155 lines
18 KiB
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155 lines
18 KiB
XML
<document id="3F79F9F2D1A0050840D3DC7EAF951205" ID-DOI="10.11646/zootaxa.5164.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6821026" ID-ZooBank="3BECB9C7-F4B5-4FA4-934B-1822BF3D1077" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657611643055" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2022" docId="03CE851E921CE943EBF949B3FA8FFE0D" docLanguage="en" docName="zootaxa.5164.1.1.pdf" docOrigin="Zootaxa 5164 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5164.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Milteliphaster Alcock 1893" docType="treatment" docVersion="3" lastPageNumber="57" masterDocId="FFF7FD66922BE97BEB6E4C15FFC4FFDC" masterDocTitle="New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition" masterLastPageNumber="75" masterPageNumber="1" pageNumber="56" updateTime="1699345508031" updateUser="plazi">
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<mods:title id="4F0F32C09790BA882B8F8BCF6AA42607">New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition</mods:title>
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<mods:namePart id="700773CCBEE8C3A9D353CE9C0BEF09EC">Mah, Christopher L.</mods:namePart>
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<mods:title id="C0F1997CDB81A24826094B451B92842E">Zootaxa</mods:title>
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<mods:part id="1842B9884AF4A89C123B7A9284B1C691">
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<mods:date id="14850AB524DA200158DE83E1110EDA18">2022</mods:date>
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<mods:identifier id="768FE62637DED66025DBC2FD1C703B48" type="DOI">10.11646/zootaxa.5164.1.1</mods:identifier>
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<subSubSection id="C37D6783921CE94CEBF949B3FD65F9DF" pageId="55" pageNumber="56" type="nomenclature">
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<paragraph id="8BD83408921CE94CEBF949B3FE25FA1C" blockId="55.[151,481,1446,1473]" box="[151,481,1446,1473]" pageId="55" pageNumber="56">
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<taxonomicName id="4C674F8B921CE94CEBF949B3FE25FA1C" ID-CoL="7PJPW" authority="Alcock 1893" authorityName="Alcock" authorityYear="1893" box="[151,481,1446,1473]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A921CE94CEBF949B3FE25FA1C" bold="true" box="[151,481,1446,1473]" pageId="55" pageNumber="56">
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<emphasis id="B913E81A921CE94CEBF949B3FE85FA1D" bold="true" box="[151,321,1446,1473]" italics="true" pageId="55" pageNumber="56">Milteliphaster</emphasis>
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<bibRefCitation id="EFF649F9921CE94CEA2649B3FE25FA1C" author="Alcock, A." box="[328,481,1446,1472]" pageId="55" pageNumber="56" pagination="73 - 121" refId="ref45221" refString="Alcock, A. (1893) Natural History notes from the HM Indian Marine Survey Steamer Investigator. 7. An account of the collection of deep-sea Asteroidea. Annals of the Magazine of Natural History, Series 6, 11 (6), 73 - 121. https: // doi. org / 10.1080 / 00222939308677478" type="journal article" year="1893">Alcock 1893</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BD83408921CE94CEBF949FEFD65F9DF" blockId="55.[151,673,1515,1539]" box="[151,673,1515,1539]" pageId="55" pageNumber="56">
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<bibRefCitation id="EFF649F9921CE94CEBF949FEFE87F9DF" author="Alcock, A." box="[151,323,1515,1539]" pageId="55" pageNumber="56" pagination="73 - 121" refId="ref45221" refString="Alcock, A. (1893) Natural History notes from the HM Indian Marine Survey Steamer Investigator. 7. An account of the collection of deep-sea Asteroidea. Annals of the Magazine of Natural History, Series 6, 11 (6), 73 - 121. https: // doi. org / 10.1080 / 00222939308677478" type="journal article" year="1893">Alcock 1893: 91</bibRefCitation>
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; Macan 1938: 391;
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<bibRefCitation id="EFF649F9921CE94CE96249FEFD59F9DF" author="Mah, C. L." box="[524,669,1515,1539]" pageId="55" pageNumber="56" pagination="1 - 116" refId="ref47769" refString="Mah, C. L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1 - 116. https: // doi. org / 10.11646 / zootaxa. 4539.1.1" type="journal article" year="2018">Mah 2018: 63</bibRefCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37D6783921CE94CEBF94A39FDA7F89E" pageId="55" pageNumber="56" type="diagnosis">
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<paragraph id="8BD83408921CE94CEBF94A39FB31F99A" blockId="55.[151,1437,1580,2039]" box="[151,1269,1580,1607]" pageId="55" pageNumber="56">
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<emphasis id="B913E81A921CE94CEBF94A39FED6F99A" bold="true" box="[151,274,1580,1606]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
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Body stellate (R/r=2.6–3.6), arms strap-like, triangular, with strongly upturned arm curl.
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</paragraph>
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<paragraph id="8BD83408921CE94CEBA94A45FDA7F89E" blockId="55.[151,1437,1580,2039]" pageId="55" pageNumber="56">
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interradial arcs straight to weakly curved. Abactinal surface covered by sharp, prominent spines, especially along carinal series. Abactinal plates without spines, polygonal to irregular in shape, relatively small with approximately 10–15 across arm base in
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<typeStatus id="54DC8AAA921CE94CE9624A8DFDAAF96E" box="[524,622,1688,1714]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
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and specimen described herein, surface smooth, flat to concave. Papular pores between plates, distinct. Marginal plates bearing 1 to many spines with raised base, surface otherwise flat or weakly concave, smooth forming lateral-facing series. Actinal plates with elongate, prominent pointed to blunt spines of varying abundance. Paddle-shaped pedicellariae present. Furrow spines 6–10, narrow with blunt tips, forming straight to weakly curved series.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37D6783921CE943EBA94B59FA8FFE0D" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="discussion">
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<paragraph id="8BD83408921CE94CEBA94B59FB19F872" blockId="55.[151,1437,1580,2039]" pageId="55" pageNumber="56">
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<emphasis id="B913E81A921CE94CEBA94B59FE8BF8BA" bold="true" box="[199,335,1868,1894]" pageId="55" pageNumber="56">Comments.</emphasis>
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<taxonomicName id="4C674F8B921CE94CEA3B4B59FE30F8BA" authorityName="Alcock" authorityYear="1893" box="[341,500,1868,1894]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A921CE94CEA3B4B59FE30F8BA" box="[341,500,1868,1894]" italics="true" pageId="55" pageNumber="56">Milteliphaster</emphasis>
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</taxonomicName>
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is part of the
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<taxonomicName id="4C674F8B921CE94CE9E44B59FD3EF8BA" authorityName="Gray" authorityYear="1840" box="[650,762,1868,1894]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A921CE94CE9E44B59FD3EF8BA" box="[650,762,1868,1894]" italics="true" pageId="55" pageNumber="56">Calliaster</emphasis>
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</taxonomicName>
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“group”, a cluster of similar
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<taxonomicName id="4C674F8B921CE94CEF524B59FB1EF8BA" authorityName="Forbes" authorityYear="1841" box="[1084,1242,1868,1894]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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, outlined by
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<bibRefCitation id="EFF649F9921CE94CEE094B59FEA5F857" author="Aziz, A. & Jangoux, M." pageId="55" pageNumber="56" pagination="585 - 586" refId="ref45387" refString="Aziz, A. & Jangoux, M. (1985) On the status and affinities of the goniasterid genus Calliaster Gray, 1840. In: Keegan, B. F. & O'Connor, B. D. S. (Eds.), Proceedings of the 5 th International Echinoderm Conference, Galway, 1984. Balkema, Rotterdam, pp 585 - 586." type="book chapter" year="1985">Aziz & Jangoux (1985)</bibRefCitation>
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and Mah (2005, 2018) based on, but not limited to, the presence of prominent spines on the abactinal, marginal and actinal surfaces, as well as the transverse series of large subambulacral spines.
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</paragraph>
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<paragraph id="8BD83408921CE943EBA94BADFE5BFEC1" blockId="55.[151,1437,1580,2039]" lastBlockId="56.[151,1436,151,465]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
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The name ‘
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<emphasis id="B913E81A921CE94CEA2B4BADFE2AF80E" box="[325,494,1976,2002]" italics="true" pageId="55" pageNumber="56">Milteliphaster’</emphasis>
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has been historically misapplied to other “
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<taxonomicName id="4C674F8B921CE94CE8A64BADFBFCF80E" authorityName="Gray" authorityYear="1840" box="[968,1080,1976,2002]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A921CE94CE8A64BADFBFCF80E" box="[968,1080,1976,2002]" italics="true" pageId="55" pageNumber="56">Calliaster</emphasis>
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</taxonomicName>
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”-like goniasterids with no clear basis.
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<emphasis id="B913E81A921CE94CEBB04BC9FE1BF82A" box="[222,479,2012,2038]" italics="true" pageId="55" pageNumber="56">
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<taxonomicName id="4C674F8B921CE94CEBB04BC9FE1EF82A" authorityName="Doderlein" authorityYear="1922" box="[222,474,2012,2038]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="species" species="regenerator">Calliaster regenerator</taxonomicName>
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,
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</emphasis>
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for example, was referred to
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<taxonomicName id="4C674F8B921CE94CE85E4BC9FC0BF82A" authorityName="Alcock" authorityYear="1893" box="[816,975,2012,2038]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="55" pageNumber="56" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A921CE94CE85E4BC9FC0BF82A" box="[816,975,2012,2038]" italics="true" pageId="55" pageNumber="56">Milteliphaster</emphasis>
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</taxonomicName>
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in
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<bibRefCitation id="EFF649F9921CE94CE89A4BC8FB34F82B" author="Rowe, F. W. E. & Gates, J." box="[1012,1264,2012,2039]" pageId="55" pageNumber="56" refId="ref48760" refString="Rowe, F. W. E. & Gates, J. (1995) Echinodermata in WELLS, A. (Ed.), Zoological Catalogue of Australia. Vol. 33. CSIRO, Melbourne, 510 pp." type="book" year="1995">Rowe & Gates (1995)</bibRefCitation>
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, a catalog with no figures or taxonomic argumentation showing the basis for the decision. Examination of the
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<typeStatus id="54DC8AAA9213E943EF8B4C82FAD1FF6D" box="[1253,1301,151,177]" pageId="56" pageNumber="57">type</typeStatus>
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material of the name-bearing species,
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<taxonomicName id="4C674F8B9213E943EAAA4CAEFD35FF09" box="[452,753,187,213]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="species" species="woodmason">
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<emphasis id="B913E81A9213E943EAAA4CAEFD35FF09" box="[452,753,187,213]" italics="true" pageId="56" pageNumber="57">Milteliphaster woodmason</emphasis>
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</taxonomicName>
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i
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<bibRefCitation id="EFF649F99213E943E9914CAEFC54FF09" author="Alcock, A." box="[767,912,187,213]" pageId="56" pageNumber="57" pagination="73 - 121" refId="ref45221" refString="Alcock, A. (1893) Natural History notes from the HM Indian Marine Survey Steamer Investigator. 7. An account of the collection of deep-sea Asteroidea. Annals of the Magazine of Natural History, Series 6, 11 (6), 73 - 121. https: // doi. org / 10.1080 / 00222939308677478" type="journal article" year="1893">Alcock 1893</bibRefCitation>
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led to restriction of the genus pending further understanding of
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<taxonomicName id="4C674F8B9213E943EA0F4CCAFE15FF25" authorityName="Gray" authorityYear="1840" box="[353,465,223,249]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A9213E943EA0F4CCAFE15FF25" box="[353,465,223,249]" italics="true" pageId="56" pageNumber="57">Calliaster</emphasis>
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</taxonomicName>
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and boundaries of related genera. A re-description of this species has recently been performed (
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<bibRefCitation id="EFF649F99213E943EA744D16FE57FEC1" author="Mah, C. L." box="[282,403,259,285]" pageId="56" pageNumber="57" pagination="1 - 116" refId="ref47769" refString="Mah, C. L. (2018) New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean. Zootaxa, 4539 (1), 1 - 116. https: // doi. org / 10.11646 / zootaxa. 4539.1.1" type="journal article" year="2018">Mah 2018</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BD834089213E943EBA94D32FA8FFE0D" blockId="56.[151,1436,151,465]" pageId="56" pageNumber="57">
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<taxonomicName id="4C674F8B9213E943EBA94D32FEA2FE9D" authorityName="Alcock" authorityYear="1893" box="[199,358,295,321]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A9213E943EBA94D32FEA2FE9D" box="[199,358,295,321]" italics="true" pageId="56" pageNumber="57">Milteliphaster</emphasis>
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</taxonomicName>
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is differentiated from
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<taxonomicName id="4C674F8B9213E943E90B4D32FD11FE9D" authorityName="Gray" authorityYear="1840" box="[613,725,295,321]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A9213E943E90B4D32FD11FE9D" box="[613,725,295,321]" italics="true" pageId="56" pageNumber="57">Calliaster</emphasis>
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</taxonomicName>
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on the basis of the shape and morphology of the abactinal and marginal plates. Superomarginal and inferomarginal plates in
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<taxonomicName id="4C674F8B9213E943E8244D5EFC2DFEB9" authorityName="Alcock" authorityYear="1893" box="[842,1001,331,357]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A9213E943E8244D5EFC2DFEB9" box="[842,1001,331,357]" italics="true" pageId="56" pageNumber="57">Milteliphaster</emphasis>
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||
</taxonomicName>
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are lateral-facing and displays a flat to weakly concave surface versus those in
|
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<taxonomicName id="4C674F8B9213E943E93C4D7AFD06FE55" authorityName="Gray" authorityYear="1840" box="[594,706,367,393]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A9213E943E93C4D7AFD06FE55" box="[594,706,367,393]" italics="true" pageId="56" pageNumber="57">Calliaster</emphasis>
|
||
</taxonomicName>
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which displays strongly convex surfaces. Abactinal plates show a parallel plate surface morphology with
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<taxonomicName id="4C674F8B9213E943E93F4D86FD34FE71" authorityName="Alcock" authorityYear="1893" box="[593,752,403,429]" class="Asteroidea" family="Goniasteridae" genus="Milteliphaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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||
<emphasis id="B913E81A9213E943E93F4D86FD34FE71" box="[593,752,403,429]" italics="true" pageId="56" pageNumber="57">Milteliphaster</emphasis>
|
||
</taxonomicName>
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displaying flat to weakly concave surfaces versus more convex surfaces in
|
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<taxonomicName id="4C674F8B9213E943EA284DA2FE72FE0D" authorityName="Gray" authorityYear="1840" box="[326,438,439,465]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
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<emphasis id="B913E81A9213E943EA284DA2FE72FE0D" box="[326,438,439,465]" italics="true" pageId="56" pageNumber="57">Calliaster</emphasis>
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</taxonomicName>
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. A manuscript detailing characters in
|
||
<taxonomicName id="4C674F8B9213E943E8304DA2FC0AFE0D" authorityName="Gray" authorityYear="1840" box="[862,974,439,465]" class="Asteroidea" family="Goniasteridae" genus="Calliaster" kingdom="Animalia" order="Valvatida" pageId="56" pageNumber="57" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B913E81A9213E943E8304DA2FC0AFE0D" box="[862,974,439,465]" italics="true" pageId="56" pageNumber="57">Calliaster</emphasis>
|
||
</taxonomicName>
|
||
and related taxa is in preparation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |