="1904" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B939EADAFFF1F84BFB0EFDFA0044FDCB" box="[1276,1293,547,572]" italics="true" pageId="9" pageNumber="245">P</emphasis>
.
<emphasis id="B939EADAFFF1F84BFAE8FDFA00D4FDCA" box="[1306,1437,547,573]" italics="true" pageId="9" pageNumber="245">punctulatus</emphasis>
(
<bibRefCitation id="EFDC4B39FFF1F84BFF6DFD9E0456FD96" author="Luze, G." box="[159,287,583,609]" pageId="9" pageNumber="245" pagination="74 - 115" refId="ref6751" refString="Luze, G. (1904) Beitrag zur Staphyliniden-Fauna von Russisch-Central-Asien (Coleoptera). Horae Societatis Entomologicae Rossicae, 37, 74 - 115." type="journal article" year="1904">Luze, 1904</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4D4D4BFFF1F84BFEAFFD9E07D3FD96" authority="Zerche, 1992" authorityName="Zerche" authorityYear="1992" box="[349,666,583,609]" class="Insecta" family="Staphylinidae" genus="Paratrichodromeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="245" phylum="Arthropoda" rank="species" species="sukranensis">
<emphasis id="B939EADAFFF1F84BFEAFFD9E0427FD97" box="[349,366,583,608]" italics="true" pageId="9" pageNumber="245">P</emphasis>
.
<emphasis id="B939EADAFFF1F84BFE8EFD9E0749FD96" box="[380,512,583,609]" italics="true" pageId="9" pageNumber="245">sukranensis</emphasis>
<bibRefCitation id="EFDC4B39FFF1F84BFDF5FD9E07D3FD96" author="Zerche, L." box="[519,666,583,609]" pageId="9" pageNumber="245" pagination="105 - 142" refId="ref7195" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche, 1992</bibRefCitation>
</taxonomicName>
only by the shape of the narrower apical portion of the aedeagus and different structure of the internal sac.
</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FD5601D7FD5E" blockId="9.[151,1437,151,861]" box="[199,1182,655,681]" pageId="9" pageNumber="245">
For illustrations of the aedeagi of the compared species see
<figureCitation id="13762A4DFFF1F84BFC96FD56069AFD5E" box="[868,979,655,681]" captionStart="FIGURES 611" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[185,1403,184,1904]" captionTargetId="figure-16@4.[181,1406,181,1919]" captionTargetPageId="4" captionText="FIGURES 611. Aedeagus of Hygrodromicus (Figs. 67: H. klapperichi, Figs. 89: H. latilobatus (holotype)) and Paratrichodromeus (Figs. 1011: P. angustilobatus): 6, 8, 10—parameral view, 7, 9, 11—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.11101706" httpUri="https://zenodo.org/record/11101706/files/figure.png" pageId="9" pageNumber="245">
Figs 9,
<quantity id="4CB59B2DFFF1F84BFC45FD56069AFD5E" box="[951,979,655,681]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="245" unit="in" value="10.0">10</quantity>
</figureCitation>
in
<bibRefCitation id="EFDC4B39FFF1F84BFC05FD5601D0FD5E" author="Zerche, L." box="[1015,1177,655,681]" pageId="9" pageNumber="245" pagination="105 - 142" refId="ref7195" refString="Zerche, L. (1992) Zur Taxonomie, Phylogenie und Verbreitung der Hygrogeus - Gruppe Mittel- und Zentralasiens (Coleoptera: Staphylinidae: Omaliinae). Annalen des Naturhistorischen Museums in Wien, 93 (B), 105 - 142." type="journal article" year="1992">Zerche (1992)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3576543FFF1F84BFF35FD6B044EFCE2" pageId="9" pageNumber="245" type="distribution">
<paragraph id="8BF236C8FFF1F84BFF35FD6B0004FD3A" blockId="9.[151,1437,151,861]" box="[199,1357,690,717]" pageId="9" pageNumber="245">
<emphasis id="B939EADAFFF1F84BFF35FD6B0428FD3B" bold="true" box="[199,353,690,716]" pageId="9" pageNumber="245">Distribution.</emphasis>
The species is known only from the
<typeStatus id="54F6886AFFF1F84BFD0CFD6A0667FD3A" box="[766,814,691,717]" pageId="9" pageNumber="245">type</typeStatus>
locality in north-eastern Afghaninstan (
<figureCitation id="13762A4DFFF1F84BFB1EFD6A0076FD3A" box="[1260,1343,690,717]" captionStart="FIGURE 15" captionStartId="9.[151,250,1610,1634]" captionTargetBox="[309,1278,885,1586]" captionTargetId="figure-308@9.[309,1278,885,1586]" captionTargetPageId="9" captionText="FIGURE 15. Distribution of Paratrichodromeus in Central Asia: P. angustilobatus (triangle), P. punctulatus (circles), P. sukranensis (squares)." figureDoi="http://doi.org/10.5281/zenodo.11101712" httpUri="https://zenodo.org/record/11101712/files/figure.png" pageId="9" pageNumber="245">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BF236C8FFF1F84BFF35FD0F044EFCE2" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="245">
<emphasis id="B939EADAFFF1F84BFF35FD0F0402FD07" bold="true" box="[199,331,726,752]" pageId="9" pageNumber="245">Bionomics.</emphasis>
The specimen was collected at elevation
<quantity id="4CB59B2DFFF1F84BFCC4FD0E069CFD06" box="[822,981,727,753]" metricMagnitude="3" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="9" pageNumber="245" unit="m" value="2850.0" valueMax="2900.0" valueMin="2800.0">28002900 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3576543FFF1F84BFF35FCC601FAFCAA" pageId="9" pageNumber="245" type="etymology">
<paragraph id="8BF236C8FFF1F84BFF35FCC601FAFCAA" blockId="9.[151,1437,151,861]" pageId="9" pageNumber="245">
<emphasis id="B939EADAFFF1F84BFF35FCC60407FCCE" bold="true" box="[199,334,799,825]" pageId="9" pageNumber="245">Etymology.</emphasis>
The specific epithet is a combination of Latin adjective
<emphasis id="B939EADAFFF1F84BFC4DFCC6016AFCCF" box="[959,1059,799,824]" italics="true" pageId="9" pageNumber="245">angustus</emphasis>
, -
<emphasis id="B939EADAFFF1F84BFBCBFCC6010EFCCF" box="[1081,1095,799,824]" italics="true" pageId="9" pageNumber="245">a</emphasis>
, -
<emphasis id="B939EADAFFF1F84BFBACFCC601C9FCCF" box="[1118,1152,799,824]" italics="true" pageId="9" pageNumber="245">um</emphasis>
(narrow) and
<emphasis id="B939EADAFFF1F84BFAECFCC60039FCCE" box="[1310,1392,799,825]" italics="true" pageId="9" pageNumber="245">lobatus</emphasis>
, -
<emphasis id="B939EADAFFF1F84BFA75FCC600DCFCCF" box="[1415,1429,799,824]" italics="true" pageId="9" pageNumber="245">a</emphasis>
, -
<emphasis id="B939EADAFFF1F84BFF53FC9A058AFCAB" box="[161,195,835,860]" italics="true" pageId="9" pageNumber="245">um</emphasis>
(lobed). It refers to the moderately narrow apical part of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>