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<document id="363D1FC3EA9313C78152654BDFE67375" ID-CLB-Dataset="302762" ID-DOI="10.17161/jom.v0i25.4659" ID-GBIF-Dataset="a954f134-084d-41e5-8a41-61af66929a3b" ID-ISSN="2325-4467" ID-Zenodo-Dep="13736871" ID-ZooBank="urn:lsid:zoobank.org:pub:9AFCD1E3-42B6-4CB3-8321-35C60579289C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1722892427188" checkinUser="felipe" docAuthor="Engel, Michael S., Ortega-Blanco, Jaime, Nascimbene, Paul C. &amp; Singh, Hukam" docDate="2013" docId="03E487BF7578FFC4FE07FA9CFE7822F8" docLanguage="en" docName="JMelittology.2013.25.1-12.pdf" docOrigin="Journal of Melittology 2013 (25)" docSource="https://doi.org/10.17161/jom.v0i25.4659" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Protobombus Cockerell 1908" docType="treatment" docVersion="4" lastPageNumber="3" masterDocId="FFDDFFC7757AFFC7FFCFFFBDFF8D2740" masterDocTitle="The bees of Early Eocene Cambay amber (Hymenoptera: Apidae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1725892524702" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="28F1D7352B1C6DB8A311D5CB3BE7387F">The bees of Early Eocene Cambay amber (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="59890D85BB050C3C7CD0B7855B77A742">Engel, Michael S.</mods:namePart>
<mods:affiliation id="B922CE2F36434127B78D89E12EFDA06C">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu; jaimeortega @ ku. edu). &amp; Division of Invertebrate Zoology, American Museum of Natural History, Central Park West</mods:affiliation>
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<mods:namePart id="A050DFB255F0CB8E2A774AE5A3374CCE">Ortega-Blanco, Jaime</mods:namePart>
<mods:affiliation id="B6E67088B63858508E8F680616727262">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu; jaimeortega @ ku. edu).</mods:affiliation>
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<mods:namePart id="E3F0AD0122B991730BC3C6EB74B87545">Nascimbene, Paul C.</mods:namePart>
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<mods:namePart id="3FE84227D2C8DA6977851FDDD4F64A3B">Singh, Hukam</mods:namePart>
<mods:affiliation id="97FA4AD4FC2C357296CE5A22D67AF437">Birbal Sahni Institute of Palaeobotany, 53 University Road, Lucknow 226 007, India (hukams @ gmail. com).</mods:affiliation>
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<subSubSection id="C35765227578FFC5FE07FA9CFCE52279" box="[456,872,1313,1337]" pageId="2" pageNumber="3" type="nomenclature">
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<taxonomicName id="4C4D4D2A7578FFC5FE07FA9CFDDB2279" authority="Cockerell" authorityName="Cockerell" authorityYear="1908" box="[456,598,1313,1337]" class="Insecta" family="Apidae" genus="Protobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B939EABB7578FFC5FE07FA9CFDDB2279" box="[456,598,1313,1337]" italics="true" pageId="2" pageNumber="3">Protobombus</emphasis>
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? species indeterminate
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MATERIAL:
<collectionCode id="ED5CAE6C7578FFC5FE9BFADFFE23223A" box="[340,430,1378,1402]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
<materialsCitation id="3B253CF47578FFC5FE77FADCFCA522D9" collectingDate="2009-01-07" collectingDateMax="2009-01-12" collectingDateMin="2009-01-07" collectionCode="AMNH" country="India" county="Formation" location="Gujurat" municipality="Tadkeshwar" pageId="2" pageNumber="3" specimenCount="1">
Tad-41-A;
<collectingCountry id="F35A76397578FFC5FDF9FADCFDFA2239" box="[566,631,1377,1401]" name="India" pageId="2" pageNumber="3">India</collectingCountry>
:
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,
<collectingMunicipality id="6B96ACD37578FFC5FD3EFADCFC0C2239" box="[753,897,1377,1401]" pageId="2" pageNumber="3">Tadkeshwar</collectingMunicipality>
lignite mine; Cambay
<collectingCounty id="62934E257578FFC5FF6FFA3CFE9622D9" box="[160,283,1409,1433]" pageId="2" pageNumber="3">Formation</collectingCounty>
(Paleo-Eocene),
<date id="FFF310697578FFC5FE12FA3CFD3F22D9" box="[477,690,1409,1434]" pageId="2" pageNumber="3" value="2009-01-07" valueMax="2009-01-12" valueMin="2009-01-07">
<collectingDate id="EFB7E9817578FFC5FE12FA3CFD3F22D9" box="[477,690,1409,1434]" pageId="2" pageNumber="3" value="2009-01-07" valueMax="2009-01-12" valueMin="2009-01-07">712 January 2009</collectingDate>
</date>
(
<collectionCode id="ED5CAE6C7578FFC5FD0EFA3CFCAD22D9" box="[705,800,1409,1433]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
)
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.
