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<mods:title id="32FCFB3311C788A825A142AC59FE6287">Phylogenetic systematics of day geckos, genus Phelsuma, based on molecular and morphological data (Squamata: Gekkonidae)</mods:title>
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<treatment id="03E487B6FFB3FFD0D5CB91E5FCC7FA99" ID-DOI="http://doi.org/10.5281/zenodo.5615795" ID-GBIF-Taxon="119384225" ID-Zenodo-Dep="5615795" LSID="urn:lsid:plazi:treatment:03E487B6FFB3FFD0D5CB91E5FCC7FA99" httpUri="http://treatment.plazi.org/id/03E487B6FFB3FFD0D5CB91E5FCC7FA99" lastPageNumber="23" pageId="22" pageNumber="23">
<subSubSection id="C357652BFFB3FFD0D5CB91E5FC26FC92" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8BF236A0FFB3FFD0D5CB91E5FE50FD77" blockId="22.[151,943,711,772]" box="[151,473,711,737]" pageId="22" pageNumber="23">
<heading id="D0BA81CCFFB3FFD0D5CB91E5FE50FD77" bold="true" box="[151,473,711,737]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B939EAB2FFB3FFD0D5CB91E5FE50FD77" bold="true" box="[151,473,711,737]" pageId="22" pageNumber="23">
<taxonomicName id="4C4D4D23FFB3FFD0D5CB91E5FE0EFD77" box="[151,391,711,737]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="B939EAB2FFB3FFD0D5CB91E5FE0EFD77" bold="true" box="[151,391,711,737]" italics="true" pageId="22" pageNumber="23">Phelsuma laticauda</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF236A0FFB3FFD0D5CB91C8FC26FC92" blockId="22.[151,943,711,772]" box="[151,943,746,772]" pageId="22" pageNumber="23">
(contains
<emphasis id="B939EAB2FFB3FFD0D45691C8FD70FC95" box="[266,761,746,771]" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C4D4D23FFB3FFD0D45691C8FE1EFC95" box="[266,407,746,771]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="laticauda">P. laticauda</taxonomicName>
,
<taxonomicName id="4C4D4D23FFB3FFD0D4FF91C8FDAAFC95" box="[419,547,746,771]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pasteuri">P. pasteuri</taxonomicName>
,
<taxonomicName id="4C4D4D23FFB3FFD0D77391C8FD70FC95" box="[559,761,746,771]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="robertmertensi">P. robertmertensi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C4D4D23FFB3FFD0D66E91C8FC2CFC95" box="[818,933,746,771]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="v-nigra">
<emphasis id="B939EAB2FFB3FFD0D66E91C8FC2CFC95" box="[818,933,746,771]" italics="true" pageId="22" pageNumber="23">P. v-nigra</emphasis>
</taxonomicName>
)
</paragraph>
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<subSubSection id="C357652BFFB3FFD0D5CB9017FCC7FA99" pageId="22" pageNumber="23" type="description">
<paragraph id="8BF236A0FFB3FFD0D5CB9017FAB1FC0C" blockId="22.[151,1437,821,1295]" pageId="22" pageNumber="23">
None of the members of
<taxonomicName id="4C4D4D23FFB3FFD0D4E69017FDCDFCD8" box="[442,580,821,846]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="B939EAB2FFB3FFD0D4E69017FDCDFCD8" box="[442,580,821,846]" italics="true" pageId="22" pageNumber="23">P. laticauda</emphasis>
</taxonomicName>
group as defined by
<bibRefCitation id="EFDC4B51FFB3FFD0D6669017FB99FCD9" box="[826,1040,821,847]" pageId="22" pageNumber="26" refString="Glaw, F., Vences, M. &amp; Ziegler, T. (1999) Bemerkungen zu Phelsuma dubia (Boettger, 1881): Wiederentdeckung des Holotypus, Verwandtschaftsbeziehungen und Daten zur Fortpflanzung. Salamandra, 35, 267 - 278." type="journal article">
Glaw
<emphasis id="B939EAB2FFB3FFD0D6DD9017FC32FCD8" box="[897,955,821,846]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
(1999)
</bibRefCitation>
(
<taxonomicName id="4C4D4D23FFB3FFD0D17D9017FB5AFCD8" box="[1057,1235,821,846]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="serraticauda">
<emphasis id="B939EAB2FFB3FFD0D17D9017FB5AFCD8" box="[1057,1235,821,846]" italics="true" pageId="22" pageNumber="23">P. serraticauda</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D23FFB3FFD0D0509017FA1BFCD8" box="[1292,1426,821,846]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="antanosy">
<emphasis id="B939EAB2FFB3FFD0D0509017FA1BFCD8" box="[1292,1426,821,846]" italics="true" pageId="22" pageNumber="23">P. antanosy</emphasis>
</taxonomicName>
) is actually closely related to
<taxonomicName id="4C4D4D23FFB3FFD0D4B99078FDF8FCE5" box="[485,625,858,883]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="B939EAB2FFB3FFD0D4B99078FDF8FCE5" box="[485,625,858,883]" italics="true" pageId="22" pageNumber="23">P. laticauda</emphasis>
</taxonomicName>
, according to our analysis:
<taxonomicName id="4C4D4D23FFB3FFD0D6E89078FBB2FCE5" box="[948,1083,858,883]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="antanosy">
<emphasis id="B939EAB2FFB3FFD0D6E89078FBB2FCE5" box="[948,1083,858,883]" italics="true" pageId="22" pageNumber="23">P. antanosy</emphasis>
</taxonomicName>
is in fact more closely related to
<taxonomicName id="4C4D4D23FFB3FFD0D5E990A2FEF7FC0F" box="[181,382,896,921]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="quadriocellata">
<emphasis id="B939EAB2FFB3FFD0D5E990A2FEF7FC0F" box="[181,382,896,921]" italics="true" pageId="22" pageNumber="23">P. quadriocellata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D23FFB3FFD0D4EB90A2FDE3FC0F" box="[439,618,896,921]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="serraticauda">
<emphasis id="B939EAB2FFB3FFD0D4EB90A2FDE3FC0F" box="[439,618,896,921]" italics="true" pageId="22" pageNumber="23">P. serraticauda</emphasis>
</taxonomicName>
cannot be clearly allocated to any species group (see below).
</paragraph>
<paragraph id="8BF236A0FFB3FFD0D5989087FCC7FA99" blockId="22.[151,1437,821,1295]" pageId="22" pageNumber="23">
Instead,
<taxonomicName id="4C4D4D23FFB3FFD0D4729087FE4AFC28" box="[302,451,933,958]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="B939EAB2FFB3FFD0D4729087FE4AFC28" box="[302,451,933,958]" italics="true" pageId="22" pageNumber="23">P. laticauda</emphasis>
</taxonomicName>
is recovered as the sister species to a clade containing the Comoran species
<taxonomicName id="4C4D4D23FFB3FFD0D0DA9087FEC0FC75" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="robertmertensi">
<emphasis id="B939EAB2FFB3FFD0D0DA9087FEC0FC75" italics="true" pageId="22" pageNumber="23">P. robertmertensi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4D4D23FFB3FFD0D40590E8FE53FC75" box="[345,474,970,995]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pasteuri">
<emphasis id="B939EAB2FFB3FFD0D40590E8FE53FC75" box="[345,474,970,995]" italics="true" pageId="22" pageNumber="23">P. pasteuri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D23FFB3FFD0D74B90E8FD06FC75" box="[535,655,970,995]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="v-nigra">
<emphasis id="B939EAB2FFB3FFD0D74B90E8FD06FC75" box="[535,655,970,995]" italics="true" pageId="22" pageNumber="23">P. v-nigra</emphasis>
</taxonomicName>
(
<figureCitation id="13762A25FFB3FFD0D7FC90E8FD62FC72" box="[672,747,970,996]" captionStart="FIGURE 7" captionStartId="17.[151,258,1686,1710]" captionTargetBox="[151,1436,194,1663]" captionTargetId="figure@17.[151,1436,194,1665]" captionTargetPageId="17" captionText="FIGURE 7. Molecular phylogeny of the Phelsuma lineata group (clade J), the P. laticauda group (clade K) and P. klemmeri as calculated from the multi-gene dataset of Rocha et al. (2009). Shown are the respective clades from the Bayesian 50 % majority-rule consensus trees based on (a) the combined dataset of mitochondrial and nuclear genes, (b) the mitochondrial genes only, and (c) the nuclear genes only. See caption to Fig. 2 for further explanations." httpUri="https://zenodo.org/record/194700/files/figure.png" pageId="22" pageNumber="23">Fig. 7</figureCitation>
; 100 PP in the mtDNA and combined analysis). Nuclear markers
<emphasis id="B939EAB2FFB3FFD0D45B90D2FEDFFB9F" box="[263,342,1008,1033]" italics="true" pageId="22" pageNumber="23">per se</emphasis>
do not provide enough resolution to recover these groups relationship, however, the monophyly of these three Comoroan species is strongly supported by both mtDNA and combined phylogenies (100 PP; clade L in
<figureCitation id="13762A25FFB3FFD0D4DB9718FE46FBC2" box="[391,463,1082,1108]" captionStart="FIGURE 7" captionStartId="17.[151,258,1686,1710]" captionTargetBox="[151,1436,194,1663]" captionTargetId="figure@17.[151,1436,194,1665]" captionTargetPageId="17" captionText="FIGURE 7. Molecular phylogeny of the Phelsuma lineata group (clade J), the P. laticauda group (clade K) and P. klemmeri as calculated from the multi-gene dataset of Rocha et al. (2009). Shown are the respective clades from the Bayesian 50 % majority-rule consensus trees based on (a) the combined dataset of mitochondrial and nuclear genes, (b) the mitochondrial genes only, and (c) the nuclear genes only. See caption to Fig. 2 for further explanations." httpUri="https://zenodo.org/record/194700/files/figure.png" pageId="22" pageNumber="23">Fig. 7</figureCitation>
a).
<taxonomicName id="4C4D4D23FFB3FFD0D4A89718FD5BFBC5" box="[500,722,1082,1107]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pasteuri">
<emphasis id="B939EAB2FFB3FFD0D4A89718FD5BFBC5" box="[500,722,1082,1107]" italics="true" pageId="22" pageNumber="23">Phelsuma pasteuri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D23FFB3FFD0D6519718FC53FBC5" box="[781,986,1082,1107]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="robertmertensi">
<emphasis id="B939EAB2FFB3FFD0D6519718FC53FBC5" box="[781,986,1082,1107]" italics="true" pageId="22" pageNumber="23">P. robertmertensi</emphasis>
</taxonomicName>
which are both endemic to the oldest island of the
<collectingCountry id="F35A7630FFB3FFD0D46F9742FE29FBEC" box="[307,416,1120,1146]" name="Comoros" pageId="22" pageNumber="23">Comoros</collectingCountry>
archipelago (
<collectingCountry id="F35A7630FFB3FFD0D71E9742FD21FBEC" box="[578,680,1120,1146]" name="Mayotte" pageId="22" pageNumber="23">Mayotte</collectingCountry>
) form a monophyletic group, strongly corroborated by all gene phylogenies, confirming previous results (
<bibRefCitation id="EFDC4B51FFB3FFD0D7D897A7FCDCFB09" author="Rocha" box="[644,853,1157,1183]" pageId="22" pageNumber="28" refString="Rocha S., Posada D., Carretero M. A., Harris D. J., 2007. Phylogenetic affinities of Comoroan and East African day geckos (genus Phelsuma): multiple natural colonisations, introductions and island radiations. Molecular Phylogenetics and Evolution, 43, 685 - 692." type="journal article" year="2007">
Rocha
<emphasis id="B939EAB2FFB3FFD0D78B97A7FC9BFB08" box="[727,786,1157,1182]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2007
</bibRefCitation>
) and supporting the specific status of
<taxonomicName id="4C4D4D23FFB3FFD0D04D97A7FA04FB08" box="[1297,1421,1157,1182]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="pasteuri">
<emphasis id="B939EAB2FFB3FFD0D04D97A7FA04FB08" box="[1297,1421,1157,1182]" italics="true" pageId="22" pageNumber="23">P. pasteuri</emphasis>
</taxonomicName>
. The
<taxonomicName id="4C4D4D23FFB3FFD0D5909789FEDEFB52" box="[204,343,1195,1220]" class="Reptilia" family="Gekkonidae" genus="Phelsuma" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="laticauda">
<emphasis id="B939EAB2FFB3FFD0D5909789FEDEFB52" box="[204,343,1195,1220]" italics="true" pageId="22" pageNumber="23">P. laticauda</emphasis>
</taxonomicName>
group species (clade K in
<figureCitation id="13762A25FFB3FFD0D7D39788FD5DFB52" box="[655,724,1194,1220]" captionStart="FIGURE 7" captionStartId="17.[151,258,1686,1710]" captionTargetBox="[151,1436,194,1663]" captionTargetId="figure@17.[151,1436,194,1665]" captionTargetPageId="17" captionText="FIGURE 7. Molecular phylogeny of the Phelsuma lineata group (clade J), the P. laticauda group (clade K) and P. klemmeri as calculated from the multi-gene dataset of Rocha et al. (2009). Shown are the respective clades from the Bayesian 50 % majority-rule consensus trees based on (a) the combined dataset of mitochondrial and nuclear genes, (b) the mitochondrial genes only, and (c) the nuclear genes only. See caption to Fig. 2 for further explanations." httpUri="https://zenodo.org/record/194700/files/figure.png" pageId="22" pageNumber="23">Fig. 7</figureCitation>
a) are characterized by being non-gluers with smooth ventral and sub-caudal scales, three nasalia and without nostril-rostral contact. They are mostly bright green coloured and have no lateral or dorsal pattern of stripes (
<tableCitation id="C6CF031BFFB3FFD0D7E397D7FCC9FA99" box="[703,832,1269,1295]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="4.[151,242,151,175]" captionStartId-1="6.[151,239,725,749]" captionTargetBox-0="[151,1437,426,1746]" captionTargetPageId-0="4" captionText-0="TABLE 1. Body size, colour characters and egg-laying behaviour of recognized Phelsuma species and subspecies. Abbreviations: TL, maximum total length; SVL, maximum snout-vent length (both in mm). Colour is coded as follows: green, predominant colour green or blue; grey, predominant colour grey or brown (not green); interm., intermediate or mixed colour. Pattern: strong, contrast of main pattern compared with ground colour strong respectively weak, or intermediate. Stripes: tendency of dorsal or dorsolateral stripes present, absent, or intermediate; elongated patches on head, neck and lower back not considered as stripes. Lateral stripes: dark lateral / sublateral stripes present, absent or weak. Phelsuma can either glue their eggs to the substrate (+) or not (-). Missing data are represented by question marks (?)." captionText-1="TABLE 2. Selected scale characters in species and subspecies of day geckos. Abbreviations: SL, number of supralabials" httpUri-0="http://table.plazi.org/id/DF326628FFA1FFC2D5CB93B5FF32FE19" pageId="22" pageNumber="23" tableUuid-0="DF326628FFA1FFC2D5CB93B5FF32FE19">Tables 12</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>