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<document id="6021CEBBAB700228F733A1FEBFFA76FF" ID-CLB-Dataset="6096" ID-DOI="10.11646/zootaxa.4980.1.8" ID-GBIF-Dataset="465ec546-64e0-4ba3-84bb-b485feab12ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="4883127" ID-ZooBank="DF5C6B16-642A-4238-A4C2-AFDF9B2A1A29" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622463809428" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Cz-, Petr Baňař" docDate="2021" docId="03E487B6FFD7FFC8FF41880BFF2978C2" docLanguage="en" docName="zootaxa.4980.1.8.pdf" docOrigin="Zootaxa 4980 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Astenus grebennikovi Janak &amp; Banar 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="132" masterDocId="FFDDFFCEFFD4FFCDFFD68B57FFDD7A0F" masterDocTitle="Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="141" masterPageNumber="127" pageNumber="130" updateTime="1698995102808" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="FB4151C3DAC663EC5EB7F5072CC736CC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C66197C74C4F86903DC5FB3AD41E91AB">
<mods:title id="4FBF107066F2A42228D30338C5132652">Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae)</mods:title>
</mods:titleInfo>
<mods:name id="C4515BA367F1BB203D5D474617EB5A7A" type="personal">
<mods:role id="40577E32EAE184870B181592A4A68E74">
<mods:roleTerm id="2295703235D233E3CDB407230F4C5BFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DEBE07A6C23CB6F9724B726452551D17">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="12E1188F349D8AE05EEB4064FC961BB9" type="personal">
<mods:role id="92E0019895BE648695E94E33CD6A37AB">
<mods:roleTerm id="B437C2E62A27DB1599E28DF3372283ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="63A22ACAB533FAA56A982D9094E15956">Cz-, Petr Baňař</mods:namePart>
</mods:name>
<mods:typeOfResource id="75699A614EB826300E0E23308E71A06F">text</mods:typeOfResource>
<mods:relatedItem id="305A62533351F7B75C95A1CC39BAA0AE" type="host">
<mods:titleInfo id="4AC3BE4420A9DE0E31B5228206E77782">
<mods:title id="6FEF03AEE05E0FC2B70DE197696075F2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EBEEAF299D580AFE39ACE83C7D2007AB">
<mods:date id="4ABC996680AD427ADD72FE7A9D2072C5">2021</mods:date>
<mods:detail id="8F07C35DDC6EC7C68D7E2AF3FCCBB399" type="pubDate">
<mods:number id="2420D76ADBF1BB86BBBCB840DBBAF474">2021-05-31</mods:number>
</mods:detail>
<mods:detail id="D5E9B3684A7B0C6EDD0D1232D82CA503" type="volume">
<mods:number id="E8BE0E00825D961012A82F363A0101B8">4980</mods:number>
</mods:detail>
<mods:detail id="E2AB8FB3FFE300D11CA5A493CD1FEF91" type="issue">
<mods:number id="B679866364A4DC9384DFF65D07916CFF">1</mods:number>
</mods:detail>
<mods:extent id="C854B054F1224FEBEC852E3CC4BF7CCA" unit="page">
<mods:start id="B32079FB9BBD5DEF8A1D4674D49A5C3D">127</mods:start>
<mods:end id="CCBE6FFEF56334F330D580C0FA9409FD">141</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="26CC9547FD8C30C5AD4452EFBDBC9F3C">journal article</mods:classification>
<mods:identifier id="30B3C316E06A3B37DF1B2D63985FC13F" type="CLB-Dataset">6096</mods:identifier>
<mods:identifier id="2EF8375D5B53FE11BA6101CBAAFA7D6F" type="DOI">10.11646/zootaxa.4980.1.8</mods:identifier>
<mods:identifier id="733AFBE92CD4E9B5461E1AC613713657" type="GBIF-Dataset">465ec546-64e0-4ba3-84bb-b485feab12ab</mods:identifier>
<mods:identifier id="9F45DF1686FA6C0AC1B387496EBFE450" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="451C4D0DF3A9D382553916F0D8A2D0C2" type="Zenodo-Dep">4883127</mods:identifier>
<mods:identifier id="E4FB76DC7C2D42491493A691F1F91F3A" type="ZooBank">DF5C6B16-642A-4238-A4C2-AFDF9B2A1A29</mods:identifier>
</mods:mods>
<treatment id="03E487B6FFD7FFC8FF41880BFF2978C2" ID-DOI="http://doi.org/10.5281/zenodo.4945830" ID-GBIF-Taxon="182741902" ID-Zenodo-Dep="4945830" LSID="urn:lsid:plazi:treatment:03E487B6FFD7FFC8FF41880BFF2978C2" httpUri="http://treatment.plazi.org/id/03E487B6FFD7FFC8FF41880BFF2978C2" lastPageId="5" lastPageNumber="132" pageId="3" pageNumber="130">
<subSubSection id="C357652BFFD7FFCEFF41880BFD0F7979" box="[151,722,859,886]" pageId="3" pageNumber="130" type="nomenclature">
<paragraph id="8BF236A0FFD7FFCEFF41880BFD0F7979" blockId="3.[151,722,859,923]" box="[151,722,859,886]" pageId="3" pageNumber="130">
<heading id="D0BA81CCFFD7FFCEFF41880BFD0F7979" bold="true" box="[151,722,859,886]" fontSize="11" level="1" pageId="3" pageNumber="130" reason="1">
<emphasis id="B939EAB2FFD7FFCEFF41880BFD0F7979" bold="true" box="[151,722,859,886]" pageId="3" pageNumber="130">
<taxonomicName id="4C4D4D23FFD7FFCEFF41880BFDB57979" ID-CoL="8XV58" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" box="[151,616,859,886]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">
<emphasis id="B939EAB2FFD7FFCEFF41880BFE407979" bold="true" box="[151,413,859,886]" italics="true" pageId="3" pageNumber="130">Astenus grebennikovi</emphasis>
Janák &amp; Baňař
</taxonomicName>
,
<taxonomicNameLabel id="A20A57C9FFD7FFCEFDA3880BFD0F7979" box="[629,722,860,886]" pageId="3" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD7FFCEFF4188D7FE5F7994" box="[151,386,896,923]" pageId="3" pageNumber="130" type="description">
<paragraph id="8BF236A0FFD7FFCEFF4188D7FE5F7994" blockId="3.[151,722,859,923]" box="[151,386,896,923]" pageId="3" pageNumber="130">
(
<figureCitation id="13762A25FFD7FFCEFF4988D7FEFA7995" box="[159,295,896,922]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Figs. 12–23</figureCitation>
,
<figureCitation id="13762A25FFD7FFCEFEE588D7FEA67994" box="[307,379,896,923]" captionStart-0="FIGURES 42–43" captionStart-1="FIGURES 44–45" captionStartId-0="10.[151,264,1943,1967]" captionStartId-1="11.[151,264,1908,1932]" captionTargetBox-0="[173,1415,200,1920]" captionTargetBox-1="[154,1434,181,1884]" captionTargetId-0="figure-20@10.[173,1415,181,1920]" captionTargetId-1="figure-17@11.[154,1434,181,1884]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 42–43. Forest of Anjozorobe-Angavo. 42, View from camping site; 43, forest detail (both photos by M. Trýzna)." captionText-1="FIGURES 44–45. Forest of Anjozorobe-Angavo, microhabitats of Astenus grebennikovi Janák &amp; Baňař sp. nov. 42, detail of forest undergrowth; 43, detail of microhabitat of sift sample nr. 4 under Pandanus plant." figureDoi-0="http://doi.org/10.5281/zenodo.4883141" figureDoi-1="http://doi.org/10.5281/zenodo.4883145" httpUri-0="https://zenodo.org/record/4883141/files/figure.png" httpUri-1="https://zenodo.org/record/4883145/files/figure.png" pageId="3" pageNumber="130">42–45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD7FFCEFF41889FFA8C7F45" pageId="3" pageNumber="130" type="materials_examined">
<paragraph id="8BF236A0FFD7FFCEFF41889FFD6879ED" blockId="3.[151,1437,968,2038]" box="[151,693,968,994]" pageId="3" pageNumber="130">
<materialsCitation id="3B253CFDFFD7FFCEFF41889FFD6879ED" ID-GBIF-Occurrence="3131331327" box="[151,693,968,994]" country="Madagascar" location="Anjozorobe-Angavo" pageId="3" pageNumber="130" specimenCount="1" typeStatus="holotype">
<emphasis id="B939EAB2FFD7FFCEFF41889FFEEB79ED" bold="true" box="[151,310,968,994]" pageId="3" pageNumber="130">
<typeStatus id="54F68802FFD7FFCEFF41889FFF0F79ED" box="[151,210,968,994]" pageId="3" pageNumber="130">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F35A7630FFD7FFCEFEEA889FFE1879ED" box="[316,453,968,994]" name="Madagascar" pageId="3" pageNumber="130">Madagascar</collectingCountry>
,
<location id="8E92607BFFD7FFCEFE19889FFD6879ED" LSID="urn:lsid:plazi:treatment:03E487B6FFD7FFC8FF41880BFF2978C2:8E92607BFFD7FFCEFE19889FFD6879ED" box="[463,693,968,994]" country="Madagascar" name="Anjozorobe-Angavo" pageId="3" pageNumber="130">Anjozorobe-Angavo</location>
</materialsCitation>
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF1188BBFA8C7F45" blockId="3.[151,1437,968,2038]" pageId="3" pageNumber="130">
<emphasis id="B939EAB2FFD7FFCEFF1188BBFEAB7E09" bold="true" box="[199,374,1004,1030]" pageId="3" pageNumber="130">Type material.</emphasis>
<materialsCitation id="3B253CFDFFD7FFCEFEA888BBFE047E41" ID-GBIF-Occurrence="3131331354" collectingDate="2019-01-08" collectionCode="JJRC" collectorName="V. Grebennikov" country="Madagascar" determinerName="Banar" elevation="1424" latitude="-18.4118" location="Anjozorobe" longLatPrecision="7" longitude="47.9504" pageId="3" pageNumber="130" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B939EAB2FFD7FFCEFEA888BBFE367E09" bold="true" box="[382,491,1004,1030]" pageId="3" pageNumber="130">
<typeStatus id="54F68802FFD7FFCEFEA888BBFE367E09" box="[382,491,1004,1030]" pageId="3" pageNumber="130" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D4BFD29FFD7FFCEFE2488BAFDD77E08" box="[498,522,1005,1031]" pageId="3" pageNumber="130" type="male">♂</specimenCount>
:
<emphasis id="B939EAB2FFD7FFCEFDC188BBFD397E09" bold="true" box="[535,740,1004,1030]" pageId="3" pageNumber="130">
<collectingCountry id="F35A7630FFD7FFCEFDC188BBFD397E09" box="[535,740,1004,1030]" name="Madagascar" pageId="3" pageNumber="130">MADAGASCAR</collectingCountry>
</emphasis>
: “MADAGAS., Anjozorob. [
<location id="8E92607BFFD7FFCEFBE688BCFB607E09" LSID="urn:lsid:plazi:treatment:03E487B6FFD7FFC8FF41880BFF2978C2:8E92607BFFD7FFCEFBE688BCFB607E09" box="[1072,1213,1003,1030]" country="Madagascar" latitude="-18.4118" longLatPrecision="7" longitude="47.9504" name="Anjozorobe" pageId="3" pageNumber="130">Anjozorobe</location>
],
<geoCoordinate id="EE795067FFD7FFCEFB1988BBFAEF7E09" box="[1231,1330,1004,1030]" degrees="18.4118" direction="south" orientation="latitude" pageId="3" pageNumber="130" precision="5" value="-18.4118">-18.4118</geoCoordinate>
<geoCoordinate id="EE795067FFD7FFCEFAEC88BBFA457E08" box="[1338,1432,1004,1031]" degrees="47.9504" direction="east" orientation="longitude" pageId="3" pageNumber="130" precision="5" value="47.9504">47.9504</geoCoordinate>
,
<quantity id="4CB59B45FFD7FFCEFF418F47FF3A7E25" box="[151,231,1040,1066]" metricMagnitude="3" metricUnit="m" metricValue="1.424" pageId="3" pageNumber="130" unit="m" value="1424.0">
<elevation id="0060D193FFD7FFCEFF418F47FF3A7E25" box="[151,231,1040,1066]" metricMagnitude="3" metricUnit="m" metricValue="1.424" pageId="3" pageNumber="130" unit="m" value="1424.0">1424m</elevation>
</quantity>
,
<date id="FFF31060FFD7FFCEFF2E8F47FE887E25" box="[248,341,1040,1066]" pageId="3" pageNumber="130" value="2019-01-08">
<collectingDate id="EFB7E988FFD7FFCEFF2E8F47FE887E25" box="[248,341,1040,1066]" pageId="3" pageNumber="130" value="2019-01-08">8.i.2019</collectingDate>
</date>
, sift [sifting], MD36,
<collectorName id="26B85376FFD7FFCEFD8B8F46FCCE7E25" box="[605,787,1040,1067]" pageId="3" pageNumber="130">V. Grebennikov</collectorName>
//
<typeStatus id="54F68802FFD7FFCEFCEF8F47FC027E25" box="[825,991,1040,1066]" pageId="3" pageNumber="130" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="4C4D4D23FFD7FFCEFC3C8F47FB397E25" authority="Janák &amp; Cz-, 2021" authorityName="Janák &amp; Cz-" authorityYear="2021" box="[1002,1252,1040,1066]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">Astenus grebennikovi</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD7FFCEFB398F46FA907E24" box="[1263,1357,1041,1067]" pageId="3" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp;
<determinerName id="7E2EDA64FFD7FFCEFF618F63FF277E41" box="[183,250,1076,1102]" pageId="3" pageNumber="130">Baňař</determinerName>
det. 2020” (
<collectionCode id="ED5CAE65FFD7FFCEFE468F63FE0C7E41" box="[400,465,1076,1102]" pageId="3" pageNumber="130">JJRC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B253CFDFFD7FFCEFE318F63FA867E7D" ID-GBIF-Occurrence="3131331306" collectingDate="2019-01" collectingDateMax="2019-01-15" collectingDateMin="2019-01" collectionCode="JJRC" collectorName="P. Banar" country="Madagascar" elevation="1381" latitude="-18.410055" longLatPrecision="1" longitude="47.945084" pageId="3" pageNumber="130" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B939EAB2FFD7FFCEFE318F63FD827E41" bold="true" box="[487,607,1076,1102]" pageId="3" pageNumber="130">
<typeStatus id="54F68802FFD7FFCEFE318F63FD827E41" box="[487,607,1076,1102]" pageId="3" pageNumber="130" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D4BFD29FFD7FFCEFDA78F63FD7C7E40" box="[625,673,1076,1103]" pageId="3" pageNumber="130" type="male">1 ♂</specimenCount>
: “AJR/
<date id="FFF31060FFD7FFCEFD218F63FC877E40" box="[759,858,1076,1103]" pageId="3" pageNumber="130" value="2019-01">
<collectingDate id="EFB7E988FFD7FFCEFD218F63FC877E40" box="[759,858,1076,1103]" pageId="3" pageNumber="130" value="2019-01">Jan.2019</collectingDate>
</date>
/sift5 C MADAGASCAR, ANJOZOROBE forest,
<date id="FFF31060FFD7FFCEFF418F0FFEDE7E7C" box="[151,259,1112,1139]" pageId="3" pageNumber="130" value="2019-01-15">
<collectingDate id="EFB7E988FFD7FFCEFF418F0FFEDE7E7C" box="[151,259,1112,1139]" pageId="3" pageNumber="130" value="2019-01-15">15.i.2019</collectingDate>
</date>
, sifting forest litter
<quantity id="4CB59B45FFD7FFCEFE3E8F0FFDE57E7D" box="[488,568,1112,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.3809999999999998" pageId="3" pageNumber="130" unit="m" value="1381.0">
<elevation id="0060D193FFD7FFCEFE3E8F0FFDE57E7D" box="[488,568,1112,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.3809999999999998" pageId="3" pageNumber="130" unit="m" value="1381.0">1381m</elevation>
</quantity>
,
<geoCoordinate id="EE795067FFD7FFCEFD938F0FFD027E7D" box="[581,735,1112,1138]" degrees="18" direction="south" minutes="24" orientation="latitude" pageId="3" pageNumber="130" precision="1" seconds="36.2" value="-18.410055">S18°24’36.2”</geoCoordinate>
<geoCoordinate id="EE795067FFD7FFCEFD3E8F0FFC587E7C" box="[744,901,1112,1139]" degrees="47" direction="east" minutes="56" orientation="longitude" pageId="3" pageNumber="130" precision="1" seconds="42.3" value="47.945084">E47°56’42.3”</geoCoordinate>
, Winkler app. extr.
<collectorName id="26B85376FFD7FFCEFBBE8F0EFB1A7E7D" box="[1128,1223,1112,1139]" pageId="3" pageNumber="130">P. Baňař</collectorName>
lgt.” (
<collectionCode id="ED5CAE65FFD7FFCEFAC48F0FFA8E7E7D" box="[1298,1363,1112,1138]" pageId="3" pageNumber="130">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B253CFDFFD7FFCEFABF8F0FFC457EB5" ID-GBIF-Occurrence="3131331303" collectingDate="2019-01" collectingDateMax="2019-01-17" collectingDateMin="2019-01" collectionCode="MMBC" collectorName="P. Banar" country="Madagascar" elevation="1394" latitude="-18.40986" longLatPrecision="1" longitude="47.94528" pageId="3" pageNumber="130" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4BFD29FFD7FFCEFABF8F0FFA4A7E7C" box="[1385,1431,1112,1139]" pageId="3" pageNumber="130" type="male">1 ♂</specimenCount>
: “AJR/
<date id="FFF31060FFD7FFCEFF0B8F2BFEE27E99" box="[221,319,1148,1174]" pageId="3" pageNumber="130" value="2019-01">
<collectingDate id="EFB7E988FFD7FFCEFF0B8F2BFEE27E99" box="[221,319,1148,1174]" pageId="3" pageNumber="130" value="2019-01">Jan.2019</collectingDate>
</date>
/sift 8 C MADAGASCAR, ANJOZOROBE forest,
<date id="FFF31060FFD7FFCEFC548F2BFC337E99" box="[898,1006,1148,1174]" pageId="3" pageNumber="130" value="2019-01-17">
<collectingDate id="EFB7E988FFD7FFCEFC548F2BFC337E99" box="[898,1006,1148,1174]" pageId="3" pageNumber="130" value="2019-01-17">17.i.2019</collectingDate>
</date>
, sifting litter by big
<taxonomicName id="4C4D4D23FFD7FFCEFB078F2BFA457E99" box="[1233,1432,1148,1174]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="genus">
<emphasis id="B939EAB2FFD7FFCEFB078F2BFA9C7E99" box="[1233,1345,1148,1174]" italics="true" pageId="3" pageNumber="130">Pandanus</emphasis>
<quantity id="4CB59B45FFD7FFCEFA918F2BFA457E99" box="[1351,1432,1148,1174]" metricMagnitude="3" metricUnit="m" metricValue="1.394" pageId="3" pageNumber="130" unit="m" value="1394.0">
<elevation id="0060D193FFD7FFCEFA918F2BFA457E99" box="[1351,1432,1148,1174]" metricMagnitude="3" metricUnit="m" metricValue="1.394" pageId="3" pageNumber="130" unit="m" value="1394.0">1394m</elevation>
</quantity>
</taxonomicName>
,
<geoCoordinate id="EE795067FFD7FFCEFF418FF7FEEC7EB4" box="[151,305,1184,1211]" degrees="18" direction="south" minutes="24" orientation="latitude" pageId="3" pageNumber="130" precision="1" seconds="35.5" value="-18.40986">S18°24’35.5”</geoCoordinate>
<geoCoordinate id="EE795067FFD7FFCEFEE58FF7FE0C7EB4" box="[307,465,1184,1211]" degrees="47" direction="east" minutes="56" orientation="longitude" pageId="3" pageNumber="130" precision="1" seconds="43.0" value="47.94528">E47°56’43.0”</geoCoordinate>
,Winkler app.extr.
<collectorName id="26B85376FFD7FFCEFD4D8FF6FD297EB5" box="[667,756,1184,1211]" pageId="3" pageNumber="130">P.Baňař</collectorName>
lgt.”(
<collectionCode id="ED5CAE65FFD7FFCEFCE58FF7FC4D7EB5" box="[819,912,1184,1210]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="130">MMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B253CFDFFD7FFCEFC498FF7FE397F0D" ID-GBIF-Occurrence="3131331352" collectingDate="2019-01" collectingDateMax="2019-01-15" collectingDateMin="2019-01" collectionCode="MMBC" collectorName="P. Banar" country="Madagascar" elevation="1331" latitude="-18.413166" longLatPrecision="1" longitude="47.942528" pageId="3" pageNumber="130" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4BFD29FFD7FFCEFC498FF7FC157EB4" box="[927,968,1184,1211]" pageId="3" pageNumber="130" type="male">1♂</specimenCount>
:“AJR/
<date id="FFF31060FFD7FFCEFBC38FF7FBAA7EB5" box="[1045,1143,1184,1210]" pageId="3" pageNumber="130" value="2019-01">
<collectingDate id="EFB7E988FFD7FFCEFBC38FF7FBAA7EB5" box="[1045,1143,1184,1210]" pageId="3" pageNumber="130" value="2019-01">Jan.2019</collectingDate>
</date>
/sift7C MADAGASCAR, ANJOZOROBE forest,
<date id="FFF31060FFD7FFCEFE758F93FDD27ED0" box="[419,527,1220,1247]" pageId="3" pageNumber="130" value="2019-01-15">
<collectingDate id="EFB7E988FFD7FFCEFE758F93FDD27ED0" box="[419,527,1220,1247]" pageId="3" pageNumber="130" value="2019-01-15">15.i.2019</collectingDate>
</date>
, sifting forest litter by big tree
<quantity id="4CB59B45FFD7FFCEFCBE8F93FC657ED1" box="[872,952,1220,1246]" metricMagnitude="3" metricUnit="m" metricValue="1.331" pageId="3" pageNumber="130" unit="m" value="1331.0">
<elevation id="0060D193FFD7FFCEFCBE8F93FC657ED1" box="[872,952,1220,1246]" metricMagnitude="3" metricUnit="m" metricValue="1.331" pageId="3" pageNumber="130" unit="m" value="1331.0">1331m</elevation>
</quantity>
,
<geoCoordinate id="EE795067FFD7FFCEFC158F93FB807ED1" box="[963,1117,1220,1246]" degrees="18" direction="south" minutes="24" orientation="latitude" pageId="3" pageNumber="130" precision="1" seconds="47.4" value="-18.413166">S18°24’47.4”</geoCoordinate>
<geoCoordinate id="EE795067FFD7FFCEFBB58F93FADC7ED0" box="[1123,1281,1220,1247]" degrees="47" direction="east" minutes="56" orientation="longitude" pageId="3" pageNumber="130" precision="1" seconds="33.1" value="47.942528">E47°56’33.1”</geoCoordinate>
, Winkler app. extr.
