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<mods:title id="9A11A3DCDDCAA4397C2E96F042B2B5B5">On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala</mods:title>
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<mods:namePart id="8C7852737088EB2D689C5F39F8C8A301">Hari, P. Praved</mods:namePart>
<mods:affiliation id="791BB4C0B70901DB6C970227A7D9F53B">Department of Marine Biology, Microbiology and Biochemistry, School of Marine Sciences, Cochin University of Science and Technology, Cochin 682016 Kerala, India</mods:affiliation>
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<mods:affiliation id="ACE2BFDBBF2B947570AEA7CDA54D193A">Department of Marine Biology, Microbiology and Biochemistry, School of Marine Sciences, Cochin University of Science and Technology, Cochin 682016 Kerala, India &amp; Department of Zoology, N. S. S. Hindu College, Changanassery 686 102 Kerala, India</mods:affiliation>
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<mods:namePart id="A490BB5A64323261EC5447D256F7E5B1">Mendoza, Jose Christopher E.</mods:namePart>
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<taxonomicName id="4C4D4D02FFE5FF97FF214EDFC3E5FE64" ID-CoL="7NPWK" authority="Ng, 2002" authorityName="Ng" authorityYear="2002" box="[151,418,439,465]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FF214EDFC3E5FE64" bold="true" box="[151,418,439,465]" pageId="1" pageNumber="262">
<emphasis id="B939EA93FFE5FF97FF214EDFC36AFE64" bold="true" box="[151,301,439,465]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FE834EDFC3E5FE64" author="Ng, P. K. L." box="[309,418,439,465]" pageId="1" pageNumber="262" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng, 2002</bibRefCitation>
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<paragraph id="8BF23681FFE5FF97FF214E94C6BAFDA1" blockId="1.[151,1277,507,532]" box="[151,1277,507,532]" pageId="1" pageNumber="262">
<treatmentCitationGroup id="AB5D11AFFFE5FF97FF214E94C6BAFDA1" box="[151,1277,507,532]" pageId="1" pageNumber="262">
<treatmentCitation id="0AEC1090FFE5FF97FF214E94C3E5FDA1" author="Ng, P. K. L." box="[151,418,508,532]" page="213" pageId="1" pageNumber="262" year="2002">
<taxonomicName id="4C4D4D02FFE5FF97FF214E94C3E5FDA1" ID-CoL="7NPWK" authority="Ng 2002: 213" authorityName="Ng" authorityPageNumber="213" authorityYear="2002" box="[151,418,508,532]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FF214E94C357FDA6" box="[151,272,508,531]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FEA14E94C3E5FDA1" author="Ng, P. K. L." box="[279,418,508,532]" pageId="1" pageNumber="262" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng 2002: 213</bibRefCitation>
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.—
<treatmentCitation id="0AEC1090FFE5FF97FE774E94C0EFFDA1" author="Ng, P. K. L. &amp; Clark, P. F." box="[449,680,508,532]" page="900" pageId="1" pageNumber="262" year="2008">
<bibRefCitation id="EFDC4B70FFE5FF97FE774E94C0EFFDA1" author="Ng, P. K. L. &amp; Clark, P. F." box="[449,680,508,532]" pageId="1" pageNumber="262" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark 2008: 900</bibRefCitation>
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, 901 (discussion).—
<treatmentCitation id="0AEC1090FFE5FF97FCC04E94C632FDA6" author="Mendoza, J. C. E. &amp; Ng, P. K. L." box="[886,1141,507,532]" page="59" pageId="1" pageNumber="262" year="2011">
<bibRefCitation id="EFDC4B70FFE5FF97FCC04E94C632FDA6" author="Mendoza, J. C. E. &amp; Ng, P. K. L." box="[886,1141,507,532]" pageId="1" pageNumber="262" pagination="58 - 66" refId="ref7639" refString="Mendoza, J. C. E. &amp; Ng, P. K. L. (2011) Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae). Zootaxa, 2809 (1), 58 - 66. https: // doi. org / 10.11646 / zootaxa. 2809.1.5" type="journal article" year="2011">Mendoza &amp; Ng 2011: 59</bibRefCitation>
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(discussion).
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<emphasis id="B939EA93FFE5FF97FF214D2AC375FDEE" bold="true" box="[151,306,577,604]" pageId="1" pageNumber="262">
<typeStatus id="54F68823FFE5FF97FF214D2AC295FDE9" box="[151,210,578,604]" pageId="1" pageNumber="262">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C4D4D02FFE5FF97FE8F4D2AC1F0FDE8" ID-CoL="4HVR8" authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[313,951,578,605]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="quadridentatus" subGenus="Parapilumnus">
<emphasis id="B939EA93FFE5FF97FE8F4D2AC3E4FDE9" box="[313,419,578,604]" italics="true" pageId="1" pageNumber="262">Pilumnus</emphasis>
(
<emphasis id="B939EA93FFE5FF97FE054D2AC015FDE9" box="[435,594,578,604]" italics="true" pageId="1" pageNumber="262">Parapilumnus</emphasis>
)
<emphasis id="B939EA93FFE5FF97FDD44D2AC14BFDE9" box="[610,780,578,604]" italics="true" pageId="1" pageNumber="262">quadridentatus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FCA54D2AC1F0FDE8" author="De Man, J. G." box="[787,951,578,605]" pageId="1" pageNumber="262" pagination="485 - 609" refId="ref7241" refString="De Man, J. G. (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BF23681FFE5FF97FF714D0DC15CFD11" blockId="1.[151,1437,577,2009]" pageId="1" pageNumber="262">
<emphasis id="B939EA93FFE5FF97FF714D0DC3A7FDCA" bold="true" box="[199,480,613,639]" pageId="1" pageNumber="262">Other included species.</emphasis>
<taxonomicName id="4C4D4D02FFE5FF97FE5F4D0EC0FFFD35" ID-CoL="a2549b20-f23f-403a-afc0-e9b90cc72e71" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[489,696,614,640]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE5FF97FE5F4D0EC0FFFD35" box="[489,696,614,640]" italics="true" pageId="1" pageNumber="262">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE5FF97FD764D0EC15BFD35" bold="true" box="[704,796,614,640]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A20A57E8FFE5FF97FD764D0EC15BFD35" box="[704,796,614,640]" pageId="1" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C4D4D02FFE5FF97FC9D4D0EC6FBFD35" authority="(Garth &amp; Kim, 1983)" baseAuthorityName="Garth &amp; Kim" baseAuthorityYear="1983" box="[811,1212,614,640]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="nefissurus">
<emphasis id="B939EA93FFE5FF97FC9D4D0EC1F9FD35" box="[811,958,614,640]" italics="true" pageId="1" pageNumber="262">A. nefissurus</emphasis>
(
<bibRefCitation id="EFDC4B70FFE5FF97FC784D0EC6F4FD35" author="Garth, J. S. &amp; Kim, H. S." box="[974,1203,614,640]" pageId="1" pageNumber="262" pagination="663 - 729" refId="ref7385" refString="Garth, J. S. &amp; Kim, H. S. (1983) Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1909 - 1909. Journal of Natural History, 17, 663 - 729. https: // doi. org / 10.1080 / 00222938300770561" type="journal article" year="1983">Garth &amp; Kim, 1983</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4D4D02FFE5FF97FB7C4D0EC29FFD10" ID-CoL="84FW9" authority="(Deb, 1995)" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" baseAuthorityName="Deb" baseAuthorityYear="1995" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="neolaevis" status="comb. nov.">
<emphasis id="B939EA93FFE5FF97FB7C4D0EC712FD35" box="[1226,1365,614,640]" italics="true" pageId="1" pageNumber="262">A. neolaevis</emphasis>
(
<bibRefCitation id="EFDC4B70FFE5FF97FAD34D0EC297FD10" author="Deb, M." pageId="1" pageNumber="262" pagination="217 - 228" refId="ref7139" refString="Deb, M. (1995) Crustacea: Xanthidae. In: Ghosh, A. K. (Ed.), Hugli Matla Estuary West Bengal. Estuarine Ecosystem Series 2. Zoological Survey of India, Kolkata, pp. 217 - 228." type="book chapter" year="1995">Deb, 1995</bibRefCitation>
)
</taxonomicName>
<emphasis id="B939EA93FFE5FF97FF564DE2C327FD11" bold="true" box="[224,352,650,676]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A20A57E8FFE5FF97FF564DE2C327FD11" box="[224,352,650,676]" pageId="1" pageNumber="262" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C4D4D02FFE5FF97FE2B4DE2C150FD11" ID-CoL="66PXJ" authority="Ng &amp; Clark, 2008" authorityName="Ng &amp; Clark" authorityYear="2008" box="[413,791,650,676]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B939EA93FFE5FF97FE2B4DE2C005FD11" box="[413,578,650,676]" italics="true" pageId="1" pageNumber="262">A. vietnamicus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FDFF4DE2C150FD11" author="Ng, P. K. L. &amp; Clark, P. F." box="[585,791,650,676]" pageId="1" pageNumber="262" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark, 2008</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C357650AFFE5FF97FF714DC5C664FC5D" pageId="1" pageNumber="262" type="diagnosis">
<paragraph id="8BF23681FFE5FF97FF714DC5C0EEFC15" blockId="1.[151,1437,577,2009]" pageId="1" pageNumber="262">
<emphasis id="B939EA93FFE5FF97FF714DC5C305FD72" bold="true" box="[199,322,685,711]" pageId="1" pageNumber="262">Diagnosis.</emphasis>
Adult: Carapace transversely subhexagonal, broader than long; anterolateral margins divided into 4 teeth posterior to exorbital angle, first low, lobiform, second to fourth more projecting. Male thoracic sternum moderately broad, smooth; sternite 2 clearly separated from sternite 3 by transverse suture; small section of sternite 8 sometimes exposed when pleon locked onto thoracic sternum. Chelipeds distinctly unequal. Ambulatory legs without dactylo-propodal lock; posterodistal margin of P5 basis-ischium, with several large, closely packed spines or conical tubercles; posterior margin of P5 merus with spines and/or conical tubercles proximally. G1 sinuous, slender; G2 short, about one-fifth length of G1.
</paragraph>
<paragraph id="8BF23681FFE5FF97FF714CC2C664FC5D" blockId="1.[151,1437,577,2009]" pageId="1" pageNumber="262">Zoea I: Rostral spine much shorter than antennal protopods; antennae extremely long, much longer than carapace; furcae of telson prominently elongate, inner margin armed with spines.</paragraph>
</subSubSection>
<subSubSection id="C357650AFFE5FF94FF714C9AC1C4FF4C" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262" type="discussion">
<paragraph id="8BF23681FFE5FF97FF714C9AC00FFAE5" blockId="1.[151,1437,577,2009]" pageId="1" pageNumber="262">
<emphasis id="B939EA93FFE5FF97FF714C9AC37CFBB9" bold="true" box="[199,315,1010,1036]" pageId="1" pageNumber="262">Remarks.</emphasis>
<taxonomicName id="4C4D4D02FFE5FF97FEF54C9AC07AFBB9" authority="Ng, 2002" authorityName="Ng" authorityYear="2002" box="[323,573,1010,1036]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FEF54C9AC38EFBB9" box="[323,457,1010,1036]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FE674C9AC07AFBB9" author="Ng, P. K. L." box="[465,573,1010,1036]" pageId="1" pageNumber="262" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng, 2002</bibRefCitation>
</taxonomicName>
, has primarily been diagnosed by the condition of the fourth ambulatory leg (P5) where the basis-ischium has a comb-like postero-distal margin due to the peculiar placement of some spines or conical tubercles, and the posterior margin of the merus is armed with spines and/or conical tubercles proximally (
<bibRefCitation id="EFDC4B70FFE5FF97FF294B36C340FBCD" author="Ng, P. K. L." box="[159,263,1118,1144]" pageId="1" pageNumber="262" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng 2002</bibRefCitation>
;
<bibRefCitation id="EFDC4B70FFE5FF97FEA54B36C3A7FBCD" author="Ng, P. K. L. &amp; Clark, P. F." box="[275,480,1118,1144]" pageId="1" pageNumber="262" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark 2008</bibRefCitation>
). Subsequent to the establishment of the genus,
<bibRefCitation id="EFDC4B70FFE5FF97FBB54B36C69AFBCD" author="Ng, P. K. L. &amp; Clark, P. F." box="[1027,1245,1118,1144]" pageId="1" pageNumber="262" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark (2008)</bibRefCitation>
added two more species,
<taxonomicName id="4C4D4D02FFE5FF97FF414BEAC03BFB29" authority="(Garth &amp; Kim 1983)" baseAuthorityName="Garth &amp; Kim" baseAuthorityYear="1983" box="[247,636,1154,1180]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="nefissurus">
<emphasis id="B939EA93FFE5FF97FF414BEAC3CEFB29" box="[247,393,1154,1180]" italics="true" pageId="1" pageNumber="262">A. nefissurus</emphasis>
(
<bibRefCitation id="EFDC4B70FFE5FF97FE2F4BEAC034FB29" author="Garth, J. S. &amp; Kim, H. S." box="[409,627,1154,1180]" pageId="1" pageNumber="262" pagination="663 - 729" refId="ref7385" refString="Garth, J. S. &amp; Kim, H. S. (1983) Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1909 - 1909. Journal of Natural History, 17, 663 - 729. https: // doi. org / 10.1080 / 00222938300770561" type="journal article" year="1983">Garth &amp; Kim 1983</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C4D4D02FFE5FF97FD024BEAC673FB29" authority="Ng &amp; Clark, 2008" authorityName="Ng &amp; Clark" authorityYear="2008" box="[692,1076,1154,1180]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B939EA93FFE5FF97FD024BEAC11CFB29" box="[692,859,1154,1180]" italics="true" pageId="1" pageNumber="262">A. vietnamicus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FCD44BEAC673FB29" author="Ng, P. K. L. &amp; Clark, P. F." box="[866,1076,1154,1180]" pageId="1" pageNumber="262" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark, 2008</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="EFDC4B70FFE5FF97FB894BEAC75CFB29" author="Ghory, F. &amp; Ng, P. K. L. &amp; Kazmi, Q. B." box="[1087,1307,1154,1180]" pageId="1" pageNumber="262" pagination="301 - 312" refId="ref7458" refString="Ghory, F., Ng, P. K. L. &amp; Kazmi, Q. B. (2013) On the identities of Pilumnopeus convexus Maccgno, 1936, and P. salomonensis Ward, 1942, from the Indian Ocean, with a note on P. riui Takeda, 2001 (Decapoda, Brachyura, Pilumnidae). Crustaceana, 86 (3), 301 - 312. https: // doi. org / 10.1163 / 15685403 - 00003180" type="journal article" year="2013">
Ghory
<emphasis id="B939EA93FFE5FF97FB394BEBC686FB29" box="[1167,1217,1154,1180]" italics="true" pageId="1" pageNumber="262">et al</emphasis>
. (2013)
</bibRefCitation>
considered
<taxonomicName id="4C4D4D02FFE5FF97FF214BCEC3BAFB75" authority="Takeda, 2001" authorityName="Takeda" authorityYear="2001" box="[151,509,1190,1216]" class="Malacostraca" family="Pilumnidae" genus="Pilumnopeus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="riui">
<emphasis id="B939EA93FFE5FF97FF214BCEC31DFB75" box="[151,346,1190,1216]" italics="true" pageId="1" pageNumber="262">Pilumnopeus riui</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FED44BCEC3BAFB75" author="Takeda, M." box="[354,509,1190,1216]" pageId="1" pageNumber="262" pagination="87 - 93" refId="ref7918" refString="Takeda, M. (2001) On two estuarine crabs from Hainan Island, South China Sea. In: Matsuura, K. (Ed.), Marine fauna of the shallow waters around Hainan Island, South China Sea. National Science Museum Monographs, 21, 87 - 93." type="journal article" year="2001">Takeda, 2001</bibRefCitation>
</taxonomicName>
, described from
<collectingRegion id="4989F863FFE5FF97FD0A4BCEC14AFB75" box="[700,781,1190,1216]" country="China" name="Hainan" pageId="1" pageNumber="262">Hainan</collectingRegion>
Island,
<collectingCountry id="F35A7611FFE5FF97FCDC4BCEC1F7FB75" box="[874,944,1190,1216]" name="China" pageId="1" pageNumber="262">China</collectingCountry>
, as a junior synonym of
<taxonomicName id="4C4D4D02FFE5FF97FB7D4BCEC7D1FB75" authorityName="De Man" authorityYear="1895" box="[1227,1430,1190,1216]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFE5FF97FB7D4BCEC7D1FB75" box="[1227,1430,1190,1216]" italics="true" pageId="1" pageNumber="262">A. quadridentatus</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFDC4B70FFE5FF97FF214BA2C3CEFB51" author="Trivedi, J. &amp; Mitra, S. &amp; Patel, P. &amp; Maheta, N. &amp; Patel, K. &amp; Ng, P. K. L." box="[151,393,1226,1252]" pageId="1" pageNumber="262" pagination="1 - 18" refId="ref7973" refString="Trivedi, J., Mitra, S., Patel, P., Maheta, N., Patel, K. &amp; Ng, P. K. L. (2021) On the Indian species of Eurycarcinus A. MilneEdwards, 1867, Heteropanope Stimpson, 1858, and Pilumnopeus A. Milne-Edwards, 1867 (Decapoda: Brachyura: Pilumnidae). Nauplius, 29 (e 2021004), 1 - 18. https: // doi. org / 10.1590 / 2358 - 2936 e 2021004" type="journal article" year="2021">
Trivedi
<emphasis id="B939EA93FFE5FF97FF434BA3C375FB51" box="[245,306,1226,1252]" italics="true" pageId="1" pageNumber="262">et al.</emphasis>
(2021)
</bibRefCitation>
reported
<taxonomicName id="4C4D4D02FFE5FF97FE494BA2C08AFB51" authorityName="De Man" authorityYear="1895" box="[511,717,1226,1252]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFE5FF97FE494BA2C08AFB51" box="[511,717,1226,1252]" italics="true" pageId="1" pageNumber="262">A. quadridentatus</emphasis>
</taxonomicName>
in the Indian Ocean (
<collectingRegion id="4989F863FFE5FF97FC6E4BA2C637FB51" box="[984,1136,1226,1252]" country="India" name="West Bengal" pageId="1" pageNumber="262">West Bengal</collectingRegion>
,
<collectingCountry id="F35A7611FFE5FF97FB364BA2C6FCFB51" box="[1152,1211,1226,1252]" name="India" pageId="1" pageNumber="262">India</collectingCountry>
) by virtue of their considering it to be a senior synonym of
<taxonomicName id="4C4D4D02FFE5FF97FDD04B87C134FABC" authority="Deb, 1985" authorityName="Deb" authorityYear="1985" box="[614,883,1262,1289]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="neolaevis">
<emphasis id="B939EA93FFE5FF97FDD04B87C0B4FABD" box="[614,755,1262,1288]" italics="true" pageId="1" pageNumber="262">H. neolaevis</emphasis>
Deb, 1985
</taxonomicName>
. The identity of
<taxonomicName id="4C4D4D02FFE5FF97FB984B87C6FCFABD" authorityName="Deb" authorityYear="1985" box="[1070,1211,1262,1288]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="neolaevis">
<emphasis id="B939EA93FFE5FF97FB984B87C6FCFABD" box="[1070,1211,1262,1288]" italics="true" pageId="1" pageNumber="262">H. neolaevis</emphasis>
</taxonomicName>
is further discussed herein. Finally,
<bibRefCitation id="EFDC4B70FFE5FF97FEF04A7AC0CCFA99" author="Widyastuti, E. &amp; Rahayu, D. L." box="[326,651,1298,1324]" pageId="1" pageNumber="262" pagination="15 - 22" refId="ref8067" refString="Widyastuti, E. &amp; Rahayu, D. L. (2022) The new record of Heteropilumnus sasekumari (Serene, 1971) and Aniptumnus quadridentatus (De Man, 1895) (Crustacea, Decapoda, Brachyura, Pilumnidae) from mangrove habitat in Papua, Indonesia. Oseanologi dan Limnologi di Indonesia, 7 (1), 15 - 22." type="journal article" year="2022">Widyastuti &amp; Rahayu (2022)</bibRefCitation>
recently reported a range extension for
<taxonomicName id="4C4D4D02FFE5FF97FBFC4A7AC753FA99" authorityName="De Man" authorityYear="1895" box="[1098,1300,1298,1324]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFE5FF97FBFC4A7AC753FA99" box="[1098,1300,1298,1324]" italics="true" pageId="1" pageNumber="262">A. quadridentatus</emphasis>
</taxonomicName>
, with a new record from Timika,
<collectingRegion id="4989F863FFE5FF97FE374A5EC38FFAE5" box="[385,456,1334,1360]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="262">Papua</collectingRegion>
,
<collectingCountry id="F35A7611FFE5FF97FE654A5EC003FAE5" box="[467,580,1334,1360]" name="Indonesia" pageId="1" pageNumber="262">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="8BF23681FFE5FF97FF714A32C01DF9F9" blockId="1.[151,1437,577,2009]" pageId="1" pageNumber="262">
It is important to note, however, that
<taxonomicName id="4C4D4D02FFE5FF97FDD14A32C0BFFAC1" baseAuthorityName="Garth &amp; Kim" baseAuthorityYear="1983" box="[615,760,1370,1396]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="nefissurus">
<emphasis id="B939EA93FFE5FF97FDD14A32C0BFFAC1" box="[615,760,1370,1396]" italics="true" pageId="1" pageNumber="262">A. nefissurus</emphasis>
</taxonomicName>
is atypical in apparently lacking the comb-like posterodistal margin on the P5 basis-ischium, and both
<taxonomicName id="4C4D4D02FFE5FF97FDC24A16C141FA2D" baseAuthorityName="Garth &amp; Kim" baseAuthorityYear="1983" box="[628,774,1406,1432]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="nefissurus">
<emphasis id="B939EA93FFE5FF97FDC24A16C141FA2D" box="[628,774,1406,1432]" italics="true" pageId="1" pageNumber="262">A. nefissurus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D02FFE5FF97FC884A16C1A3FA2D" authorityName="Ng &amp; Clark" authorityYear="2008" box="[830,996,1406,1432]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B939EA93FFE5FF97FC884A16C1A3FA2D" box="[830,996,1406,1432]" italics="true" pageId="1" pageNumber="262">A. vietnamicus</emphasis>
</taxonomicName>
have the distal tip of their G1 tapering to a point (but with varying degrees of curvature), which is typical of many pilumnines, whereas in the
<typeStatus id="54F68823FFE5FF97FAA34ACAC702FA09" box="[1301,1349,1442,1468]" pageId="1" pageNumber="262">type</typeStatus>
species and
<taxonomicName id="4C4D4D02FFE5FF97FF704AAEC363FA55" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[198,292,1478,1504]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE5FF97FF704AAEC363FA55" box="[198,292,1478,1504]" italics="true" pageId="1" pageNumber="262">A. bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE5FF97FE9C4AAEC3C5FA55" bold="true" box="[298,386,1478,1504]" pageId="1" pageNumber="262">
<taxonomicNameLabel id="A20A57E8FFE5FF97FE9C4AAEC3C5FA55" box="[298,386,1478,1504]" pageId="1" pageNumber="262" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the distal tip of the G1 has an apical lobe that is bluntly round and scoop-shaped. The position of
<taxonomicName id="4C4D4D02FFE5FF97FF0C4A82C308F9B1" baseAuthorityName="Garth &amp; Kim" baseAuthorityYear="1983" box="[186,335,1514,1540]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="nefissurus">
<emphasis id="B939EA93FFE5FF97FF0C4A82C308F9B1" box="[186,335,1514,1540]" italics="true" pageId="1" pageNumber="262">A. nefissurus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D02FFE5FF97FE3B4A82C071F9B1" authorityName="Ng &amp; Clark" authorityYear="2008" box="[397,566,1514,1540]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B939EA93FFE5FF97FE3B4A82C071F9B1" box="[397,566,1514,1540]" italics="true" pageId="1" pageNumber="262">A. vietnamicus</emphasis>
</taxonomicName>
in
<taxonomicName id="4C4D4D02FFE5FF97FDD44A82C0AFF9B1" authorityName="Ng" authorityYear="2002" box="[610,744,1514,1540]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FDD44A82C0AFF9B1" box="[610,744,1514,1540]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
will have to be evaluated further. Also, given that only
<taxonomicName id="4C4D4D02FFE5FF97FA324A82C306F99D" authorityName="De Man" authorityYear="1895" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFE5FF97FA324A82C306F99D" italics="true" pageId="1" pageNumber="262">A. quadridentatus</emphasis>
</taxonomicName>
has the first zoea morphology known, it will be critical to also determine the zoeal morphology of the other species of
<taxonomicName id="4C4D4D02FFE5FF97FEC1495AC3BAF9F9" authorityName="Ng" authorityYear="2002" box="[375,509,1586,1612]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FEC1495AC3BAF9F9" box="[375,509,1586,1612]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
as well.
</paragraph>
<paragraph id="8BF23681FFE5FF94FF71493EC1C4FF4C" blockId="1.[151,1437,577,2009]" lastBlockId="2.[151,1437,150,249]" lastPageId="2" lastPageNumber="263" pageId="1" pageNumber="262">
The systematic position of
<taxonomicName id="4C4D4D02FFE5FF97FE49493EC0C2F9C5" authorityName="Ng" authorityYear="2002" box="[511,645,1622,1648]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FE49493EC0C2F9C5" box="[511,645,1622,1648]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
within
<taxonomicName id="4C4D4D02FFE5FF97FD69493EC119F9C5" authorityName="Samouelle" authorityYear="1819" box="[735,862,1622,1648]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="family">Pilumnidae</taxonomicName>
/
<taxonomicName id="4C4D4D02FFE5FF97FCD1493EC1A1F9C5" authorityName="Samouelle" authorityYear="1819" box="[871,998,1622,1648]" class="Malacostraca" family="Pilumnidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="subFamily" subFamily="Pilumninae">Pilumninae</taxonomicName>
is also an interesting issue to resolve. It is, thus far, the only pilumnid reported to have such long antennae on the first zoea, although another pilumnid genus,
<taxonomicName id="4C4D4D02FFE5FF97FF5D49F6C035F90D" authority="Mendoza &amp; Ng, 2011" authorityName="Mendoza &amp; Ng" authorityYear="2011" box="[235,626,1694,1720]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FF5D49F6C323F90D" box="[235,356,1694,1720]" italics="true" pageId="1" pageNumber="262">Leelumnus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FED949F6C035F90D" author="Mendoza, J. C. E. &amp; Ng, P. K. L." box="[367,626,1694,1720]" pageId="1" pageNumber="262" pagination="58 - 66" refId="ref7639" refString="Mendoza, J. C. E. &amp; Ng, P. K. L. (2011) Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae). Zootaxa, 2809 (1), 58 - 66. https: // doi. org / 10.11646 / zootaxa. 2809.1.5" type="journal article" year="2011">Mendoza &amp; Ng, 2011</bibRefCitation>
</taxonomicName>
, also has a similar feature (Mendoza unpubl. data). In addition, both
<taxonomicName id="4C4D4D02FFE5FF97FF2149AAC35AF969" authorityName="Ng" authorityYear="2002" box="[151,285,1730,1756]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FF2149AAC35AF969" box="[151,285,1730,1756]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4D4D02FFE5FF97FEE349AAC389F969" authorityName="Mendoza &amp; Ng" authorityYear="2011" box="[341,462,1730,1756]" class="Malacostraca" family="Pilumnidae" genus="Leelumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FEE349AAC389F969" box="[341,462,1730,1756]" italics="true" pageId="1" pageNumber="262">Leelumnus</emphasis>
</taxonomicName>
appear to favor estuarine areas and are a common component of fouling communities there (
<bibRefCitation id="EFDC4B70FFE5FF97FF68498EC397F8B5" author="Mendoza, J. C. E. &amp; Ng, P. K. L." box="[222,464,1766,1792]" pageId="1" pageNumber="262" pagination="58 - 66" refId="ref7639" refString="Mendoza, J. C. E. &amp; Ng, P. K. L. (2011) Leelumnus radium, a new genus and species of pilumnid crab from marine encrusting communities in Singapore (Crustacea: Decapoda: Brachyura: Pilumnidae). Zootaxa, 2809 (1), 58 - 66. https: // doi. org / 10.11646 / zootaxa. 2809.1.5" type="journal article" year="2011">Mendoza &amp; Ng 2011</bibRefCitation>
). Another genus,
<taxonomicName id="4C4D4D02FFE5FF97FD25498EC63AF8B5" authority="Hsueh, Huang &amp; Ng, 2009" authorityName="Hsueh, Huang &amp; Ng" authorityYear="2009" box="[659,1149,1766,1792]" class="Malacostraca" family="Pilumnidae" genus="Cryptopilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FD25498EC100F8B5" box="[659,839,1766,1792]" italics="true" pageId="1" pageNumber="262">Cryptopilumnus</emphasis>
<bibRefCitation id="EFDC4B70FFE5FF97FCF8498EC63AF8B5" author="Hsueh, P. - W. &amp; Huang, J. - F. &amp; Ng, P. K. L." box="[846,1149,1766,1792]" pageId="1" pageNumber="262" pagination="323 - 334" refId="ref7545" refString="Hsueh, P. - W., Huang, J. - F. &amp; Ng, P. K. L. (2009) On a new genus and new species of pilumnid crab from Taiwan, and the generic placements of Heteropanope changensis (Rathbun, 1909) and Pilumnopeus pereiodontus Davie and Ghani, 1993 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 43 (5 - 6), 323 - 334. https: // doi. org / 10.1080 / 00222930802585802" type="journal article" year="2009">Hsueh, Huang &amp; Ng, 2009</bibRefCitation>
</taxonomicName>
, has a similar granulation pattern on the P5 basis-ischium and merus, although it can be separated from
<taxonomicName id="4C4D4D02FFE5FF97FB974862C6E0F891" authorityName="Ng" authorityYear="2002" box="[1057,1191,1802,1828]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FB974862C6E0F891" box="[1057,1191,1802,1828]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
by the presence of a dactylo-propodal lock on the ambulatory legs, the complete fusion of thoracic sternites 13, and the predisposition to live in holes and crevices within rocks (
<bibRefCitation id="EFDC4B70FFE5FF97FDDD483AC172F8D9" author="Hsueh, P. - W. &amp; Huang, J. - F. &amp; Ng, P. K. L." box="[619,821,1874,1900]" pageId="1" pageNumber="262" pagination="323 - 334" refId="ref7545" refString="Hsueh, P. - W., Huang, J. - F. &amp; Ng, P. K. L. (2009) On a new genus and new species of pilumnid crab from Taiwan, and the generic placements of Heteropanope changensis (Rathbun, 1909) and Pilumnopeus pereiodontus Davie and Ghani, 1993 (Crustacea: Decapoda: Brachyura). Journal of Natural History, 43 (5 - 6), 323 - 334. https: // doi. org / 10.1080 / 00222930802585802" type="journal article" year="2009">
Hsueh
<emphasis id="B939EA93FFE5FF97FD0D483BC0B4F8D9" box="[699,755,1874,1900]" italics="true" pageId="1" pageNumber="262">et al.</emphasis>
2009
</bibRefCitation>
), whereas
<taxonomicName id="4C4D4D02FFE5FF97FC1C483AC677F8D9" authorityName="Ng" authorityYear="2002" box="[938,1072,1874,1900]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FC1C483AC677F8D9" box="[938,1072,1874,1900]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
has no dactylo-propodal lock on the ambulatory legs, has fused thoracic sternites 1 and 2 but with sternite 3 separated from sternite 2 by a distinct suture, and it has not been recorded to live within rocks.
<taxonomicName id="4C4D4D02FFE5FF97FC8248F2C1FDF801" authorityName="Ng" authorityYear="2002" box="[820,954,1946,1972]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FC8248F2C1FDF801" box="[820,954,1946,1972]" italics="true" pageId="1" pageNumber="262">Aniptumnus</emphasis>
</taxonomicName>
has also been compared to
<taxonomicName id="4C4D4D02FFE5FF97FAB448F2C3B2F86C" authority="Turkay &amp; Schuhmacher, 1985" authorityName="Turkay &amp; Schuhmacher" authorityYear="1985" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE5FF97FAB448F2C7DBF801" box="[1282,1436,1946,1972]" italics="true" pageId="1" pageNumber="262">Latopilumnus</emphasis>
Türkay &amp; Schuhmacher, 1985
</taxonomicName>
, primarily due to similarities in the general form of the carapace and pereopods, although in the case of
<taxonomicName id="4C4D4D02FFE6FF94FE2E4FFFC075FF04" authorityName="Turkay &amp; Schuhmacher" authorityYear="1985" box="[408,562,151,177]" class="Malacostraca" family="Pilumnidae" genus="Latopilumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE6FF94FE2E4FFFC075FF04" box="[408,562,151,177]" italics="true" pageId="2" pageNumber="263">Latopilumnus</emphasis>
</taxonomicName>
, the spines and conical tubercles on the P5 basis-ischium and merus are absent and the first zoea, which has short antennae, is typical of the family (
<bibRefCitation id="EFDC4B70FFE6FF94FC284FD3C62EFF60" author="Ng, P. K. L. &amp; Clark, P. F." box="[926,1129,187,213]" pageId="2" pageNumber="263" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark 2008</bibRefCitation>
). These genera will have to be included in future phylogenetic studies concerning
<taxonomicName id="4C4D4D02FFE6FF94FD404FB7C13BFF4C" authorityName="Ng" authorityYear="2002" box="[758,892,223,249]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
<emphasis id="B939EA93FFE6FF94FD404FB7C13BFF4C" box="[758,892,223,249]" italics="true" pageId="2" pageNumber="263">Aniptumnus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>