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<paragraph id="8BF236A97578FFC4FF1FFA1CFE7822F8" blockId="2.[160,1168,1377,1785]" lastBlockId="3.[160,1168,1376,1464]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
COMMENTS: A poorly preserved female worker (length approximately
<quantity id="4CB59B4C7578FFC5FBE1FA1CFB0622F9" box="[1070,1163,1441,1465]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="3" unit="mm" value="7.5">7.5 mm</quantity>
, forewing length approximately
<quantity id="4CB59B4C7578FFC5FDEAFA7CFD0F229A" box="[549,642,1473,1498]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="3" unit="mm" value="6.7">6.7 mm</quantity>
) putatively of the electrapine genus
<taxonomicName id="4C4D4D2A7578FFC5FB83FA7FFEE122B9" authority="Cockerell" authorityName="Cockerell" authorityYear="1908" class="Insecta" family="Apidae" genus="Protobombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B939EABB7578FFC5FB83FA7FFF7F22B9" italics="true" pageId="2" pageNumber="3">Protobombus</emphasis>
Cockerell
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as indicated by the shape of the metabasitarsus. The specimen is poorly preserved with the wings outstretched from the body, the mesosoma of which is largely opened ventrally and cleared. The head is also only partially preserved, most of the right side and ventral portions being incomplete at the amber surface, with only the scape, pedicel, and basalmost flagellomeres preserved from the bees left side. The wings themselves are also somewhat partial with the anterior-apical portions of the bees right forewing missing and the left forewing largely crumpled and obscured. Given the diversity of these bees in the somewhat contemporaneous amber of the Baltic region (
<bibRefCitation id="EFDC4B587578FFC5FE5AF95CFDA021B9" author="Engel, M. S." box="[405,557,1761,1785]" pageId="2" pageNumber="3" pagination="1 - 192" refId="ref3854" refString="Engel, M. S. 2001 a. A monograph of the Baltic amber bees and evolution of the Apoidea (Hymenoptera). Bulletin of the American Museum of Natural History 259: 1 - 192." type="journal article" year="2001">Engel, 2001a</bibRefCitation>
,
<bibRefCitation id="EFDC4B587578FFC5FDF4F95CFDFF21B9" author="Engel, M. S." box="[571,626,1761,1785]" pageId="2" pageNumber="3" pagination="9 - 33" refId="ref3941" refString="Engel, M. S. 2004. Geological history of the bees (Hymenoptera: Apoidea). Revista de Tecnologia e Ambiente 10 (2): 9 - 33." type="journal article" year="2004">2004</bibRefCitation>
) as well as other deposits (
<bibRefCitation id="EFDC4B587578FFC4FC76F95CFF542238" author="Wappler, T. &amp; M. S. Engel" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" pagination="908 - 921" refId="ref5013" refString="Wappler, T., &amp; M. S. Engel. 2003. The middle Eocene bee faunas of Eckfeld and Messel, Germany (Hymenoptera: Apoidea). Journal of Paleontology 77 (5): 908 - 921." type="journal article" year="2003">Wappler &amp; Engel, 2003</bibRefCitation>
), it is perhaps not surprising that such similar species should be discovered in
<collectingCountry id="F35A76397579FFC4FF6FFA3DFE8E22D8" box="[160,259,1408,1432]" name="India" pageId="3" pageNumber="4">Cambay</collectingCountry>
amber. It is greatly hoped that more complete and well-preserved specimens will eventually be recovered.
</paragraph>
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