189 lines
22 KiB
XML
189 lines
22 KiB
XML
<document id="F4DAA95004A85FDCE282DBC144AECD34" ID-CLB-Dataset="6167" ID-DOI="10.11646/zootaxa.4975.2.4" ID-GBIF-Dataset="3d8639f0-544a-4ca5-83a8-b729cf1c3e56" ID-ISSN="1175-5326" ID-Zenodo-Dep="4925219" ID-ZooBank="4F22565A-7089-49FD-9313-63195491B3EA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622022232820" checkinUser="plazi" docAuthor="Henry, R. & Bøggild, Esben" docDate="2021" docId="03E487AEFFDA7B1978B7E502FCA2AA4E" docLanguage="en" docName="zootaxa.4975.2.4.pdf" docOrigin="Zootaxa 4975 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaselia neuter Henry & Bøggild 2021, n. sp." docType="treatment" docVersion="6" lastPageNumber="328" masterDocId="FFDDFFD6FFCE7B0F7820E251FFF9AE78" masterDocTitle="Nineteen new species of Megaselia Rondani (Diptera, Phoridae) from Denmark" masterLastPageNumber="342" masterPageNumber="306" pageNumber="326" updateTime="1698991867495" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2AA92A57F49DB8B20836D21151140059">Nineteen new species of Megaselia Rondani (Diptera, Phoridae) from Denmark</mods:title>
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<mods:name id="6F48591B9E883AF5B3FF3B7A4CFE32AF" type="personal">
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<mods:namePart id="22E8897D0C75F1007E9E9057EB4CCE75">Henry, R.</mods:namePart>
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<mods:affiliation id="7AB5F8C586479C872D6F56015DD1C83B">University Department of Zoology, Downing Street, Cambridge CB 2 3 EJ, UK ..</mods:affiliation>
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<mods:namePart id="E2D293854B1378CF7FEEBDCCC7BC8261">Bøggild, Esben</mods:namePart>
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<mods:affiliation id="7A751AFE9F5F1CCC217F60C212242945">Fayesgade 10, DK- 9500, Hobro, Denmark.</mods:affiliation>
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<mods:title id="AB96E5F44BC0A9082DCEB9E2C6BCDB5B">Zootaxa</mods:title>
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<mods:date id="EED21BDEFD748E4BD862967C6849C9AE">2021</mods:date>
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<mods:number id="C9805E513F6B48A90CF59DB0D5758EDB">2021-05-25</mods:number>
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<treatment id="03E487AEFFDA7B1978B7E502FCA2AA4E" ID-DOI="http://doi.org/10.5281/zenodo.4925340" ID-GBIF-Taxon="182645722" ID-Zenodo-Dep="4925340" LSID="urn:lsid:plazi:treatment:03E487AEFFDA7B1978B7E502FCA2AA4E" httpUri="http://treatment.plazi.org/id/03E487AEFFDA7B1978B7E502FCA2AA4E" lastPageId="22" lastPageNumber="328" pageId="20" pageNumber="326">
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<subSubSection id="C3576533FFDA7B1B78B7E502FEBAA9EB" pageId="20" pageNumber="326" type="nomenclature">
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<paragraph id="8BF236B8FFDA7B1B78B7E502FE4BA916" blockId="20.[151,434,1875,1939]" box="[151,434,1875,1902]" pageId="20" pageNumber="326">
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<heading id="D0BA81D4FFDA7B1B78B7E502FE4BA916" bold="true" box="[151,434,1875,1902]" fontSize="11" level="1" pageId="20" pageNumber="326" reason="1">
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<emphasis id="B939EAAAFFDA7B1B78B7E502FE4BA916" bold="true" box="[151,434,1875,1902]" pageId="20" pageNumber="326">
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<taxonomicName id="4C4D4D3BFFDA7B1B78B7E502FE91A916" ID-CoL="9HX8V" authority="Henry & Bøggild, 2021" authorityName="Henry & Bøggild" authorityYear="2021" box="[151,360,1875,1902]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="326" phylum="Arthropoda" rank="species" species="neuter" status="sp. nov.">
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<emphasis id="B939EAAAFFDA7B1B78B7E502FE91A916" bold="true" box="[151,360,1875,1902]" italics="true" pageId="20" pageNumber="326">Megaselia neuter</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20A57D1FFDA7B1B7950E505FE4BA916" box="[368,434,1876,1902]" pageId="20" pageNumber="326" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF236B8FFDA7B1B78B7E529FEBAA9EB" blockId="20.[151,434,1875,1939]" box="[151,323,1912,1939]" pageId="20" pageNumber="326">(Figs 157–168)</paragraph>
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</subSubSection>
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<subSubSection id="C3576533FFDA7B1978B7E5E8FE70AF8D" lastPageId="22" lastPageNumber="328" pageId="20" pageNumber="326" type="diagnosis">
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<paragraph id="8BF236B8FFDA7B1978B7E5E8FE70AF8D" blockId="20.[151,1436,1977,2040]" lastBlockId="22.[151,1437,150,1078]" lastPageId="22" lastPageNumber="328" pageId="20" pageNumber="326">
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<emphasis id="B939EAAAFFDA7B1B78B7E5E8FEF3A9AB" bold="true" box="[151,266,1977,2003]" pageId="20" pageNumber="326">Diagnosis</emphasis>
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. In the key of
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<bibRefCitation id="EFDC4B49FFDA7B1B798BE5E8FD95A9AB" author="Lundbeck, W." box="[427,620,1977,2003]" pageId="20" pageNumber="326" refId="ref14900" refString="Lundbeck, W. (1922) Diptera Danica Part VI Pipunculidae, Phoridae. Wesley, London, 447 pp." type="book" year="1922">Lundbeck (1922)</bibRefCitation>
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for Group IV it runs to couplet
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<specimenCount id="9D4BFD31FFDA7B1B7BEDE5E8FBF7A9AB" box="[973,1038,1977,2003]" pageId="20" pageNumber="326" type="generic">
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33
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<taxonomicName id="4C4D4D3BFFDA7B1B7BCFE5EBFBF7A9AB" authority="(Wood)" baseAuthorityName="Wood" baseAuthorityYear="1909" box="[1007,1038,1978,2003]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="326" phylum="Arthropoda" rank="species" species="barbulata">
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<emphasis id="B939EAAAFFDA7B1B7BCFE5EBFBF7A9AB" box="[1007,1038,1978,2003]" italics="true" pageId="20" pageNumber="326">M.</emphasis>
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</taxonomicName>
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</specimenCount>
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barbulata (Wood), but it has a very different hypopygium (
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<figureCitation id="13762A3DFFDA7B1B79BBE58CFE24A98F" box="[411,477,2013,2039]" captionStart="FIGURES 1–11" captionStartId="1.[151,264,1943,1967]" captionTargetBox="[271,1312,181,1913]" captionTargetId="figure-31@1.[268,1320,181,1919]" captionTargetPageId="1" captionText="FIGURES 1–11. Megaselia culmenensis n. sp. male. 1, frons; 2, antennae, palps and proboscis; 3, postpedicel; 4, side of thorax; 5, dorsal view of abdominal tergites 2–6; 6; left face of hypopygioum; 7–8, right faces of hypopygium; 9, front tarsus; 10, hind femur; 11, wing." figureDoi="http://doi.org/10.5281/zenodo.4925221" httpUri="https://zenodo.org/record/4925221/files/figure.png" pageId="20" pageNumber="326">
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Fig.
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</figureCitation>
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in
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<bibRefCitation id="EFDC4B49FFDA7B1B7A21E58CFD60A98F" author="Disney, R. H. L." box="[513,665,2013,2039]" pageId="20" pageNumber="326" pagination="93 - 94" refId="ref14059" refString="Disney, R. H. L. (1991) Two rare scuttle flies (Diptera, Phoridae) from Yorkshire, including one new for Britain. Naturalist, Hull, 116, 93 - 94." type="journal article" year="1991">Disney, 1991</bibRefCitation>
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). In the key to the males of the British species (
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<bibRefCitation id="EFDC4B49FFDA7B1B7C8BE58CFABBA98F" author="Disney R. H. L." box="[1195,1346,2013,2039]" pageId="20" pageNumber="326" pagination="1 - 155" refId="ref14009" refString="Disney R. H. L. (1989) Scuttle Flies-Diptera Phoridae Genus Megaselia. Handbooks for the Identification of British Insects 10 (8), Royal Entomologicai Society of London, 1 - 155. https: // doi. org / 10.1163 / 187631287 X 00115" type="book chapter" year="1989">Disney, 1989</bibRefCitation>
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) it runs to couplet 102, but its hypopygium differs from both species. In the keys of
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<bibRefCitation id="EFDC4B49FFD87B197BCBE2C6FB06AEC9" author="Schmitz H. & Beyer" box="[1003,1279,150,177]" pageId="22" pageNumber="328" pagination="513 - 608" refId="ref15062" refString="Schmitz H. & Beyer (1965) Phoridae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region 4 (33) (Lieferung 258, 260). E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart, pp. 513 - 608." type="book chapter" year="1965">Schmitz & Beyer (1965)</bibRefCitation>
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for Abteilung IV Zweite Reihe it runs couplet 71. However, the supposed difference in the number of costal cilia between the two options is negated by reference to their photo of the wing for
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<taxonomicName id="4C4D4D3BFFD87B197B82E28EFB45AE81" authority="Schmitz. The" authorityName="Schmitz. The" authorityYear="1934" box="[930,1212,223,249]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="raetica">
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<emphasis id="B939EAAAFFD87B197B82E28EFBE0AE80" box="[930,1049,223,248]" italics="true" pageId="22" pageNumber="328">M. raetica</emphasis>
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Schmitz. The
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</taxonomicName>
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latter has a clearly different hypopygium from the two species encountered by proceeding to couplet 72. Our species has a hypopygium similar but different from those of
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<taxonomicName id="4C4D4D3BFFD87B197A3CE376FCEFAF39" authority="Schmitz" authorityName="Schmitz" authorityYear="1935" box="[540,790,295,321]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="fusciclava">
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<emphasis id="B939EAAAFFD87B197A3CE376FD4DAF39" box="[540,692,295,321]" italics="true" pageId="22" pageNumber="328">M. fusciclava</emphasis>
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Schmitz
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</taxonomicName>
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and
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<taxonomicName id="4C4D4D3BFFD87B197B6DE376FB80AF3A" authority="(Lundbeck)" baseAuthorityName="Lundbeck" baseAuthorityYear="1920" box="[845,1145,295,322]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="fuscipalpis">
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<emphasis id="B939EAAAFFD87B197B6DE376FC12AF39" box="[845,1003,295,321]" italics="true" pageId="22" pageNumber="328">M. fuscipalpis</emphasis>
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(Lundbeck)
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</taxonomicName>
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. The latter is immediately distinguished by its legs being entirely dark brown.
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<taxonomicName id="4C4D4D3BFFD87B197AF8E31AFC96AF1D" authorityName="Schmitz" authorityYear="1935" box="[728,879,331,357]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="fusciclava">
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<emphasis id="B939EAAAFFD87B197AF8E31AFC96AF1D" box="[728,879,331,357]" italics="true" pageId="22" pageNumber="328">M. fusciclava</emphasis>
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</taxonomicName>
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differs from our species in having a posterodorsal hair palisade on all 5 segments of the front tarsus and its costal index exceeding 0.48. In Bormeier’s (1964) key for Nearctic Group IV species it runs to couplet 17 lead
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<specimenCount id="9D4BFD31FFD87B197AD6E3C2FCD2AFD4" box="[758,811,403,429]" pageId="22" pageNumber="328" type="generic">
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1
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<taxonomicName id="4C4D4D3BFFD87B197B2CE3C2FCD2AFD4" authority="(Malloch)" baseAuthorityName="Malloch" baseAuthorityYear="1912" box="[780,811,403,428]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="difficilis">
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<emphasis id="B939EAAAFFD87B197B2CE3C2FCD2AFD4" box="[780,811,403,428]" italics="true" pageId="22" pageNumber="328">M.</emphasis>
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</taxonomicName>
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</specimenCount>
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difficilis (Malloch), but its wing has 6 axillary bristles and clearly longer costal cilia at
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. It fails to run down in keys for the rest of the world’s fauna and subsequent additions.
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</paragraph>
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</subSubSection>
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<caption id="DF326630FFDB7B1A78B7E5C6FB42A980" ID-DOI="http://doi.org/10.5281/zenodo.4925249" ID-Zenodo-Dep="4925249" httpUri="https://zenodo.org/record/4925249/files/figure.png" pageId="21" pageNumber="327" startId="21.[151,264,1943,1967]" targetBox="[157,1433,187,1908]" targetPageId="21">
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<paragraph id="8BF236B8FFDB7B1A78B7E5C6FB42A980" blockId="21.[151,1437,1943,2040]" pageId="21" pageNumber="327">
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<emphasis id="B939EAAAFFDB7B1A78B7E5C6FDDAA9D7" bold="true" box="[151,547,1943,1967]" pageId="21" pageNumber="327">
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FIGURES 157–168
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<taxonomicName id="4C4D4D3BFFDB7B1A794EE5C6FDDAA9D7" authority="Henry & Bøggild, 2021" authorityName="Henry & Bøggild" authorityYear="2021" box="[366,547,1943,1967]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="327" phylum="Arthropoda" rank="species" species="neuter" status="sp. nov.">
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<emphasis id="B939EAAAFFDB7B1A794EE5C6FDDAA9D7" bold="true" box="[366,547,1943,1967]" italics="true" pageId="21" pageNumber="327">Megaselia neuter</emphasis>
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</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A20A57D1FFDB7B1A7A0AE5C9FD99A9D7" box="[554,608,1944,1967]" pageId="21" pageNumber="327" rank="species">sp. n.</taxonomicNameLabel>
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male. 157, frons; 158, detail of part of frons; 159, antennae, palps and proboscis; 160, proboscis from below; 161, side of thorax; 162, abdomen; 163, left face of hypopygium; 164, left face of epandrium and hypandrium; 165, right face of hypopygium; 166, front tarsus; 167, hind femur; 168, wing; 162, Haltere.
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</paragraph>
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</caption>
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<subSubSection id="C3576533FFD87B1978E7E3AFFCCAADDE" pageId="22" pageNumber="328" type="description">
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<paragraph id="8BF236B8FFD87B1978E7E3AFFCCAADDE" blockId="22.[151,1437,150,1078]" pageId="22" pageNumber="328">
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<emphasis id="B939EAAAFFD87B1978E7E3AFFEA0AC60" bold="true" box="[199,345,510,536]" pageId="22" pageNumber="328">Description.</emphasis>
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Male. Frons as Fig. 157, and lacking fine microtrichia except at front (Fig. 158). Cheek with 3 bristles and jowl with 2 that are longer and more robust. Postpedicels, with a few SPS vesicles, palps and proboscis as Fig. 159. The labella with some but not very many short spinules below (Fig. 160). Thorax brown, with 3 notopleural bristles, with no cleft in front of these, and mesopleuron with hairs (Fig. 161). Scutellum with an anterior pair of small hairs and a posterior pair of bristles. Abdominal tergites and venter as Fig. 162, the latter with hairs on segments 3–6. Hypopygium as Figs 163–165. Legs yellowish but the hind femora yellowish brown. Fore tarsus with posterodorsal hair palisades on segments 1 to 4 (Fig. 166). Mid tibia with dorsal hair palisade about 0.63 times its length. Hairs below basal half of hind femur marginally shorter than those of anteroventral row of outer half (Fig. 167). Hind tibia with about a dozen moderately differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (Fig. 168)
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<quantity id="4CB59B5DFFD87B197AF5E112FCD5AD25" box="[725,812,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="328" unit="mm" value="1.4">1.4 mm</quantity>
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long. Costal index 0.46. Costal ratios 3.4–3.5: 1.6–1.7: 1. Costal cilia (of section 3)
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long. Vein 3 hair
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long. 3 axillary bristles, the outermost being
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long. Sc not reaching R1. Haltere knob pale (Fig. 162).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576533FFD87B1978E7E1FEFF19AD95" pageId="22" pageNumber="328" type="materials_examined">
|
||
<paragraph id="8BF236B8FFD87B1978E7E1FEFF19AD95" blockId="22.[151,1437,150,1078]" pageId="22" pageNumber="328">
|
||
<emphasis id="B939EAAAFFD87B1978E7E1FEFE95ADB0" bold="true" box="[199,364,942,969]" pageId="22" pageNumber="328">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B253CE5FFD87B197958E1FEFF19AD95" ID-GBIF-Occurrence="3128593320" collectingDate="2017-05-21" collectingDateMax="2017-06-05" collectingDateMin="2017-05-21" collectionCode="DK, EJ, UCMZ" country="Denmark" location="Onsild Mose" pageId="22" pageNumber="328" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54F6881AFFD87B197958E1FEFE19ADB1" box="[376,480,943,969]" pageId="22" pageNumber="328" type="holotype">Holotype</typeStatus>
|
||
male,
|
||
<collectingCountry id="F35A7628FFD87B197A07E1FEFD4CADB1" box="[551,693,943,969]" name="Denmark" pageId="22" pageNumber="328">DENMARK</collectingCountry>
|
||
,
|
||
<collectionCode id="ED5CAE7DFFD87B197AE1E1FEFD10ADB1" box="[705,745,943,969]" pageId="22" pageNumber="328">DK</collectionCode>
|
||
,
|
||
<collectionCode id="ED5CAE7DFFD87B197AD5E1FEFCEDADB1" box="[757,788,943,969]" country="Israel" name="Ein Yabrud collection catalogue entries at The Hebrew University" pageId="22" pageNumber="328">EJ</collectionCode>
|
||
,
|
||
<location id="8E926063FFD87B197B3DE1FEFC57ADB1" LSID="urn:lsid:plazi:treatment:03E487AEFFDA7B1978B7E502FCA2AA4E:8E926063FFD87B197B3DE1FEFC57ADB1" box="[797,942,943,969]" country="Denmark" name="Onsild Mose" pageId="22" pageNumber="328">Onsild Mose</location>
|
||
,
|
||
<date id="FFF31078FFD87B197B97E1FFFB8DADB1" box="[951,1140,942,969]" pageId="22" pageNumber="328" value="2017-05-21" valueMax="2017-06-05" valueMin="2017-05-21">
|
||
<collectingDate id="EFB7E990FFD87B197B97E1FFFB8DADB1" box="[951,1140,942,969]" pageId="22" pageNumber="328" value="2017-05-21" valueMax="2017-06-05" valueMin="2017-05-21">21.V.–5.VI.2017</collectingDate>
|
||
</date>
|
||
, Esben Bøggild (
|
||
<collectionCode id="ED5CAE7DFFD87B197D0DE1FEFA86ADB1" box="[1325,1407,943,969]" pageId="22" pageNumber="328">UCMZ</collectionCode>
|
||
— 9–18).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3576533FFD87B1978E7E1A6FCA2AA4E" pageId="22" pageNumber="328" type="etymology">
|
||
<paragraph id="8BF236B8FFD87B1978E7E1A6FCA2AA4E" blockId="22.[151,1437,150,1078]" pageId="22" pageNumber="328">
|
||
<emphasis id="B939EAAAFFD87B1978E7E1A6FEB1AA69" bold="true" box="[199,328,1015,1041]" pageId="22" pageNumber="328">Etymology</emphasis>
|
||
. Named after it being neither one nor the other (
|
||
<taxonomicName id="4C4D4D3BFFD87B197B5BE1A6FC3AAA68" authorityName="Henry & Bøggild" authorityYear="2021" box="[891,963,1015,1040]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="neuter">
|
||
<emphasis id="B939EAAAFFD87B197B5BE1A6FC3AAA68" box="[891,963,1015,1040]" italics="true" pageId="22" pageNumber="328">neuter</emphasis>
|
||
</taxonomicName>
|
||
) with its hypopygium being intermediate between
|
||
<taxonomicName id="4C4D4D3BFFD87B1978DBE64AFE0CAA4D" authority="Schmitz" authorityName="Schmitz" authorityYear="1935" box="[251,501,1051,1077]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="fusciclava">
|
||
<emphasis id="B939EAAAFFD87B1978DBE64AFE6AAA4D" box="[251,403,1051,1077]" italics="true" pageId="22" pageNumber="328">M. fusciclava</emphasis>
|
||
Schmitz
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D3BFFD87B197A0BE64AFCA1AA4E" authority="(Lundbeck)" baseAuthorityName="Lundbeck" baseAuthorityYear="1920" box="[555,856,1051,1078]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="328" phylum="Arthropoda" rank="species" species="fuscipalpis">
|
||
<emphasis id="B939EAAAFFD87B197A0BE64AFD33AA4D" box="[555,714,1051,1077]" italics="true" pageId="22" pageNumber="328">M. fuscipalpis</emphasis>
|
||
(Lundbeck)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |