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<document id="8C62A811C28F5AD51962BF47F4908A18" ID-DOI="10.11646/zootaxa.5297.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8004996" ID-ZooBank="ABF87F77-6C54-4460-919E-78D67CFAB176" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685959583509" checkinUser="plazi" docAuthor="Mikhaljova, Elena V." docDate="2023" docId="03E48797B823F71264CF1D0361D9599C" docLanguage="en" docName="zootaxa.5297.3.3.pdf" docOrigin="Zootaxa 5297 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5297.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nepalmatoiulus alternus Mikhaljova 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="385" masterDocId="FFDDFFEFB822F71764581F216535584D" masterDocTitle="Descriptions of the new species of the genus Nepalmatoiulus Mauriès, 1983 from China, with notes on the closely coexisting species (Diplopoda, Julida, Julidae)" masterLastPageNumber="392" masterPageNumber="380" pageNumber="381" updateTime="1685960003826" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1077ACE59E09C8328B5FB9AF379B5000">Descriptions of the new species of the genus Nepalmatoiulus Mauriès, 1983 from China, with notes on the closely coexisting species (Diplopoda, Julida, Julidae)</mods:title>
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<mods:namePart id="FEE7F877D439B772292D5BFBAA23AFF6">Mikhaljova, Elena V.</mods:namePart>
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<mods:title id="1714748B29FE6DFDD302C9190CD07689">Zootaxa</mods:title>
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<mods:date id="8926483B9A2D7C010FA533C77055D780">2023</mods:date>
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<mods:number id="1D353B95E3565ABAC2E0A31F1BDAFFCF">2023-06-02</mods:number>
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<mods:identifier id="9C96DFFD2B44B348B98795ACFEA75911" type="DOI">10.11646/zootaxa.5297.3.3</mods:identifier>
<mods:identifier id="220697EF63209E3DAD9B425E71938B17" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E48797B823F71264CF1D0361D9599C" ID-DOI="http://doi.org/10.5281/zenodo.8004998" ID-Zenodo-Dep="8004998" LSID="urn:lsid:plazi:treatment:03E48797B823F71264CF1D0361D9599C" httpUri="http://treatment.plazi.org/id/03E48797B823F71264CF1D0361D9599C" lastPageId="5" lastPageNumber="385" pageId="1" pageNumber="381">
<subSubSection id="C357650AB823F71664CF1D0367125A70" box="[151,551,546,573]" pageId="1" pageNumber="381" type="nomenclature">
<paragraph id="8BF23681B823F71664CF1D0367125A70" blockId="1.[151,551,546,609]" box="[151,551,546,573]" pageId="1" pageNumber="381">
<heading id="D0BA81EDB823F71664CF1D0367125A70" bold="true" box="[151,551,546,573]" fontSize="11" level="1" pageId="1" pageNumber="381" reason="1">
<emphasis id="B939EA93B823F71664CF1D0367125A70" bold="true" box="[151,551,546,573]" pageId="1" pageNumber="381">
<taxonomicName id="4C4D4D02B823F71664CF1D0364F75A70" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[151,450,546,573]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="381" phylum="Arthropoda" rank="species" species="alternus" status="sp. nov.">
<emphasis id="B939EA93B823F71664CF1D0364F75A70" bold="true" box="[151,450,546,573]" italics="true" pageId="1" pageNumber="381">Nepalmatoiulus alternus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20A57E8B823F71665921D0267125A70" box="[458,551,547,573]" pageId="1" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C357650AB823F71664CF1D6664055A2C" box="[151,304,583,609]" pageId="1" pageNumber="381" type="description">
<paragraph id="8BF23681B823F71664CF1D6664055A2C" blockId="1.[151,551,546,609]" box="[151,304,583,609]" pageId="1" pageNumber="381">
<figureCitation id="13762A04B823F71664CF1D66643D5A2C" box="[151,264,583,609]" captionStart-0="FIGURES 15" captionStart-1="FIGURES 610" captionStart-2="FIGURES 1114" captionStartId-0="2.[152,270,1732,1756]" captionStartId-1="3.[152,270,1823,1847]" captionStartId-2="4.[152,270,1725,1749]" captionTargetBox-0="[151,1436,192,1686]" captionTargetBox-1="[151,1436,199,1786]" captionTargetBox-2="[154,1434,181,1693]" captionTargetId-0="figure-15@2.[151,1436,192,1686]" captionTargetId-1="figure-33@3.[151,1436,199,1786]" captionTargetId-2="figure-15@4.[154,1434,181,1701]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 15. Nepalmatoiulus alternus sp. nov., male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes. 1. Leg pair 1, anterior view. 2. Leg 2, anterior view (mesapical oral seta broken off). 3. Coxae 2, anterior view. 4. Claw 2, anterior view. 5. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5)." captionText-1="FIGURES 610. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 6. Penis, caudal view. 7. Left promere, caudal view. 8. Right promere, caudal view. 9. Distal part of left promere, caudal view. 10. Distal part of right promere, caudal view. Abbreviations: a, protrusion; la, lateral angle. Scales: 20 μm (Figs 6, 910), 100 μm (Figs 78)." captionText-2="FIGURES 1114. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 11. Opisthomere, mesal view. 12. Anterior part of opisthomere, mesal view. 13. Solenomere. 14. Opisthomere, anteromesal view. Abbreviations: bb, basal blade of solenomere; d, outgrowth of mesomeral process; ex, excavation; k, knobs. Scales: 10 μm (Figs 1213), 100 μm (Fig. 11), 0.5 mm (Fig. 14)." figureDoi-0="http://doi.org/10.5281/zenodo.8005000" figureDoi-1="http://doi.org/10.5281/zenodo.8005002" figureDoi-2="http://doi.org/10.5281/zenodo.8005006" httpUri-0="https://zenodo.org/record/8005000/files/figure.png" httpUri-1="https://zenodo.org/record/8005002/files/figure.png" httpUri-2="https://zenodo.org/record/8005006/files/figure.png" pageId="1" pageNumber="381">Figs 114</figureCitation>
,
<figureCitation id="13762A04B823F716654C1D6664055A2C" box="[276,304,583,609]" captionStart="FIGURE 39" captionStartId="12.[152,255,852,876]" captionTargetBox="[225,1362,187,822]" captionTargetId="figure-14@12.[219,1369,181,828]" captionTargetPageId="12" captionText="FIGURE 39. Size relations of coexisting Nepalmatoiulus alternus sp. nov. (continuous lines, square - ♁, diamond - ♀) and Nepalmatoiulus simultaneus sp. nov. (stippled lines, heart - ♁, oval - ♀). H, midbody vertical diameter (mm), p.s., number of podous rings." figureDoi="http://doi.org/10.5281/zenodo.8005020" httpUri="https://zenodo.org/record/8005020/files/figure.png" pageId="1" pageNumber="381">39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C357650AB823F71664CF1DA964885B43" pageId="1" pageNumber="381" type="materials_examined">
<paragraph id="8BF23681B823F71664CF1DA964885B43" blockId="1.[151,1437,648,2043]" pageId="1" pageNumber="381">
<emphasis id="B939EA93B823F71664CF1DA964B15AEF" bold="true" box="[151,388,648,674]" pageId="1" pageNumber="381">Material examined.</emphasis>
<materialsCitation id="3B253CDCB823F71665D71DA967705AA7" collectingDate="2014-05-06" collectionCode="ZMUM" collectorName="Laojunshan E &amp; I. Belousov &amp; Belousov, I." country="China" elevation="4075" latitude="26.882778" location="Laotzyunshan Mt." longLatPrecision="20" longitude="99.62889" pageId="1" pageNumber="381" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54F68823B823F71665D71DA964C95AEF" box="[399,508,648,674]" pageId="1" pageNumber="381">Holotype</typeStatus>
: male (
<collectionCode id="ED5CAE44B823F716660A1DA9679A5AEF" box="[594,687,648,674]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="381">ZMUM</collectionCode>
),
<collectingCountry id="F35A7611B823F716669E1DA966395AEF" box="[710,780,648,674]" name="China" pageId="1" pageNumber="381">China</collectingCountry>
,
<collectingRegion id="4989F863B823F71667411DA866EA5AEF" box="[793,991,648,675]" country="China" name="Yunnan" pageId="1" pageNumber="381">Yunnan Province</collectingRegion>
,
<collectorName id="26B85357B823F71667B71DA961B85AEE" box="[1007,1165,648,675]" pageId="1" pageNumber="381">Laojunshan E</collectorName>
slope of
<location id="8E92605AB823F71660A71DA965F75A8A" LSID="urn:lsid:plazi:treatment:03E48797B823F71264CF1D0361D9599C:8E92605AB823F71660A71DA965F75A8A" country="China" latitude="26.882778" longLatPrecision="20" longitude="99.62889" name="Laotzyunshan Mt." pageId="1" pageNumber="381" stateProvince="Yunnan">Laotzyunshan Mt.</location>
,
<geoCoordinate id="EE795046B823F71664951D8D64535A8A" box="[205,358,684,711]" degrees="26" direction="north" minutes="52" orientation="latitude" pageId="1" pageNumber="381" precision="15" seconds="58" value="26.882778">26°52´58´´ N</geoCoordinate>
,
<geoCoordinate id="EE795046B823F71665291D8D67335A8A" box="[369,518,684,711]" degrees="99" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="381" precision="15" seconds="44" value="99.62889">99°37´44´´ E</geoCoordinate>
, H =
<quantity id="4CB59B64B823F716661C1D8D67AE5A8A" box="[580,667,684,711]" metricMagnitude="3" metricUnit="m" metricValue="4.075" pageId="1" pageNumber="381" unit="m" value="4075.0">
<elevation id="0060D1B2B823F716661C1D8D67AE5A8A" box="[580,667,684,711]" metricMagnitude="3" metricUnit="m" metricValue="4.075" pageId="1" pageNumber="381" unit="m" value="4075.0">4075 m</elevation>
</quantity>
, crooked
<taxonomicName id="4C4D4D02B823F71667501D8D66985A8B" box="[776,941,684,710]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="381" phylum="Tracheophyta" rank="genus">
<emphasis id="B939EA93B823F71667501D8D66985A8B" box="[776,941,684,710]" italics="true" pageId="1" pageNumber="381">Rhododendron</emphasis>
</taxonomicName>
forest, wet litter + numerous streams, sifter,
<date id="FFF31041B823F71664CF1DF1642D5AA6" box="[151,280,720,747]" pageId="1" pageNumber="381" value="2014-05-06">
<collectingDate id="EFB7E9A9B823F71664CF1DF1642D5AA6" box="[151,280,720,747]" pageId="1" pageNumber="381" value="2014-05-06">05.06.2014</collectingDate>
</date>
, leg.
<collectorName id="26B85357B823F716650A1DF064E65AA7" box="[338,467,720,747]" pageId="1" pageNumber="381">I. Belousov</collectorName>
, I. Kabak.
</materialsCitation>
<materialsCitation id="3B253CDCB823F71666121DF061515AA7" box="[586,1124,720,747]" collectionCode="ZMUM" country="China" pageId="1" pageNumber="381" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="54F68823B823F71666121DF0678E5AA6" box="[586,699,721,747]" pageId="1" pageNumber="381">Paratypes</typeStatus>
:
<specimenCount id="9D4BFD08B823F716669D1DF166395AA7" box="[709,780,720,746]" pageId="1" pageNumber="381" type="male">1male</specimenCount>
,
<specimenCount id="9D4BFD08B823F716674F1DF166445AA7" box="[791,881,720,746]" pageId="1" pageNumber="381" type="female">1female</specimenCount>
(dissected) (
<collectionCode id="ED5CAE44B823F71667A71DF161695AA7" box="[1023,1116,720,746]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="1" pageNumber="381">ZMUM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B253CDCB823F71660361DF1648C5B43" collectingDate="2014-05-06" collectionCode="FSCB" collectorName="Laojunshan E &amp; I. Belousov &amp; Belousov, I." country="China" elevation="4075" latitude="26.882778" location="Laotzyunshan Mt." longLatPrecision="20" longitude="99.62889" pageId="1" pageNumber="381" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D4BFD08B823F71660361DF1618D5AA7" box="[1134,1208,720,746]" pageId="1" pageNumber="381" type="male">1 male</specimenCount>
(dissected) (
<collectionCode id="ED5CAE44B823F716611E1DF160A55AA7" box="[1350,1424,720,746]" pageId="1" pageNumber="381">FSCB</collectionCode>
), same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357650AB823F716649F1C3967DC5BAB" pageId="1" pageNumber="381" type="diagnosis">
<paragraph id="8BF23681B823F716649F1C3967DC5BAB" blockId="1.[151,1437,648,2043]" pageId="1" pageNumber="381">
<emphasis id="B939EA93B823F716649F1C3964775B7F" bold="true" box="[199,322,792,818]" pageId="1" pageNumber="381">Diagnosis.</emphasis>
Differs from congeners mainly by the relatively flattened apical part of the opisthomere mesomeral process with a longitudinal frontal, distinct broad excavation (
<emphasis id="B939EA93B823F71667171C1D665C5B1B" bold="true" box="[847,873,828,854]" pageId="1" pageNumber="381">ex</emphasis>
in
<figureCitation id="13762A04B823F71667D51C1D613E5B1B" box="[909,1035,828,854]" captionStart="FIGURES 1114" captionStartId="4.[152,270,1725,1749]" captionTargetBox="[154,1434,181,1693]" captionTargetId="figure-15@4.[154,1434,181,1701]" captionTargetPageId="4" captionText="FIGURES 1114. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 11. Opisthomere, mesal view. 12. Anterior part of opisthomere, mesal view. 13. Solenomere. 14. Opisthomere, anteromesal view. Abbreviations: bb, basal blade of solenomere; d, outgrowth of mesomeral process; ex, excavation; k, knobs. Scales: 10 μm (Figs 1213), 100 μm (Fig. 11), 0.5 mm (Fig. 14)." figureDoi="http://doi.org/10.5281/zenodo.8005006" httpUri="https://zenodo.org/record/8005006/files/figure.png" pageId="1" pageNumber="381">Figs 11, 14</figureCitation>
), by the relatively broad, somewhat flattened subapical outgrowth (
<emphasis id="B939EA93B823F71665AD1C4167305B37" bold="true" box="[501,517,864,890]" pageId="1" pageNumber="381">d</emphasis>
in
<figureCitation id="13762A04B823F71666721C4167E75B37" box="[554,722,864,890]" captionStart="FIGURES 1114" captionStartId="4.[152,270,1725,1749]" captionTargetBox="[154,1434,181,1693]" captionTargetId="figure-15@4.[154,1434,181,1701]" captionTargetPageId="4" captionText="FIGURES 1114. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 11. Opisthomere, mesal view. 12. Anterior part of opisthomere, mesal view. 13. Solenomere. 14. Opisthomere, anteromesal view. Abbreviations: bb, basal blade of solenomere; d, outgrowth of mesomeral process; ex, excavation; k, knobs. Scales: 10 μm (Figs 1213), 100 μm (Fig. 11), 0.5 mm (Fig. 14)." figureDoi="http://doi.org/10.5281/zenodo.8005006" httpUri="https://zenodo.org/record/8005006/files/figure.png" pageId="1" pageNumber="381">Figs 1112, 14</figureCitation>
) of the mesomeral process and the absence of the velar process coupled with the apically broadly rounded, slightly obtuse promere and penis as hourglass figure. Particularly similar to
<taxonomicName id="4C4D4D02B823F716655E1C8967E75B8F" authority="Mikhaljova" authorityName="Mikhaljova" authorityYear="2020" box="[262,722,936,962]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="381" phylum="Arthropoda" rank="species" species="lanpingensis">
<emphasis id="B939EA93B823F716655E1C8967795B8F" box="[262,588,936,962]" italics="true" pageId="1" pageNumber="381">Nepalmatoiulus lanpingensis</emphasis>
Mikhaljova
</taxonomicName>
and
<taxonomicName id="4C4D4D02B823F716675E1C8961765B8F" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[774,1091,936,962]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="1" pageNumber="381" phylum="Arthropoda" rank="species" species="simultaneus" status="sp. nov.">
<emphasis id="B939EA93B823F716675E1C8961765B8F" box="[774,1091,936,962]" italics="true" pageId="1" pageNumber="381">Nepalmatoiulus simultaneus</emphasis>
</taxonomicName>
<emphasis id="B939EA93B823F71660111C8961975B8F" bold="true" box="[1097,1186,936,962]" pageId="1" pageNumber="381">
<taxonomicNameLabel id="A20A57E8B823F71660111C8961975B8F" box="[1097,1186,936,962]" pageId="1" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but differs from these species by certain characters (see Discussion below).
</paragraph>
</subSubSection>
<subSubSection id="C357650AB823F712649F1CD1648059E0" lastPageId="5" lastPageNumber="385" pageId="1" pageNumber="381" type="description">
<paragraph id="8BF23681B823F716649F1CD1605B5CD7" blockId="1.[151,1437,648,2043]" pageId="1" pageNumber="381">
<emphasis id="B939EA93B823F716649F1CD1646C5C47" bold="true" box="[199,345,1008,1034]" pageId="1" pageNumber="381">Description.</emphasis>
<emphasis id="B939EA93B823F71665381CD164AF5C47" box="[352,410,1008,1034]" italics="true" pageId="1" pageNumber="381">Male</emphasis>
. Length in alcohol 31.034.0 mm, midbody vertical diameter 2.3 2.7 mm, with 57(1) (in a male
<typeStatus id="54F68823B823F71664BF1B3564785C63" box="[231,333,1044,1070]" pageId="1" pageNumber="381">holotype</typeStatus>
), 55(1), 52(1) rings, excluding telson. Coloration in alcohol dark grey; sometimes posterior body half with hardly lighter dorso-anterior metazonital portions. This portions not visible in dried specimen. Venter (including basal parts of legs) black-brown.Anterior several rings (78) of the usual julid type pattern. Legs dorsally and antennae dark brown. Occiput and forehead marbled brown, ventral parts of head light brown. Eyes black.
</paragraph>
<paragraph id="8BF23681B823F716649F1B8564BF5E97" blockId="1.[151,1437,648,2043]" pageId="1" pageNumber="381">
Head smooth and sparsely irregularly setose (in the form of blurred individual groups of setae), 2 epicranial setae, 4 supralabral setae; not less than 18 labral setae in one of the
<typeStatus id="54F68823B823F71667CA1BE861345CAE" box="[914,1025,1225,1251]" pageId="1" pageNumber="381">paratypes</typeStatus>
. Eye patches almost oval, composed of about 50 ommatidia (4748 ommatidia in one of the
<typeStatus id="54F68823B823F716677C1BCD66A05D4B" box="[804,917,1260,1286]" pageId="1" pageNumber="381">paratypes</typeStatus>
). Antennae medium-sized, rather slender and clavate. Gnathochilarium with no less than 14 nonapical stipital setae; lamellae linguales each with not less than 6 setae; mandibular stipites each with a subtriangular rounded smooth lobe; apical portion of the lobe without well demarcated border. Collum laterally with several distinct longer lower striae at posterior margin, not reaching to anterior margin, dorsally with short striae at posterior margin only. Length of collum striae gradually decreasing toward dorsum. A transverse row of sparse setae at hind edge of collum. Body rings circular. Prozona generally smooth, but with barely noticeable, very shallow obliquely located striae. Metazona with dense, regular, longitudinal striae reaching hind margin (18 or 19 striae in an approximate square with sides equal to metazonital length of a dorsal side of midbody ring). The metazonite gradually growing somewhat narrower towards body venter. Limbus straight, smooth (of Type
<quantity id="4CB59B64B823F716659B191164C45E07" box="[451,497,1584,1610]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="381" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFDC4B70B823F71665A2191167AE5E07" author="Enghoff, H." box="[506,667,1584,1610]" pageId="1" pageNumber="381" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
). A transverse row of thin setae at hind edge of metazonites, setae gradually growing longer and denser toward telson. Ozopores small, lying immediately behind suture dividing pro- and metazona. Caudal dorsal projection of telson straight and long, covered with setae and carrying at tip a claw-shaped process curved dorsally. Preanal ring setose; anal valves densely setose; subanal scale densely setose only in posterior half.
</paragraph>
<paragraph id="8BF23681B823F716649F19C467435FB7" blockId="1.[151,1437,648,2043]" pageId="1" pageNumber="381">
Legs relatively short and slender. Very delicately serrate ventral pads on postfemur and tibia, starting from legs 2 (
<figureCitation id="13762A04B823F71664EC182965CF5F6F" box="[180,250,1800,1826]" captionStart="FIGURES 15" captionStartId="2.[152,270,1732,1756]" captionTargetBox="[151,1436,192,1686]" captionTargetId="figure-15@2.[151,1436,192,1686]" captionTargetPageId="2" captionText="FIGURES 15. Nepalmatoiulus alternus sp. nov., male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes. 1. Leg pair 1, anterior view. 2. Leg 2, anterior view (mesapical oral seta broken off). 3. Coxae 2, anterior view. 4. Claw 2, anterior view. 5. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5)." figureDoi="http://doi.org/10.5281/zenodo.8005000" httpUri="https://zenodo.org/record/8005000/files/figure.png" pageId="1" pageNumber="381">Fig. 2</figureCitation>
). Claw of all legs at base with a long (somewhat longer than claw) setiform accessory claw ventrally (
<figureCitation id="13762A04B823F7166137182965925F0B" captionStart="FIGURES 15" captionStartId="2.[152,270,1732,1756]" captionTargetBox="[151,1436,192,1686]" captionTargetId="figure-15@2.[151,1436,192,1686]" captionTargetPageId="2" captionText="FIGURES 15. Nepalmatoiulus alternus sp. nov., male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes. 1. Leg pair 1, anterior view. 2. Leg 2, anterior view (mesapical oral seta broken off). 3. Coxae 2, anterior view. 4. Claw 2, anterior view. 5. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5)." figureDoi="http://doi.org/10.5281/zenodo.8005000" httpUri="https://zenodo.org/record/8005000/files/figure.png" pageId="1" pageNumber="381">Fig. 4</figureCitation>
, the setiform accessory claw is bent in this Figure). Leg pair 1 forming hook, the distal segment not in close contact with the basal segments (“Open hook”
<typeStatus id="54F68823B823F7166607187067BA5F26" box="[607,655,1873,1899]" pageId="1" pageNumber="381">type</typeStatus>
in
<bibRefCitation id="EFDC4B70B823F71666E01871666C5F27" author="Enghoff, H." box="[696,857,1872,1898]" pageId="1" pageNumber="381" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
). Leg pair 1: postfemur with slightly scaly-rugose ventral surface; tip ventrally slightly rugosity, coxa with one seta, distal segment without tarsal remnant (
<figureCitation id="13762A04B823F7166110185560BE5FC3" box="[1352,1419,1908,1934]" captionStart="FIGURES 15" captionStartId="2.[152,270,1732,1756]" captionTargetBox="[151,1436,192,1686]" captionTargetId="figure-15@2.[151,1436,192,1686]" captionTargetPageId="2" captionText="FIGURES 15. Nepalmatoiulus alternus sp. nov., male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes. 1. Leg pair 1, anterior view. 2. Leg 2, anterior view (mesapical oral seta broken off). 3. Coxae 2, anterior view. 4. Claw 2, anterior view. 5. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5)." figureDoi="http://doi.org/10.5281/zenodo.8005000" httpUri="https://zenodo.org/record/8005000/files/figure.png" pageId="1" pageNumber="381">Fig. 1</figureCitation>
). Coxa 2 with one mesapical oral seta, a gland opening positioned in central and axial position (
<figureCitation id="13762A04B823F71660E518B960355FFF" box="[1213,1280,1944,1970]" captionStart="FIGURES 15" captionStartId="2.[152,270,1732,1756]" captionTargetBox="[151,1436,192,1686]" captionTargetId="figure-15@2.[151,1436,192,1686]" captionTargetPageId="2" captionText="FIGURES 15. Nepalmatoiulus alternus sp. nov., male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes. 1. Leg pair 1, anterior view. 2. Leg 2, anterior view (mesapical oral seta broken off). 3. Coxae 2, anterior view. 4. Claw 2, anterior view. 5. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5)." figureDoi="http://doi.org/10.5281/zenodo.8005000" httpUri="https://zenodo.org/record/8005000/files/figure.png" pageId="1" pageNumber="381">Fig. 3</figureCitation>
) according to
<bibRefCitation id="EFDC4B70B823F71664CF189D64005F9B" author="Enghoff, H." box="[151,309,1980,2006]" pageId="1" pageNumber="381" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
. Penis as hourglass figure, about 1.8 longer than wide (
<figureCitation id="13762A04B823F71667FA189D66D25F9B" box="[930,999,1980,2006]" captionStart="FIGURES 610" captionStartId="3.[152,270,1823,1847]" captionTargetBox="[151,1436,199,1786]" captionTargetId="figure-33@3.[151,1436,199,1786]" captionTargetPageId="3" captionText="FIGURES 610. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 6. Penis, caudal view. 7. Left promere, caudal view. 8. Right promere, caudal view. 9. Distal part of left promere, caudal view. 10. Distal part of right promere, caudal view. Abbreviations: a, protrusion; la, lateral angle. Scales: 20 μm (Figs 6, 910), 100 μm (Figs 78)." figureDoi="http://doi.org/10.5281/zenodo.8005002" httpUri="https://zenodo.org/record/8005002/files/figure.png" pageId="1" pageNumber="381">Fig. 6</figureCitation>
). Ventral margin of body ring VII with lobes similar to figure
<quantity id="4CB59B64B823F71665CD18C164FB5FB7" box="[405,462,2016,2042]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="1" pageNumber="381" unit="in" value="32.0">32 in</quantity>
<bibRefCitation id="EFDC4B70B823F716658D18C167475FB7" author="Enghoff, H." box="[469,626,2016,2042]" pageId="1" pageNumber="381" pagination="241 - 331" refId="ref5882" refString="Enghoff, H. (1987) Revision of Nepalmatoiulus Mauries, 1983 - a Southeast Asiatic genus of millipedes (Diplopoda: Julida: Julidae). Courier Forschung-Institut Senckenberg, 93, 241 - 331." type="journal article" year="1987">Enghoff 1987</bibRefCitation>
.
</paragraph>
<caption id="DF326609B820F71564C019E566C95F69" ID-DOI="http://doi.org/10.5281/zenodo.8005000" ID-Zenodo-Dep="8005000" httpUri="https://zenodo.org/record/8005000/files/figure.png" pageId="2" pageNumber="382" startId="2.[152,270,1732,1756]" targetBox="[151,1436,192,1686]" targetPageId="2" targetType="figure">
<paragraph id="8BF23681B820F71564C019E566C95F69" blockId="2.[151,1436,1732,1828]" pageId="2" pageNumber="382">
<emphasis id="B939EA93B820F71564C019E564745E91" bold="true" box="[152,321,1732,1756]" pageId="2" pageNumber="382">FIGURES 15.</emphasis>
<taxonomicName id="4C4D4D02B820F715651019E567775E91" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[328,578,1732,1756]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="382" phylum="Arthropoda" rank="species" species="alternus" status="sp. nov.">
<emphasis id="B939EA93B820F715651019E567775E91" box="[328,578,1732,1756]" italics="true" pageId="2" pageNumber="382">Nepalmatoiulus alternus</emphasis>
</taxonomicName>
<emphasis id="B939EA93B820F715661019E567AC5E91" bold="true" box="[584,665,1732,1756]" pageId="2" pageNumber="382">
<taxonomicNameLabel id="A20A57E8B820F715661019E567AC5E91" box="[584,665,1732,1756]" pageId="2" pageNumber="382" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes.
<emphasis id="B939EA93B820F715617519E5600F5E91" bold="true" box="[1325,1338,1732,1756]" pageId="2" pageNumber="382">1</emphasis>
. Leg pair 1, anterior view.
<emphasis id="B939EA93B820F715651A19C9647A5F4D" bold="true" box="[322,335,1768,1792]" pageId="2" pageNumber="382">2</emphasis>
. Leg 2, anterior view (mesapical oral seta broken off).
<emphasis id="B939EA93B820F715672619C966BE5F4D" bold="true" box="[894,907,1768,1792]" pageId="2" pageNumber="382">3</emphasis>
. Coxae 2, anterior view.
<emphasis id="B939EA93B820F71560D119C961A35F4D" bold="true" box="[1161,1174,1768,1792]" pageId="2" pageNumber="382">4</emphasis>
. Claw 2, anterior view.
<emphasis id="B939EA93B820F71561D119C960A35F4D" bold="true" box="[1417,1430,1768,1792]" pageId="2" pageNumber="382">5</emphasis>
. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5).
</paragraph>
</caption>
<caption id="DF326609B821F71464C0183E61595F32" ID-DOI="http://doi.org/10.5281/zenodo.8005002" ID-Zenodo-Dep="8005002" httpUri="https://zenodo.org/record/8005002/files/figure.png" pageId="3" pageNumber="383" startId="3.[152,270,1823,1847]" targetBox="[151,1436,199,1786]" targetPageId="3" targetType="figure">
<paragraph id="8BF23681B821F71464C0183E61595F32" blockId="3.[151,1437,1823,1919]" pageId="3" pageNumber="383">
<emphasis id="B939EA93B821F71464C0183E64655F7A" bold="true" box="[152,336,1823,1847]" pageId="3" pageNumber="383">FIGURES 610.</emphasis>
<taxonomicName id="4C4D4D02B821F7146501183E67635F7A" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[345,598,1823,1847]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="383" phylum="Arthropoda" rank="species" species="alternus" status="sp. nov.">
<emphasis id="B939EA93B821F7146501183E67635F7A" box="[345,598,1823,1847]" italics="true" pageId="3" pageNumber="383">Nepalmatoiulus alternus</emphasis>
</taxonomicName>
<emphasis id="B939EA93B821F7146607183E67875F7A" bold="true" box="[607,690,1823,1847]" pageId="3" pageNumber="383">
<taxonomicNameLabel id="A20A57E8B821F7146607183E67875F7A" box="[607,690,1823,1847]" pageId="3" pageNumber="383" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (FSCB).
<emphasis id="B939EA93B821F71467E3183E66FD5F7A" bold="true" box="[955,968,1823,1847]" pageId="3" pageNumber="383">6</emphasis>
. Penis, caudal view.
<emphasis id="B939EA93B821F71460FF183E61815F7A" bold="true" box="[1191,1204,1823,1847]" pageId="3" pageNumber="383">7</emphasis>
. Left promere, caudal view.
<emphasis id="B939EA93B821F714648C186265D45F16" bold="true" box="[212,225,1859,1883]" pageId="3" pageNumber="383">8</emphasis>
. Right promere, caudal view.
<emphasis id="B939EA93B821F714664E186267165F16" bold="true" box="[534,547,1859,1883]" pageId="3" pageNumber="383">9</emphasis>
. Distal part of left promere, caudal view.
<emphasis id="B939EA93B821F7146797186266DD5F16" bold="true" box="[975,1000,1859,1883]" pageId="3" pageNumber="383">10</emphasis>
. Distal part of right promere, caudal view.
<emphasis id="B939EA93B821F71464CF184664795F32" bold="true" box="[151,332,1895,1919]" pageId="3" pageNumber="383">Abbreviations: a</emphasis>
, protrusion;
<emphasis id="B939EA93B821F7146596184664D75F32" bold="true" box="[462,482,1895,1919]" pageId="3" pageNumber="383">la</emphasis>
, lateral angle. Scales: 20 μm (Figs 6, 910), 100 μm (Figs 78).
</paragraph>
</caption>
<caption id="DF326609B826F71364C0199C60A85F50" ID-DOI="http://doi.org/10.5281/zenodo.8005006" ID-Zenodo-Dep="8005006" httpUri="https://zenodo.org/record/8005006/files/figure.png" pageId="4" pageNumber="384" startId="4.[152,270,1725,1749]" targetBox="[154,1434,181,1693]" targetPageId="4" targetType="figure">
<paragraph id="8BF23681B826F71364C0199C60A85F50" blockId="4.[151,1437,1725,1822]" pageId="4" pageNumber="384">
<emphasis id="B939EA93B826F71364C0199C646C5E98" bold="true" box="[152,345,1725,1749]" pageId="4" pageNumber="384">FIGURES 1114.</emphasis>
<taxonomicName id="4C4D4D02B826F7136506199C676D5E98" authority="Mikhaljova, 2023" authorityName="Mikhaljova" authorityYear="2023" box="[350,600,1725,1749]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="384" phylum="Arthropoda" rank="species" species="alternus" status="sp. nov.">
<emphasis id="B939EA93B826F7136506199C676D5E98" box="[350,600,1725,1749]" italics="true" pageId="4" pageNumber="384">Nepalmatoiulus alternus</emphasis>
</taxonomicName>
<emphasis id="B939EA93B826F7136605199C679B5E98" bold="true" box="[605,686,1725,1749]" pageId="4" pageNumber="384">
<taxonomicNameLabel id="A20A57E8B826F7136605199C679B5E98" box="[605,686,1725,1749]" pageId="4" pageNumber="384" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male paratype (FSCB).
<emphasis id="B939EA93B826F71367F2199C66F75E98" bold="true" box="[938,962,1725,1749]" pageId="4" pageNumber="384">11</emphasis>
. Opisthomere, mesal view.
<emphasis id="B939EA93B826F7136082199C61C65E98" bold="true" box="[1242,1267,1725,1749]" pageId="4" pageNumber="384">12</emphasis>
. Anterior part of opisthomere, mesal view.
<emphasis id="B939EA93B826F71365C719C0648D5EB4" bold="true" box="[415,440,1761,1785]" pageId="4" pageNumber="384">13</emphasis>
. Solenomere.
<emphasis id="B939EA93B826F713661219C067565EB4" bold="true" box="[586,611,1761,1785]" pageId="4" pageNumber="384">14</emphasis>
. Opisthomere, anteromesal view.
<emphasis id="B939EA93B826F71367E519C061B45EB4" bold="true" box="[957,1153,1761,1785]" pageId="4" pageNumber="384">Abbreviations: bb</emphasis>
, basal blade of solenomere;
<emphasis id="B939EA93B826F71364CF182465905F50" bold="true" box="[151,165,1797,1821]" pageId="4" pageNumber="384">d</emphasis>
, outgrowth of mesomeral process;
<emphasis id="B939EA93B826F7136652182467175F50" bold="true" box="[522,546,1797,1821]" pageId="4" pageNumber="384">ex</emphasis>
, excavation;
<emphasis id="B939EA93B826F71366F21824678D5F50" bold="true" box="[682,696,1797,1821]" pageId="4" pageNumber="384">k</emphasis>
, knobs. Scales: 10 μm (Figs 1213), 100 μm (Fig. 11), 0.5 mm (Fig. 14).
</paragraph>
</caption>
<paragraph id="8BF23681B826F712649F186A66505928" blockId="4.[151,1437,1867,2038]" lastBlockId="5.[151,1436,151,465]" lastPageId="5" lastPageNumber="385" pageId="4" pageNumber="384">
Gonopods slightly protruding. Anterior gonopod flattened, with subparallel margins, distally slightly expanded, apically broadly rounded, slightly obtuse, in posterior view subapically excavated for accommodation of mesomeral process, distal margins of the excavation papillate (especially mesal margin), rugose margin of apical excavation somewhat produced into low short blunt caudad protrusion (
<emphasis id="B939EA93B826F71367121896666D5F9C" bold="true" box="[842,856,1975,2001]" pageId="4" pageNumber="384">a</emphasis>
), subcentrally with knoblike flattened rudimentary telopodite (
<figureCitation id="13762A04B826F713654D18FA64BC5FB8" box="[277,393,2011,2037]" captionStart="FIGURES 610" captionStartId="3.[152,270,1823,1847]" captionTargetBox="[151,1436,199,1786]" captionTargetId="figure-33@3.[151,1436,199,1786]" captionTargetPageId="3" captionText="FIGURES 610. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 6. Penis, caudal view. 7. Left promere, caudal view. 8. Right promere, caudal view. 9. Distal part of left promere, caudal view. 10. Distal part of right promere, caudal view. Abbreviations: a, protrusion; la, lateral angle. Scales: 20 μm (Figs 6, 910), 100 μm (Figs 78)." figureDoi="http://doi.org/10.5281/zenodo.8005002" httpUri="https://zenodo.org/record/8005002/files/figure.png" pageId="4" pageNumber="384">Figs 710</figureCitation>
). In addition, apex of anterior gonopod variable even within same male
<typeStatus id="54F68823B826F71360FD18FD603F5FBB" box="[1189,1290,2012,2038]" pageId="4" pageNumber="384">paratype</typeStatus>
: lateral angle (
<emphasis id="B939EA93B827F71264F91FB6658258FC" bold="true" box="[161,183,151,177]" pageId="5" pageNumber="385">la</emphasis>
) of top of right promere (
<figureCitation id="13762A04B827F71265B91FB6670158FC" box="[481,564,151,177]" captionStart="FIGURES 610" captionStartId="3.[152,270,1823,1847]" captionTargetBox="[151,1436,199,1786]" captionTargetId="figure-33@3.[151,1436,199,1786]" captionTargetPageId="3" captionText="FIGURES 610. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 6. Penis, caudal view. 7. Left promere, caudal view. 8. Right promere, caudal view. 9. Distal part of left promere, caudal view. 10. Distal part of right promere, caudal view. Abbreviations: a, protrusion; la, lateral angle. Scales: 20 μm (Figs 6, 910), 100 μm (Figs 78)." figureDoi="http://doi.org/10.5281/zenodo.8005002" httpUri="https://zenodo.org/record/8005002/files/figure.png" pageId="5" pageNumber="385">Fig. 10</figureCitation>
) more elongated than that of left promere (
<figureCitation id="13762A04B827F712607E1FB6615958FC" box="[1062,1132,151,177]" captionStart="FIGURES 610" captionStartId="3.[152,270,1823,1847]" captionTargetBox="[151,1436,199,1786]" captionTargetId="figure-33@3.[151,1436,199,1786]" captionTargetPageId="3" captionText="FIGURES 610. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 6. Penis, caudal view. 7. Left promere, caudal view. 8. Right promere, caudal view. 9. Distal part of left promere, caudal view. 10. Distal part of right promere, caudal view. Abbreviations: a, protrusion; la, lateral angle. Scales: 20 μm (Figs 6, 910), 100 μm (Figs 78)." figureDoi="http://doi.org/10.5281/zenodo.8005002" httpUri="https://zenodo.org/record/8005002/files/figure.png" pageId="5" pageNumber="385">Fig. 9</figureCitation>
). Flagellum slender, short, caudally covered with cuticular spikes in distal part. Opisthomere hook-shaped mesomeral process apically with longitudinal frontal broad excavation (
<emphasis id="B939EA93B827F71266101FFE675758B4" bold="true" box="[584,610,223,249]" pageId="5" pageNumber="385">ex</emphasis>
); mesal margin of excavation with small knobs (
<emphasis id="B939EA93B827F71260D11FFE61AC58B4" bold="true" box="[1161,1177,223,249]" pageId="5" pageNumber="385">k</emphasis>
); subapical mesomeral outgrowth (
<emphasis id="B939EA93B827F71265451E2264185950" bold="true" box="[285,301,259,285]" pageId="5" pageNumber="385">d</emphasis>
) somewhat flattened, apically poorly papillate (
<figureCitation id="13762A04B827F712670B1E2261375950" box="[851,1026,259,285]" captionStart="FIGURES 1114" captionStartId="4.[152,270,1725,1749]" captionTargetBox="[154,1434,181,1693]" captionTargetId="figure-15@4.[154,1434,181,1701]" captionTargetPageId="4" captionText="FIGURES 1114. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 11. Opisthomere, mesal view. 12. Anterior part of opisthomere, mesal view. 13. Solenomere. 14. Opisthomere, anteromesal view. Abbreviations: bb, basal blade of solenomere; d, outgrowth of mesomeral process; ex, excavation; k, knobs. Scales: 10 μm (Figs 1213), 100 μm (Fig. 11), 0.5 mm (Fig. 14)." figureDoi="http://doi.org/10.5281/zenodo.8005006" httpUri="https://zenodo.org/record/8005006/files/figure.png" pageId="5" pageNumber="385">Figs 1112, 14</figureCitation>
). Membranous velum with strongly sloped smooth margin. Margin of accessory membrane serrate. Solenomere spinose, basally with blade (
<emphasis id="B939EA93B827F712616D1E066061590C" bold="true" box="[1333,1364,295,321]" pageId="5" pageNumber="385">bb</emphasis>
), then tapering into slender apical part spinose anteriorly (
<figureCitation id="13762A04B827F712668F1E6A666D5928" box="[727,856,331,357]" captionStart="FIGURES 1114" captionStartId="4.[152,270,1725,1749]" captionTargetBox="[154,1434,181,1693]" captionTargetId="figure-15@4.[154,1434,181,1701]" captionTargetPageId="4" captionText="FIGURES 1114. Nepalmatoiulus alternus sp. nov., male paratype (FSCB). 11. Opisthomere, mesal view. 12. Anterior part of opisthomere, mesal view. 13. Solenomere. 14. Opisthomere, anteromesal view. Abbreviations: bb, basal blade of solenomere; d, outgrowth of mesomeral process; ex, excavation; k, knobs. Scales: 10 μm (Figs 1213), 100 μm (Fig. 11), 0.5 mm (Fig. 14)." figureDoi="http://doi.org/10.5281/zenodo.8005006" httpUri="https://zenodo.org/record/8005006/files/figure.png" pageId="5" pageNumber="385">Figs 11, 13</figureCitation>
).
</paragraph>
<paragraph id="8BF23681B827F712649F1E4E648059E0" blockId="5.[151,1436,151,465]" pageId="5" pageNumber="385">
<emphasis id="B939EA93B827F712649F1E4E642E59C4" box="[199,283,367,393]" italics="true" pageId="5" pageNumber="385">Female</emphasis>
. Length in alcohol about 31.0 mm, midbody vertical diameter 2.4 mm, with 48(1) rings, excluding telson. Vulva as in
<figureCitation id="13762A04B827F71265341EB2648559E0" box="[364,432,402,429]" captionStart="FIGURES 15" captionStartId="2.[152,270,1732,1756]" captionTargetBox="[151,1436,192,1686]" captionTargetId="figure-15@2.[151,1436,192,1686]" captionTargetPageId="2" captionText="FIGURES 15. Nepalmatoiulus alternus sp. nov., male (Figs 14, FSCB) and female (Fig. 5, ZMUM) paratypes. 1. Leg pair 1, anterior view. 2. Leg 2, anterior view (mesapical oral seta broken off). 3. Coxae 2, anterior view. 4. Claw 2, anterior view. 5. Left vulva, caudal view. Scales: 10 μm (Fig. 4), 20 μm (Fig. 1), 100 μm (Figs 23, 5)." figureDoi="http://doi.org/10.5281/zenodo.8005000" httpUri="https://zenodo.org/record/8005000/files/figure.png" pageId="5" pageNumber="385">Fig. 5</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C357650AB827F712649F1E9661D9599C" box="[199,1260,439,465]" pageId="5" pageNumber="385" type="etymology">
<paragraph id="8BF23681B827F712649F1E9661D9599C" blockId="5.[151,1436,151,465]" box="[199,1260,439,465]" pageId="5" pageNumber="385">
<emphasis id="B939EA93B827F712649F1E96647B599C" bold="true" box="[199,334,439,465]" pageId="5" pageNumber="385">Etymology.</emphasis>
The specific epithet refers to the alternation of
<taxonomicName id="4C4D4D02B827F712673A1E966146599C" box="[866,1139,439,465]" class="Diplopoda" family="Julidae" genus="Nepalmatoiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="385" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B939EA93B827F712673A1E966126599C" box="[866,1043,439,465]" italics="true" pageId="5" pageNumber="385">Nepalmatoiulus</emphasis>
species.
</taxonomicName>
Adjective.
</paragraph>
</subSubSection>
</treatment>
</document>