<collectorName id="26B85376FFD7FFCEFF048FBEFEEF7F0D" box="[210,306,1256,1283]" pageId="3" pageNumber="130">P. Baňař</collectorName>
lgt.” (
<collectionCode id="ED5CAE65FFD7FFCEFEA88FBFFE067F0D" box="[382,475,1256,1282]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="130">MMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B253CFDFFD7FFCEFE248FBFFAB77F29" ID-GBIF-Occurrence="3131331319" collectingDate="2019-01" collectingDateMax="2019-01-15" collectingDateMin="2019-01" collectionCode="MMBC" collectorName="P. Banar" country="Madagascar" elevation="1382" latitude="-18.409777" longLatPrecision="1" longitude="47.94514" pageId="3" pageNumber="130" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4BFD29FFD7FFCEFE248FBFFDFD7F0C" box="[498,544,1256,1283]" pageId="3" pageNumber="130" type="female">1 ♀</specimenCount>
: “AJR/
<date id="FFF31060FFD7FFCEFDA38FBFFD057F0D" box="[629,728,1256,1282]" pageId="3" pageNumber="130" value="2019-01">
<collectingDate id="EFB7E988FFD7FFCEFDA38FBFFD057F0D" box="[629,728,1256,1282]" pageId="3" pageNumber="130" value="2019-01">Jan.2019</collectingDate>
</date>
/sift 4 C MADAGASCAR, ANJOZOROBE forest,
<date id="FFF31060FFD7FFCEFAFD8FBFFA457F0C" box="[1323,1432,1256,1283]" pageId="3" pageNumber="130" value="2019-01-15">
<collectingDate id="EFB7E988FFD7FFCEFAFD8FBFFA457F0C" box="[1323,1432,1256,1283]" pageId="3" pageNumber="130" value="2019-01-15">15.i.2019</collectingDate>
</date>
, sifting under
<taxonomicName id="4C4D4D23FFD7FFCEFEEE8E5BFDDB7F29" box="[312,518,1292,1318]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="3" pageNumber="130" phylum="Tracheophyta" rank="genus">
<emphasis id="B939EAB2FFD7FFCEFEEE8E5BFE757F29" box="[312,424,1292,1318]" italics="true" pageId="3" pageNumber="130">Pandanus</emphasis>
<quantity id="4CB59B45FFD7FFCEFE638E5BFDDB7F29" box="[437,518,1292,1318]" metricMagnitude="3" metricUnit="m" metricValue="1.382" pageId="3" pageNumber="130" unit="m" value="1382.0">
<elevation id="0060D193FFD7FFCEFE638E5BFDDB7F29" box="[437,518,1292,1318]" metricMagnitude="3" metricUnit="m" metricValue="1.382" pageId="3" pageNumber="130" unit="m" value="1382.0">1382m</elevation>
</quantity>
</taxonomicName>
,
<geoCoordinate id="EE795067FFD7FFCEFDC18E5BFD6C7F28" box="[535,689,1292,1319]" degrees="18" direction="south" minutes="24" orientation="latitude" pageId="3" pageNumber="130" precision="1" seconds="35.2" value="-18.409777">S18°24’35.2”</geoCoordinate>
<geoCoordinate id="EE795067FFD7FFCEFD688E5BFC867F28" box="[702,859,1292,1319]" degrees="47" direction="east" minutes="56" orientation="longitude" pageId="3" pageNumber="130" precision="1" seconds="42.5" value="47.94514">E47°56’42.5”</geoCoordinate>
, Winkler app. extr.
<collectorName id="26B85376FFD7FFCEFB988E5AFB6C7F29" box="[1102,1201,1292,1319]" pageId="3" pageNumber="130">P. Baňař</collectorName>
lgt.” (
<collectionCode id="ED5CAE65FFD7FFCEFAD28E5BFABC7F29" box="[1284,1377,1292,1318]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="3" pageNumber="130">MMBC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B253CFDFFD7FFCEFAAE8E5BFA907F45" ID-GBIF-Occurrence="3131331343" country="Madagascar" determinerName="Janak &amp; Banar" location="All" pageId="3" pageNumber="130" specimenCount="2" typeStatus="paratype">
<location id="8E92607BFFD7FFCEFAAE8E5BFA417F29" LSID="urn:lsid:plazi:treatment:03E487B6FFD7FFC8FF41880BFF2978C2:8E92607BFFD7FFCEFAAE8E5BFA417F29" box="[1400,1436,1292,1318]" country="Madagascar" name="All" pageId="3" pageNumber="130">All</location>
<typeStatus id="54F68802FFD7FFCEFF418E66FEDF7F44" box="[151,258,1329,1355]" pageId="3" pageNumber="130" type="paratype">paratypes</typeStatus>
with an additional label “
<typeStatus id="54F68802FFD7FFCEFDF78E67FD1F7F45" box="[545,706,1328,1354]" pageId="3" pageNumber="130" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="4C4D4D23FFD7FFCEFD1E8E67FC607F45" authority="Janák &amp; Cz-, 2021" authorityName="Janák &amp; Cz-" authorityYear="2021" box="[712,957,1328,1354]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">Astenus grebennikovi</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD7FFCEFC128E66FBC07F44" box="[964,1053,1329,1355]" pageId="3" pageNumber="130" rank="species">sp. nov.</taxonomicNameLabel>
,
<determinerName id="7E2EDA64FFD7FFCEFBFE8E67FBB57F45" box="[1064,1128,1328,1354]" pageId="3" pageNumber="130">Janák</determinerName>
&amp;
<determinerName id="7E2EDA64FFD7FFCEFB5D8E67FB137F45" box="[1163,1230,1328,1354]" pageId="3" pageNumber="130">Baňař</determinerName>
det. 2020”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD7FFC8FF118E03FC777B4E" lastPageId="5" lastPageNumber="132" pageId="3" pageNumber="130" type="description">
<paragraph id="8BF236A0FFD7FFCEFF118E03FE707FD5" blockId="3.[151,1437,968,2038]" pageId="3" pageNumber="130">
<emphasis id="B939EAB2FFD7FFCEFF118E03FE8C7F61" bold="true" box="[199,337,1364,1390]" pageId="3" pageNumber="130">Description</emphasis>
: Body length
<quantity id="4CB59B45FFD7FFCEFE208E03FDA27F60" box="[502,639,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.5" metricValueMin="5.5" pageId="3" pageNumber="130" unit="mm" value="6.5" valueMax="7.5" valueMin="5.5">5.5–7.5 mm</quantity>
(M =
<quantity id="4CB59B45FFD7FFCEFD1D8E03FCF87F60" box="[715,805,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="3" pageNumber="130" unit="mm" value="6.4">6.4 mm</quantity>
, HT =
<quantity id="4CB59B45FFD7FFCEFCAE8E03FC097F60" box="[888,980,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="3" pageNumber="130" unit="mm" value="5.5">5.5 mm</quantity>
), forebody length 3.0–
<quantity id="4CB59B45FFD7FFCEFB018E03FAF07F60" box="[1239,1325,1364,1391]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="130" unit="mm" value="3.3">3.3 mm</quantity>
(M =
<quantity id="4CB59B45FFD7FFCEFAAF8E03FF187F9C" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="130" unit="mm" value="3.2">3.2 mm</quantity>
, HT = 3.0 mm). Dark brown, anterior part of head and sometimes also abdomen dark reddish brown, legs light yellow, antennomeres light reddish yellow with mostly slightly lighter first two antennomeres, mouthparts light reddish yellow (
<figureCitation id="13762A25FFD7FFCEFE9A8E97FE7D7FD5" box="[332,416,1472,1498]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF118EB3FC727C80" blockId="3.[151,1437,968,2038]" pageId="3" pageNumber="130">
Head (
<figureCitation id="13762A25FFD7FFCEFEC68EB3FEBD7FF1" box="[272,352,1508,1534]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Fig. 13</figureCitation>
) moderately convex, a quarter longer than wide (L/W = 1.26–1.31, M = 1.28, HT = 1.27). Surface with coarse and dense umbilicate punctures. Eyes moderately large, temples on average about 1.3 times as long as eyes (TL/EL 1.23–1.43, M = 1.33, HT = 1.43). Temples markedly narrowed posteriorly. Antennae long and slender, fifth antennomere about 3 times as long as wide (L/W = 3.00–3.32, M = 3.16, HT = 3.00), tenth antennomere about two and half as long as wide (L/W = 2.38–2.56, M = 2.44, HT = 2.56).
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF118DCEFBFB7D11" blockId="3.[151,1437,968,2038]" pageId="3" pageNumber="130">
Pronotum (
<figureCitation id="13762A25FFD7FFCEFE918DCFFE447CBD" box="[327,409,1688,1714]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Fig. 13</figureCitation>
) egg-shaped, moderately convex, with shallow lateral impressions, about a quarter longer than wide (L/W = 1.23–1.30, M = 1.28, HT = 1.28) and moderately narrower than head (PW/HW = 0.86–0.90, M = 0.87, HT = 0.87). Sides markedly narrowed posteriorly, slightly emarginated in posterior half. Surface with coarse and dense umbilicate punctures similar as on head, unpunctured midline missing.
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF118C7FFAB47DA1" blockId="3.[151,1437,968,2038]" pageId="3" pageNumber="130">
Elytra (
<figureCitation id="13762A25FFD7FFCEFECA8C7FFE457D4D" box="[284,408,1832,1858]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Fig. 13, 14</figureCitation>
) elongate with rounded sides, concave along suture, about 1.4 times longer than wide (R = 1.34–1.49, M = 1.41, HT = 1.49). Surface moderately coarsely and densely punctured. Interstices shining, without microscupture. Shoulders with 4 tubercules bearing long, protruding fragile spines (
<figureCitation id="13762A25FFD7FFCEFB8C8C27FB707D85" box="[1114,1197,1904,1930]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Fig. 14</figureCitation>
) [in most specimens including HT most spines broken and missing]. Sides with additional a few shorter and less protruding spines.
</paragraph>
<paragraph id="8BF236A0FFD7FFCEFF118CEFFE9A7DF9" blockId="3.[151,1437,968,2038]" pageId="3" pageNumber="130">
Abdomen (
<figureCitation id="13762A25FFD7FFCEFE938CEFFE4A7DDD" box="[325,407,1976,2002]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="3" pageNumber="130">Fig. 12</figureCitation>
) slightly widened to segment VI, finely and densely punctured. Interstices shining, without microsculpture.
</paragraph>
<caption id="DF326628FFD0FFC9FF418C59FDB97D9D" ID-DOI="http://doi.org/10.5281/zenodo.4883133" ID-Zenodo-Dep="4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="4" pageNumber="131" startId="4.[151,264,1806,1830]" targetBox="[162,1427,189,1775]" targetPageId="4">
<paragraph id="8BF236A0FFD0FFC9FF418C59FDB97D9D" blockId="4.[151,1436,1806,1938]" pageId="4" pageNumber="131">
<emphasis id="B939EAB2FFD0FFC9FF418C59FE927D29" bold="true" box="[151,335,1806,1830]" pageId="4" pageNumber="131">FIGURES 12–23</emphasis>
.
<taxonomicName id="4C4D4D23FFD0FFC9FE888C59FD067D29" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" box="[350,731,1806,1830]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="131" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">
<emphasis id="B939EAB2FFD0FFC9FE888C59FDE57D29" box="[350,568,1806,1830]" italics="true" pageId="4" pageNumber="131">Astenus grebennikovi</emphasis>
Janák &amp; Baňař
</taxonomicName>
<emphasis id="B939EAB2FFD0FFC9FD358C59FCEB7D29" bold="true" box="[739,822,1806,1830]" pageId="4" pageNumber="131">
<taxonomicNameLabel id="A20A57C9FFD0FFC9FD358C59FCEB7D29" box="[739,822,1806,1830]" pageId="4" pageNumber="131" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21).
</paragraph>
</caption>
<paragraph id="8BF236A0FFD1FFC8FF118BC0FC777B4E" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="132">
Male. Posterior margin of sternite VII broadly emarginate and with almost 25 moderately long spines (
<figureCitation id="13762A25FFD1FFC8FAEA8BC0FA527ABE" box="[1340,1423,151,177]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="5" pageNumber="132">Fig. 22</figureCitation>
), sternite VIII long and narrow, with triangular emargination in posterior fifth (
<figureCitation id="13762A25FFD1FFC8FC258BECFB9A7ADA" box="[1011,1095,187,213]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="5" pageNumber="132">Fig. 23</figureCitation>
). Aedeagus (
<figureCitation id="13762A25FFD1FFC8FB008BECFA817ADA" box="[1238,1372,186,213]" captionStart="FIGURES 12–23" captionStartId="4.[151,264,1806,1830]" captionTargetBox="[162,1427,189,1775]" captionTargetId="figure-20@4.[151,1436,181,1782]" captionTargetPageId="4" captionText="FIGURES 12–23. Astenus grebennikovi Janák &amp; Baňař sp. nov. 12, 13, 15, 16, 19–23, holotype; 14, 17, 18, paratypes; 12, habitus; 13, forebody; 14, elytra; 15, 17, 19, aedeagus, lateral view; 16, 18, 20, 21, aedeagus, dorsal view; 22, posterior part of male sternite VII; 23, male sternite VIII; 15, 16, 19–21, transmitted light; 17, 18, reflected light. Scale bar: 1.0 mm (Fig. 12); 0.5 mm (Figs.13, 16–18); 0.1 mm (Figs. 19–21)." figureDoi="http://doi.org/10.5281/zenodo.4883133" httpUri="https://zenodo.org/record/4883133/files/figure.png" pageId="5" pageNumber="132">Figs. 15–21</figureCitation>
) long and slender, about
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long (AL =
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, M =
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, HT =
<quantity id="4CB59B45FFD1FFC8FC368B88FB947AF6" box="[992,1097,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" pageId="5" pageNumber="132" unit="mm" value="0.99">0.99 mm</quantity>
). Median lobe in ventral view narrow, sides markedly emarginate in middle part and with relatively wide sub-rounded point. Median lobe in lateral view regularly bent. Female. Sternite VIII largely rounded posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD1FFC8FF118A1DFEBB7BDD" pageId="5" pageNumber="132" type="diagnosis">
<paragraph id="8BF236A0FFD1FFC8FF118A1DFEBB7BDD" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="132">
<emphasis id="B939EAB2FFD1FFC8FF118A1DFE107B6B" bold="true" box="[199,461,330,356]" pageId="5" pageNumber="132">Differential diagnosis.</emphasis>
The new species is to be arranged in the last key of Madagascan
<taxonomicName id="4C4D4D23FFD1FFC8FB498A1CFA907B6A" box="[1183,1357,331,357]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B939EAB2FFD1FFC8FB498A1CFB2B7B6A" box="[1183,1270,331,357]" italics="true" pageId="5" pageNumber="132">Astenus</emphasis>
species
</taxonomicName>
(
<bibRefCitation id="EFDC4B51FFD1FFC8FA8D8A1CFEE77B86" author="Janak, J. &amp; Lecoq, J. - C." pageId="5" pageNumber="132" pagination="299 - 330" refId="ref7827" refString="Janak, J. &amp; Lecoq, J. - C. (2007) Treize nouvelles especes et nouvelles recoltes du genre Astenus Dejean, 1833 de Madagascar et des iles voisines (Coleoptera, Staphylinidae, Paederinae). Bulletin de la Societe entomologique de France, 112 (3), 299 - 330." type="journal article" year="2007">Janák &amp; Lecoq 2007</bibRefCitation>
) in the couplet 10 near
<taxonomicName id="4C4D4D23FFD1FFC8FD968A38FC9D7B86" authority="Lecoq, 1996" authorityName="Lecoq" authorityYear="1996" box="[576,832,367,393]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="species" species="duflosi">
<emphasis id="B939EAB2FFD1FFC8FD968A38FD767B86" box="[576,683,367,393]" italics="true" pageId="5" pageNumber="132">A. duflosi</emphasis>
<bibRefCitation id="EFDC4B51FFD1FFC8FD678A38FC9D7B86" author="Lecoq, J. - C." box="[689,832,367,393]" pageId="5" pageNumber="132" refId="ref7794" refString="Lecoq, J. - C. (1996) Faune de Madagascar. 86. Insectes coleopteres Staphylinidae Paederinae Genre Astenus. Museum national d'histoire naturelle, Paris, 73 pp." type="book" year="1996">Lecoq, 1996</bibRefCitation>
</taxonomicName>
, but it differs externally by more rounded temples and in males by markedly emarginate posterior margin of sternite VII and different shape of the aedeagus (compare
<figureCitation id="13762A25FFD1FFC8FAB38AC4FF007BDE" captionStart="FIGURES 31–39" captionStartId="8.[151,264,1908,1932]" captionTargetBox="[169,1418,316,1871]" captionTargetId="figure-73@8.[158,1430,309,1885]" captionTargetPageId="8" captionText="FIGURES 31–39. Astenus duflosi Lecoq. 31, 32, 36–39, Vohimana; 33–35, Andasibé; 31, habitus; 32, forebody; 33, 35, aedeagus, lateral view; 34, aedeagus, ventral view; 36, 37, apical part of aedeagus, ventral view; 38, posterior part of male sternite VII; 39, male sternite VIII; 33–35, reflected light; 36, 37, transmitted light. Scale bar: 1.0 mm (Fig. 31), 0.5 mm (Figs. 33–35, 38–39), 0.1 mm (Figs. 36 –37)." figureDoi="http://doi.org/10.5281/zenodo.4945834" httpUri="https://zenodo.org/record/4945834/files/figure.png" pageId="5" pageNumber="132">Figs. 31–39</figureCitation>
for details).
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD1FFC8FF118A8CFDB67816" pageId="5" pageNumber="132" type="etymology">
<paragraph id="8BF236A0FFD1FFC8FF118A8CFDB67816" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="132">
<emphasis id="B939EAB2FFD1FFC8FF118A8CFE937BFA" bold="true" box="[199,334,475,501]" pageId="5" pageNumber="132">Etymology.</emphasis>
This species is dedicated to Vasily Grebennikov, specialist in various groups of
<taxonomicName id="4C4D4D23FFD1FFC8FB348A8CFABD7BFA" authorityName="Linnaeus" authorityYear="1758" box="[1250,1376,475,501]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="5" pageNumber="132" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
, who collected the
<typeStatus id="54F68802FFD1FFC8FEFA8AA8FE537816" box="[300,398,511,537]" pageId="5" pageNumber="132" type="holotype">holotype</typeStatus>
of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C357652BFFD1FFC8FF118975FF2978C2" pageId="5" pageNumber="132" type="distribution">
<paragraph id="8BF236A0FFD1FFC8FF118975FEF8788A" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="132">
<emphasis id="B939EAB2FFD1FFC8FF118975FEBC7833" bold="true" box="[199,353,546,572]" pageId="5" pageNumber="132">Distribution.</emphasis>
<taxonomicName id="4C4D4D23FFD1FFC8FEBF8974FCD07832" authority="Janak &amp; Banar" authorityName="Janak &amp; Banar" authorityYear="2021" box="[361,781,547,573]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="132" phylum="Arthropoda" rank="species" species="grebennikovi" status="sp. nov.">
<emphasis id="B939EAB2FFD1FFC8FEBF8974FD877832" box="[361,602,547,573]" italics="true" pageId="5" pageNumber="132">Astenus grebennikovi</emphasis>
Janák &amp; Baňař
</taxonomicName>
<taxonomicNameLabel id="A20A57C9FFD1FFC8FCC08974FCB27832" box="[790,879,547,573]" pageId="5" pageNumber="132" rank="species">sp. nov.</taxonomicNameLabel>
is currently recorded only from the Anjozorobe-Angavo [Paysage Harmonieux Protègé du Complexe Anjozorobe-Angavo] in the northern central part of
<collectingCountry id="F35A7630FFD1FFC8FF41893CFEFD788A" box="[151,288,619,645]" name="Madagascar" pageId="5" pageNumber="132">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BF236A0FFD1FFC8FF1189D9FF2978C2" blockId="5.[151,1437,150,717]" pageId="5" pageNumber="132">
<emphasis id="B939EAB2FFD1FFC8FF1189D9FE9678A7" bold="true" box="[199,331,654,680]" pageId="5" pageNumber="132">Bionomics.</emphasis>
The
<typeStatus id="54F68802FFD1FFC8FE5089D8FE6B78A6" box="[390,438,655,681]" pageId="5" pageNumber="132">type</typeStatus>
series was sifted from forest litter (
<figureCitation id="13762A25FFD1FFC8FC9D89D8FC0B78A6" box="[843,982,654,681]" captionStart-0="FIGURES 42–43" captionStart-1="FIGURES 44–45" captionStartId-0="10.[151,264,1943,1967]" captionStartId-1="11.[151,264,1908,1932]" captionTargetBox-0="[173,1415,200,1920]" captionTargetBox-1="[154,1434,181,1884]" captionTargetId-0="figure-20@10.[173,1415,181,1920]" captionTargetId-1="figure-17@11.[154,1434,181,1884]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 42–43. Forest of Anjozorobe-Angavo. 42, View from camping site; 43, forest detail (both photos by M. Trýzna)." captionText-1="FIGURES 44–45. Forest of Anjozorobe-Angavo, microhabitats of Astenus grebennikovi Janák &amp; Baňař sp. nov. 42, detail of forest undergrowth; 43, detail of microhabitat of sift sample nr. 4 under Pandanus plant." figureDoi-0="http://doi.org/10.5281/zenodo.4883141" figureDoi-1="http://doi.org/10.5281/zenodo.4883145" httpUri-0="https://zenodo.org/record/4883141/files/figure.png" httpUri-1="https://zenodo.org/record/4883145/files/figure.png" pageId="5" pageNumber="132">Figs. 42–45</figureCitation>
). One teneral specimen was collected in January.
</paragraph>
</subSubSection>
</treatment>
</document>
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