174 lines
18 KiB
XML
174 lines
18 KiB
XML
<document id="4AC23A0346306A753C2BFE3618C52684" ID-DOI="10.11646/zootaxa.2618.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5304087" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629699783393" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="03E487BBFFDD2B00FF3CFB1C386B1D33" docLanguage="en" docName="zt02618p035.pdf" docOrigin="Zootaxa 2618 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Nyctiophylax (Paranyctiophylax) kupirah Oláh & Johanson 2010, new species" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="FFDDFFC3FFD32B0FFFABFF9B39571F41" masterDocTitle="Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia" masterLastPageNumber="35" masterPageNumber="1" pageNumber="15" updateTime="1699097725960" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E40FDC1ED4E79B27BC5526F81B1CB4E3">
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<mods:title id="39A60BC6DFBA6FBE65E7733C3A251A4C">Fifteen new Trichoptera (Insecta) species from Sumatra, Indonesia</mods:title>
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</mods:titleInfo>
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<mods:name id="279C525405F2FB426AD00BC2C9FE1805" type="personal">
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<mods:roleTerm id="06A7C350EC94492B82EF250C5A6D3C5E">Author</mods:roleTerm>
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<mods:namePart id="14D67944BB2D09F6296AF9F9EDBD467A">Oláh, János</mods:namePart>
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<mods:affiliation id="87EC13810EBE206CBC4AFAFCD43BFB1C">Szent István University, Gödöll, Centre of Environmental Health. Residence postal address: Tarján u. 28, H- 4032 Debrecen, Hungary. E-mail: profolah @ gmail. com</mods:affiliation>
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<mods:roleTerm id="82BE38A2F91645B71D35A846B677BDE2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F925393E591A01A98C42F740D0E962E2">Johanson, Kjell Arne</mods:namePart>
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<mods:affiliation id="F7E2676E063FCFD2C4D9BDFA16BC50D2">Swedish Museum of Natural History, Entomology Department, Box 50007, S- 10405 Stockholm, Sweden.</mods:affiliation>
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<mods:typeOfResource id="89AEDA41A70DFDF306C5DB698468AC1E">text</mods:typeOfResource>
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<mods:title id="A7A6A7791DDA273212C34E3271C0C1DB">Zootaxa</mods:title>
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<mods:part id="15B0F9F6F754407BEDA15A70995E0602">
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<mods:date id="E11EFF23AF7D1FB7D94D4EFDA331A816">2010</mods:date>
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<mods:number id="B6DD491C354990BE323ABB7F70E87599">2010-09-20</mods:number>
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<mods:detail id="EC942BC88A7E2F02122288867EEF4C82" type="volume">
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<mods:number id="F2DCE59F4A59ABC553412CF0C8F0E5EE">2618</mods:number>
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<mods:url id="46207C57A49B61E6FFEBDACEFA08FE06">https://biotaxa.org/Zootaxa/article/view/zootaxa.2618.1.1</mods:url>
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<mods:classification id="E0E6CA09AB486165202A777B48CCB27A">journal article</mods:classification>
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<mods:identifier id="799D553771916DB5F3443AE9954D4783" type="DOI">10.11646/zootaxa.2618.1.1</mods:identifier>
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<mods:identifier id="D7210F6FB5FE1759F60D4835E8EB3AAB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E487BBFFDD2B00FF3CFB1C386B1D33" ID-DOI="http://doi.org/10.5281/zenodo.5311082" ID-Zenodo-Dep="5311082" LSID="urn:lsid:plazi:treatment:03E487BBFFDD2B00FF3CFB1C386B1D33" httpUri="http://treatment.plazi.org/id/03E487BBFFDD2B00FF3CFB1C386B1D33" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<subSubSection id="C3576526FFDD2B01FF3CFB1C3A7B1BE0" box="[151,812,1159,1185]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BF236ADFFDD2B01FF3CFB1C3A7B1BE0" blockId="14.[151,812,1159,1220]" box="[151,812,1159,1185]" pageId="14" pageNumber="15">
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<heading id="D0BA81C1FFDD2B01FF3CFB1C3A7B1BE0" bold="true" box="[151,812,1159,1185]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="B939EABFFFDD2B01FF3CFB1C3A7B1BE0" bold="true" box="[151,812,1159,1185]" pageId="14" pageNumber="15">
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<taxonomicName id="4C4D4D2EFFDD2B01FF3CFB1C3BD91BE0" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,654,1159,1185]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="kupirah" status="sp. nov." subGenus="Paranyctiophylax">
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<emphasis id="B939EABFFFDD2B01FF3CFB1C38601BE0" bold="true" box="[151,311,1159,1185]" italics="true" pageId="14" pageNumber="15">Nyctiophylax</emphasis>
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(
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<emphasis id="B939EABFFFDD2B01FEECFB1C3B4B1BE0" bold="true" box="[327,540,1159,1185]" italics="true" pageId="14" pageNumber="15">Paranyctiophylax</emphasis>
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)
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<emphasis id="B939EABFFFDD2B01FD85FB1C3BD91BE0" bold="true" box="[558,654,1159,1185]" italics="true" pageId="14" pageNumber="15">kupirah</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20A57C4FFDD2B01FD36FB1C3A7B1BE0" box="[669,812,1159,1185]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFDD2B00FF3CFB313D461ED3" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BF236ADFFDD2B01FF3CFB31384E1B85" blockId="14.[151,812,1159,1220]" box="[151,281,1194,1220]" pageId="14" pageNumber="15">
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<figureCitation id="13762A28FFDD2B01FF3CFB31384E1B85" box="[151,281,1194,1220]" captionStart="FIGURES 23–26" captionStartId="15.[151,271,1813,1837]" captionTargetBox="[177,1370,709,1756]" captionTargetId="figure-193@15.[151,1436,682,1792]" captionTargetPageId="15" captionText="FIGURES 23–26. Nyctiophylax (Paranyctiophylax) kupirah, new species, holotype. 23—genitalia, left lateral; 24— genitalia, dorsal; 25—gonocoxites, ventral; 26—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5304113" httpUri="https://zenodo.org/record/5304113/files/figure.png" pageId="14" pageNumber="15">Figs 23–26</figureCitation>
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</paragraph>
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<paragraph id="8BF236ADFFDD2B01FF3CFB6E3DE41AE5" blockId="14.[151,1436,1269,2004]" pageId="14" pageNumber="15">
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This species belongs in the
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<taxonomicName id="4C4D4D2EFFDD2B01FE4FFB6E3A971A4F" authorityName="Ulmer" authorityYear="1915" box="[484,960,1269,1295]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="flavus" subGenus="Paranyctiophylax">
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<emphasis id="B939EABFFFDD2B01FE4FFB6E3BD61A4F" box="[484,641,1269,1294]" italics="true" pageId="14" pageNumber="15">Nyctiophylax</emphasis>
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(
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<emphasis id="B939EABFFFDD2B01FD38FB6E3A301A4F" box="[659,871,1269,1294]" italics="true" pageId="14" pageNumber="15">Paranyctiophylax</emphasis>
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)
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<emphasis id="B939EABFFFDD2B01FCD2FB6E3A971A4F" box="[889,960,1269,1294]" italics="true" pageId="14" pageNumber="15">flavus</emphasis>
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</taxonomicName>
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species group and is most similar to
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<taxonomicName id="4C4D4D2EFFDD2B01FA2AFB6E38641A75" authorityName="Malicky" authorityYear="1993" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="padangensis">
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<emphasis id="B939EABFFFDD2B01FA2AFB6E38641A75" italics="true" pageId="14" pageNumber="15">N. padangensis</emphasis>
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</taxonomicName>
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from
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<collectingRegion id="4989F84FFFDD2B01FE29FA813B681A75" box="[386,575,1306,1332]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="15">North Sumatra</collectingRegion>
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, but differs by its triangular, not spatulate cerci; ventral paraproctal processes each with shorter ventrad-curving apical part; gonopods each with well-developed ventrobasal “elbow;” the shorter and basally curved ventroapical median lobe of sternite IX; and the presence of a pair of strong, anterad-directed spines and an ovoid phallotremal sclerite in the phallic apparatus.
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</paragraph>
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<paragraph id="8BF236ADFFDD2B01FF6EFA2B3A1319C5" blockId="14.[151,1436,1269,2004]" pageId="14" pageNumber="15">
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Male. Body uniformly pale brown. Maxillary palp formula IV-II-I-III-V, segment III originating from mesal face basal of apex of segment II. Forewings 4.7 mm long; brown, with hypertrophied longitudinal vein
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<emphasis id="B939EABFFFDD2B01FF3CFA6039FE1955" box="[151,169,1531,1556]" italics="true" pageId="14" pageNumber="15">R</emphasis>
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and pronounced hyaline patch pattern: small, rounded spot below discoidal cell around crossvein
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<emphasis id="B939EABFFFDD2B01FAF9FA603C2B1955" box="[1362,1404,1531,1556]" italics="true" pageId="14" pageNumber="15">r-m</emphasis>
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in subradial cell; and large, triangular spot along crossvein
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<emphasis id="B939EABFFFDD2B01FC97F9BB3A201978" box="[828,887,1568,1593]" italics="true" pageId="14" pageNumber="15">m-cu</emphasis>
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and point of basal bifurcation of
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<emphasis id="B939EABFFFDD2B01FAA3F9BB3C771978" box="[1288,1312,1568,1593]" italics="true" pageId="14" pageNumber="15">M</emphasis>
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; discoidal cell and median cell present; apical forks 2, 3, 4 and 5 present;
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<emphasis id="B939EABFFFDD2B01FC2CF9DE3AFF191F" box="[903,936,1605,1630]" italics="true" pageId="14" pageNumber="15">A1</emphasis>
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,
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<emphasis id="B939EABFFFDD2B01FC1CF9DE3A8F191F" box="[951,984,1605,1630]" italics="true" pageId="14" pageNumber="15">A2</emphasis>
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, and
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<emphasis id="B939EABFFFDD2B01FBB1F9DE3D6C191F" box="[1050,1083,1605,1630]" italics="true" pageId="14" pageNumber="15">A3</emphasis>
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looping. Hind wing discoidal cell closed, median cell open; apical forks 2 and 5 present.
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</paragraph>
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<paragraph id="8BF236ADFFDD2B01FF6EF90B38FE19EB" blockId="14.[151,1436,1269,2004]" box="[197,425,1680,1706]" pageId="14" pageNumber="15">Spur formula 3,4,4.</paragraph>
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<paragraph id="8BF236ADFFDD2B00FF6EF92E3D461ED3" blockId="14.[151,1436,1269,2004]" lastBlockId="15.[151,1436,152,626]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">Male genitalia. Sternite IX robust, triangular in lateral view with well-developed apicoventral mesal lobe having regular rectangular apex in ventral view and sinuous basal half in lateral view, shorter than ventrobasal “elbows” of gonopods. Tergite IX small, sclerotized; elongated obliquely vertical, discernible dorsal of sternite and anterior of paraproct. Segment X membranous, nearly invisible; with triangular mesal lobe in dorsal view. Cerci setose, triangular in lateral view; clavate in dorsal view; shorter than gonopods and ventral paraproctal processes. Paraproctal complex well-developed, highly sclerotized; consisting of pair of large ventral paraproctal processes curving ventrad, each bearing 4-6 setae in row on outer surface of ventradcurving apical half. Dorsal paraproctal processes forming pair of hyaline digitate processes. Gonopods elongate, slender, uniformly curving in ventral view; each with basal half broadening, forming ventrobasal “elbow;” distal half slender; apex slightly excised, located laterally. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproct and segment X; middle section of phallotheca forming short tube. Phallobase modified into long, narrow, arching dorsal apodeme and short ventral apodeme in lateral view; dorsal apodeme distinctly clear-cut, arching and narrowing into spine-like process in lateral view; weakly narrowing in dorsal view. Phallicata membranous, with pair of anterad-directed, stout spines and ovoid, horizontally elongate phallotremal sclerites embedded in phallic membrane.</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFDC2B00FF6EFE063AE11D43" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="8BF236ADFFDC2B00FF6EFE063AA51E9D" blockId="15.[151,1436,152,626]" pageId="15" pageNumber="16">
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<materialsCitation id="3B253CF0FFDC2B00FF6EFE063AA51E9D" collectingDate="2008-06-17" collectingDateMax="2008-06-19" collectingDateMin="2008-06-17" collectorName="J. Olah" country="Indonesia" county="Bukit Barisan Selatan National Park" elevation="800" location="Way Pitias" municipality="Barisan Mts." pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Sumatera Utara" typeStatus="holotype">
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<emphasis id="B939EABFFFDC2B00FF6EFE06382D1EF6" bold="true" box="[197,378,413,439]" pageId="15" pageNumber="16">
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<typeStatus id="54F6880FFFDC2B00FF6EFE0638601EF6" box="[197,311,413,439]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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:
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<emphasis id="B939EABFFFDC2B00FE21FE063B671EF6" bold="true" box="[394,560,413,439]" pageId="15" pageNumber="16">
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<collectingCountry id="F35A763DFFDC2B00FE21FE063B671EF6" box="[394,560,413,439]" name="Indonesia" pageId="15" pageNumber="16">INDONESIA</collectingCountry>
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</emphasis>
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(
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<collectingRegion id="4989F84FFFDC2B00FD94FE063BFF1EF6" box="[575,680,413,439]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="16">Sumatra</collectingRegion>
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):
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<collectingCounty id="62934E21FFDC2B00FD10FE063D301EF6" box="[699,1127,413,439]" pageId="15" pageNumber="16">Bukit Barisan Selatan National Park</collectingCounty>
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||
,
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<collectingMunicipality id="6B96ACD7FFDC2B00FBD8FE063C5C1EF6" box="[1139,1291,413,439]" pageId="15" pageNumber="16">Barisan Mts.</collectingMunicipality>
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||
,
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||
<location id="8E926076FFDC2B00FABCFE063CCF1EF6" LSID="urn:lsid:plazi:treatment:03E487BBFFDD2B00FF3CFB1C386B1D33:8E926076FFDC2B00FABCFE063CCF1EF6" box="[1303,1432,413,439]" country="Indonesia" county="Bukit Barisan Selatan National Park" municipality="Barisan Mts." name="Way Pitias" pageId="15" pageNumber="16" stateProvince="Sumatera Utara">Way Pitias</location>
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||
,
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<quantity id="4CB59B48FFDC2B00FF3CFE5939B31E9D" box="[151,228,450,476]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="800.0">
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||
<elevation id="0060D19EFFDC2B00FF3CFE5939B31E9D" box="[151,228,450,476]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="800.0">800 m</elevation>
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</quantity>
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,
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<date id="FFF3106DFFDC2B00FF44FE5938CC1E9D" box="[239,411,450,476]" pageId="15" pageNumber="16" value="2008-06-17" valueMax="2008-06-19" valueMin="2008-06-17">
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||
<collectingDate id="EFB7E985FFDC2B00FF44FE5938CC1E9D" box="[239,411,450,476]" pageId="15" pageNumber="16" value="2008-06-17" valueMax="2008-06-19" valueMin="2008-06-17">17–19.vi.2008</collectingDate>
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</date>
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,
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<collectingMethod id="520C4EBAFFDC2B00FE0CFE593B461E9D" box="[423,529,450,476]" pageId="15" pageNumber="16">light trap</collectingMethod>
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[
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<collectorName id="26B8537BFFDC2B00FD89FE593B2E1E9D" box="[546,633,450,476]" pageId="15" pageNumber="16">J. Oláh</collectorName>
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, Jr.]—
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<specimenCount id="9D4BFD24FFDC2B00FD6FFE593A441E9D" box="[708,787,450,476]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
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(OPC, in alcohol).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BF236ADFFDC2B00FF6EFE7C3AE11D43" blockId="15.[151,1436,152,626]" box="[197,950,487,514]" pageId="15" pageNumber="16">
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||
<materialsCitation id="3B253CF0FFDC2B00FF6EFE7C3AE51D43" box="[197,946,487,514]" collectingDate="2008-06-17" collectingDateMax="2008-06-19" collectingDateMin="2008-06-17" collectorName="J. Olah" country="Indonesia" county="Bukit Barisan Selatan National Park" elevation="800" location="Way Pitias" municipality="Barisan Mts." pageId="15" pageNumber="16" specimenCount="10" specimenCount-male="10" stateProvince="Sumatera Utara" typeStatus="paratype">
|
||
<emphasis id="B939EABFFFDC2B00FF6EFE7C38151D40" bold="true" box="[197,322,487,513]" pageId="15" pageNumber="16">
|
||
<typeStatus id="54F6880FFFDC2B00FF6EFE7C38151D40" box="[197,322,487,513]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
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||
: same data as holotype—
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<specimenCount id="9D4BFD24FFDC2B00FDC5FE733B801D43" box="[622,727,488,514]" pageId="15" pageNumber="16" type="male">10 males</specimenCount>
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(OPC, in alcohol)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576526FFDC2B00FF6EFD963B321D66" box="[197,613,525,551]" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="8BF236ADFFDC2B00FF6EFD963B321D66" blockId="15.[151,1436,152,626]" box="[197,613,525,551]" pageId="15" pageNumber="16">
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<emphasis id="B939EABFFFDC2B00FF6EFD9638301D66" bold="true" box="[197,359,525,551]" pageId="15" pageNumber="16">Distribution:</emphasis>
|
||
<collectingCountry id="F35A763DFFDC2B00FEC4FD9638B61D66" box="[367,481,525,551]" name="Indonesia" pageId="15" pageNumber="16">Indonesia</collectingCountry>
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(
|
||
<collectingRegion id="4989F84FFFDC2B00FE5AFD963B0E1D66" box="[497,601,525,551]" country="Indonesia" name="Sumatera Utara" pageId="15" pageNumber="16">Sumatra</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576526FFDC2B00FF6EFDA9386B1D33" pageId="15" pageNumber="16" type="etymology">
|
||
<paragraph id="8BF236ADFFDC2B00FF6EFDA9386B1D33" blockId="15.[151,1436,152,626]" pageId="15" pageNumber="16">
|
||
<emphasis id="B939EABFFFDC2B00FF6EFDA938001D0D" bold="true" box="[197,343,562,588]" pageId="15" pageNumber="16">Etymology:</emphasis>
|
||
<emphasis id="B939EABFFFDC2B00FEF4FDA938961D0A" box="[351,449,562,587]" italics="true" pageId="15" pageNumber="16">Kupirah</emphasis>
|
||
, from “kupirah,” elbow in Sanskrit, referring to the large, ventrobasal “elbows” of the gonopods.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |