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<document id="748DEFA76FB17124B145ED50CAF8B7A3" ID-CLB-Dataset="21809" ID-DOI="10.5852/ejt.2020.652" ID-GBIF-Dataset="64fd24bb-ee1d-44d7-ac6a-25e8d5f96777" ID-ISSN="2118-9773" ID-Zenodo-Dep="3860236" ID-ZooBank="DC65C142-53F1-4416-A916-8F78C27DCF93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1590592466678" checkinUser="valdenar" docAuthor="Garbin, Rafaella C., Bandyopadhyay, Saswati &amp; Joyce, Walter G." docDate="2020" docId="03C7CF0BC02F1D54FDBAAA68FB2495A7" docLanguage="en" docName="ejt-652_garbin_bandyopadhyay_joyce.pdf.imf" docOrigin="European Journal of Taxonomy 652" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hardella thurjii" docType="treatment" docVersion="8" lastPageNumber="33" masterDocId="FFFEB773C0341D74FFD1A908FFA79736" masterDocTitle="A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan" masterLastPageNumber="67" masterPageNumber="1" pageNumber="28" updateTime="1698841626966" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B2D7C9A2BC82D3DD6A9FFBBD8970139B">A taxonomic revision of geoemydid turtles from Siwalik-age of India and Pakistan</mods:title>
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<mods:namePart id="5076AC83F7FE5A060B7C330B94E28C0F">Garbin, Rafaella C.</mods:namePart>
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<mods:affiliation id="8F3C5ACA5D98D77D31A811811A9DB40C">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. &amp; urn: lsid: zoobank. org: author: A 8 FCEF 2 B- 7524 - 46 C 3 - 808 C-D 9 CBFBC 08017 &amp; Corresponding author: garbin. rafaella @ gmail. com</mods:affiliation>
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<mods:namePart id="96D2E0589050D3A2128FC39F067979B8">Bandyopadhyay, Saswati</mods:namePart>
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<mods:affiliation id="EB2C33609CDF24912CE4D008460B3ED2">Geological Studies Unit, Indian Statistical Institute, Kolkata; Barrackpore Trunk Road 233, Kolkata, India. &amp; Email: saswati @ isical. ac. in &amp; urn: lsid: zoobank. org: author: 02005 D 35 - 6 DA 9 - 4 A 7 E-B 838 - 16 F 4 FEF 108 DA</mods:affiliation>
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<mods:namePart id="727ACBCC728A8F967567A297F2FCE1D2">Joyce, Walter G.</mods:namePart>
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<mods:affiliation id="C05A44EB86250326E0CAE2E3D5EB6B26">Department of Geosciences, University of Fribourg, Fribourg; Chemin du Musée 6, 1700 Fribourg, Switzerland. &amp; Email: walter. joyce @ unifr. ch &amp; urn: lsid: zoobank. org: author: C 30 D 317 A- 0029 - 439 C- 8446 - 31281771 ADA 5</mods:affiliation>
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<treatment id="03C7CF0BC02F1D54FDBAAA68FB2495A7" ID-DOI="http://doi.org/10.5281/zenodo.3861117" ID-GBIF-Taxon="164362571" ID-Zenodo-Dep="3861117" LSID="urn:lsid:plazi:treatment:03C7CF0BC02F1D54FDBAAA68FB2495A7" httpUri="http://treatment.plazi.org/id/03C7CF0BC02F1D54FDBAAA68FB2495A7" lastPageId="32" lastPageNumber="33" pageId="27" pageNumber="28">
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<heading id="D099C971C02F1D6FFDBAAA68FC6F944D" box="[619,968,864,891]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="28" reason="2">
<taxonomicName id="4C6E059EC02F1D6FFDBAAA68FC6F944D" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[619,968,864,891]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC02F1D6FFDBAAA68FC8C944D" bold="true" box="[619,811,864,891]" italics="true" pageId="27" pageNumber="28">Hardella thurjii</emphasis>
(Gray, 1831)
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<paragraph id="8BD17E1DC02F1D6FFD09AA80FCFB9495" blockId="27.[619,968,864,931]" box="[728,860,904,931]" pageId="27" pageNumber="28">
<figureCitation id="13556298C02F1D6FFD09AA80FCFB9495" box="[728,860,904,931]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="29.[189,232,1929,1955]" captionStartId-1="30.[189,232,1897,1923]" captionStartId-2="31.[189,232,1930,1956]" captionTargetBox-0="[199,1389,543,1875]" captionTargetBox-1="[194,1391,273,1848]" captionTargetBox-2="[201,1392,512,1880]" captionTargetId-0="figure@29.[189,1398,537,1879]" captionTargetId-1="figure@30.[189,1398,265,1854]" captionTargetId-2="figure@31.[189,1398,506,1881]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionTargetPageId-2="31" captionText-0="Fig. 19. BMNH 39835, holotype of Batagur falconeri Lydekker, 1885, here identified as a synonym of Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." captionText-1="Fig. 20. BMNH R.748, holotype of Clemmys watsoni Lydekker, 1886a, here identified as synonym of Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of carapace in lateral view. D. Illustration of carapace in lateral view. E. Photograph of plastron. F. Illustration of plastron. Scale bar: 3 cm." captionText-2="Fig. 21. BMNH R.890, referred to Hardella thurgi (Gray, 1831) by Lydekker (1889b), here identified as Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi-0="http://doi.org/10.5281/zenodo.3860278" figureDoi-1="http://doi.org/10.5281/zenodo.3860280" figureDoi-2="http://doi.org/10.5281/zenodo.3860282" httpUri-0="https://zenodo.org/record/3860278/files/figure.png" httpUri-1="https://zenodo.org/record/3860280/files/figure.png" httpUri-2="https://zenodo.org/record/3860282/files/figure.png" pageId="27" pageNumber="28">Figs 19–21</figureCitation>
</paragraph>
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<treatmentCitation id="0ACF580CC02F1D6FFF6CAAC7FDB794DF" author="Gray J. E." box="[189,528,975,1001]" page="22" pageId="27" pageNumber="28" year="1831">
<taxonomicName id="4C6E059EC02F1D6FFF6CAAC7FDB794DF" authority="Gray, 1831 a: 22" authorityName="Gray" authorityPageNumber="22" authorityYear="1831" box="[189,528,975,1001]" class="Reptilia" family="Emydidae" genus="Emys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC02F1D6FFF6CAAC7FEEA94DF" box="[189,333,975,1001]" italics="true" pageId="27" pageNumber="28">Emys thurjii</emphasis>
<bibRefCitation id="EFFF03ECC02F1D6FFE84AAC7FDB794DF" author="Gray J. E." box="[341,528,975,1001]" pageId="27" pageNumber="28" refId="ref23690" refString="Gray J. E. 1831 a. Synopsys Reptilium: or Short Descriptions of the Species of the Reptiles. Part I - Cataphracta. Tortoises, Crocodiles, and Enaliosaurus. Treuttel, Wurz and Co, London." type="book" year="1831">Gray, 1831a: 22</bibRefCitation>
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(
<emphasis id="B91AA20FC02F1D6FFDF3AAC7FD6294DE" box="[546,709,975,1000]" italics="true" pageId="27" pageNumber="28">nomen novum</emphasis>
).
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<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CAAFFFD779327" box="[189,720,1014,1041]" pageId="27" pageNumber="28">
<treatmentCitation id="0ACF580CC02F1D6FFF6CAAFFFE589326" box="[189,511,1014,1040]" pageId="27" pageNumber="28">
<taxonomicName id="4C6E059EC02F1D6FFF6CAAFFFE589326" authority="Gray, 1831 c: 8" authorityName="Gray" authorityPageNumber="8" authorityYear="1831" box="[189,511,1014,1040]" class="Reptilia" family="Emydidae" genus="Emys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thuryi">
<emphasis id="B91AA20FC02F1D6FFF6CAAFFFEED9326" box="[189,330,1014,1040]" italics="true" pageId="27" pageNumber="28">Emys thuryi</emphasis>
Gray, 1831c: 8
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(
<emphasis id="B91AA20FC02F1D6FFDC1AAFFFD189326" box="[528,703,1014,1040]" italics="true" pageId="27" pageNumber="28">nomen oblitum</emphasis>
).
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<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CAD17FD0B930E" box="[189,684,1054,1081]" pageId="27" pageNumber="28">
<treatmentCitation id="0ACF580CC02F1D6FFF6CAD17FD0B930E" box="[189,684,1054,1081]" pageId="27" pageNumber="28">
<taxonomicName id="4C6E059EC02F1D6FFF6CAD17FE75930E" authority="Gray, 1831 b" authorityName="Gray" authorityYear="1831" box="[189,466,1054,1080]" class="Reptilia" family="Emydidae" genus="Emys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thuji">
<emphasis id="B91AA20FC02F1D6FFF6CAD17FE9D930E" box="[189,314,1054,1080]" italics="true" pageId="27" pageNumber="28">Emys thuji</emphasis>
<bibRefCitation id="EFFF03ECC02F1D6FFE90AD16FE75930E" author="Gray J. E." box="[321,466,1054,1080]" pageId="27" pageNumber="28" refId="ref23731" refString="Gray J. E. 1831 b. Illustrations of Indian Zoology, Chiefly Selected from the Collection of Major-General Hardwicke. Vol I, part V. Treuttel, Wurtz, Treuttel jun. and Richter, London. https: // doi. org / 10.5962 / bhl. title. 95127" type="book" year="1831">Gray, 1831b</bibRefCitation>
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(ex errore): pl. 73
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.
</paragraph>
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<treatmentCitation id="0ACF580CC02F1D6FFF6CAD4FFDF89356" author="Lesson R. P." box="[189,607,1094,1120]" page="120" pageId="27" pageNumber="28" year="1831">
<taxonomicName id="4C6E059EC02F1D6FFF6CAD4FFDF89356" authority="Lesson, 1831: 120" authorityName="Lesson" authorityPageNumber="120" authorityYear="1831" box="[189,607,1094,1120]" class="Reptilia" family="Emydidae" genus="Emys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="flavonigra">
<emphasis id="B91AA20FC02F1D6FFF6CAD4FFEDA9356" box="[189,381,1094,1120]" italics="true" pageId="27" pageNumber="28">Emys flavonigra</emphasis>
<bibRefCitation id="EFFF03ECC02F1D6FFE55AD4EFDF89356" author="Lesson R. P." box="[388,607,1094,1120]" pageId="27" pageNumber="28" pagination="119 - 123" refId="ref24335" refString="Lesson R. P. 1831. Catalogue des reptiles qui font partie d`une collection zoologique recueillie dans l`Inde continentale ou en Afrique, et apportee en Franc par M. Lamare-Piquot. Bulletin des Sciences naturelles et Geologie de Paris 25: 119 - 123." type="journal article" year="1831">Lesson, 1831: 120</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BD17E1DC02F1D6FFF6CAD66FD7893BE" blockId="27.[189,960,975,1398]" box="[189,735,1134,1160]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CAD66FD7C93BE" box="[189,731,1134,1160]" pageId="27" pageNumber="28">
<treatmentCitation id="0ACF580CC02F1D6FFF6CAD66FE6D93BE" box="[189,458,1134,1160]" pageId="27" pageNumber="28">
<taxonomicName id="4C6E059EC02F1D6FFF6CAD66FE6D93BE" authority="Gray, 1832" authorityName="Gray" authorityYear="1832" box="[189,458,1134,1160]" class="Reptilia" family="Emydidae" genus="Emys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thugi">
<emphasis id="B91AA20FC02F1D6FFF6CAD66FEE793BE" box="[189,320,1134,1160]" italics="true" pageId="27" pageNumber="28">Emys thugi</emphasis>
<bibRefCitation id="EFFF03ECC02F1D6FFE96AD66FE6D93BE" author="Gray J. E." box="[327,458,1134,1160]" pageId="27" pageNumber="28" refId="ref23785" refString="Gray J. E. 1832. Illustrations of Indian Zoology, Chiefly Selected from the Collection of Major-General Hardwicke. Vol II, part XI. Adolphus Richter and Co., London. https: // doi. org / 10.5962 / bhl. title. 95127" type="book" year="1832">Gray, 1832</bibRefCitation>
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(ex errore): Directions
</treatmentCitationGroup>
.
</paragraph>
<paragraph id="8BD17E1DC02F1D6FFF6CAD9DFC679386" blockId="27.[189,960,975,1398]" box="[189,960,1173,1200]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CAD9DFC679386" box="[189,960,1173,1200]" pageId="27" pageNumber="28">
<treatmentCitation id="0ACF580CC02F1D6FFF6CAD9DFD5D9399" author="Fitzinger L. J." box="[189,762,1173,1200]" page="123" pageId="27" pageNumber="28" year="1835">
<taxonomicName id="4C6E059EC02F1D6FFF6CAD9DFD5D9399" authority="Fitzinger, 1835: 123" authorityName="Fitzinger" authorityPageNumber="123" authorityYear="1835" box="[189,762,1173,1200]" class="Reptilia" family="Emydidae" genus="Clemmys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurgii" subGenus="Clemmys">
<emphasis id="B91AA20FC02F1D6FFF6CAD9DFDA29399" box="[189,517,1173,1199]" italics="true" pageId="27" pageNumber="28">Clemmys (Clemmys) thurgii</emphasis>
<bibRefCitation id="EFFF03ECC02F1D6FFDDDAD9DFD5D9399" author="Fitzinger L. J." box="[524,762,1173,1200]" pageId="27" pageNumber="28" pagination="105 - 128" refId="ref23246" refString="Fitzinger L. J. 1835. Entwurf einer systematischen Anordnung der Schildkroten nach den Grundsatzen der naturlichen Methode. Annalen des Wiener Museums der Naturgeschichte 1: 105 - 128." type="journal article" year="1835">Fitzinger, 1835: 123</bibRefCitation>
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(
<emphasis id="B91AA20FC02F1D6FFCDAAD9EFC089399" box="[779,943,1174,1199]" italics="true" pageId="27" pageNumber="28">nomen novum</emphasis>
).
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</paragraph>
<paragraph id="8BD17E1DC02F1D6FFF6CADB5FDFC93E1" blockId="27.[189,960,975,1398]" box="[189,603,1213,1239]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CADB5FDFC93E1" box="[189,603,1213,1239]" pageId="27" pageNumber="28">
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<emphasis id="B91AA20FC02F1D6FFF6CADEEFE2993C9" box="[189,398,1253,1279]" italics="true" pageId="27" pageNumber="28">Batagur falconeri</emphasis>
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<emphasis id="B91AA20FC02F1D6FFF6CAC05FE2C9211" box="[189,395,1293,1319]" italics="true" pageId="27" pageNumber="28">Clemmys watsoni</emphasis>
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<emphasis id="B91AA20FC02F1D6FFF6CAC3CFEFC9278" box="[189,347,1332,1358]" italics="true" pageId="27" pageNumber="28">Hardella indi</emphasis>
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<emphasis id="B91AA20FC02F1D6FFF6CAC54FE069240" box="[189,417,1372,1398]" italics="true" pageId="27" pageNumber="28">Geoemyda pilgrimi</emphasis>
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<bibRefCitation id="EFFF03ECC02F1D6FFEA1ACAAFDBB928A" author="Gray J. E." box="[368,540,1442,1468]" pageId="27" pageNumber="28" refId="ref23834" refString="Gray J. E. 1844. Catalogue of the Tortoises, Crocodiles and Amphisbaenians, in the Collection of the British Museum. Order of the Trustees, London. https: // doi. org / 10.5962 / bhl. title. 21639" type="book" year="1844">Gray 1844: 17</bibRefCitation>
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<taxonomicName id="4C6E059EC02F1D6FFF6CACC3FEEC92D2" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[189,331,1482,1508]" class="Reptilia" family="Emydidae" genus="Emys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurgi">Emys thurgi</taxonomicName>
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<bibRefCitation id="EFFF03ECC02F1D6FFEB9ACC2FE4292D3" author="Gray J. E." box="[360,485,1482,1509]" pageId="27" pageNumber="28" refId="ref23879" refString="Gray J. E. 1856 (&quot; 1855 &quot;). Catalogue of Shield Reptiles in the Collection of the British Museum, Part I - Testudinata (Tortoises). Order of the Trustees, London. https: // doi. org / 10.5962 / bhl. title. 5491" type="book" year="1856">Gray 1856</bibRefCitation>
(“1855”): 21
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<taxonomicName id="4C6E059EC02F1D6FFF6CACFAFED8913A" authorityName="Fitzinger" authorityYear="1835" box="[189,383,1522,1548]" class="Reptilia" family="Emydidae" genus="Clemmys" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurgii">Clemmys thurgii</taxonomicName>
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<treatmentCitation id="0ACF580CC02F1D6FFE4DACFAFDCF913A" author="Strauch A." box="[412,616,1522,1548]" page="32" pageId="27" pageNumber="28" year="1862">
<bibRefCitation id="EFFF03ECC02F1D6FFE4DACFAFDCF913A" author="Strauch A." box="[412,616,1522,1548]" pageId="27" pageNumber="28" pagination="1 - 197" refId="ref25675" refString="Strauch A. 1862. Chelonologische Studien, mit besonderer Beziehung auf die Schildkrotensammlung der Kaiserlichen Akademie der Wissenschaften zu St. - Petersburg. Memoires de l'Academie imperiale des Sciences de St. - Petersbourg, 7 ieme serie 5 (7): 1 - 197." type="journal article" year="1862">Strauch 1862: 32</bibRefCitation>
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<taxonomicName id="4C6E059EC02F1D6FFF6CAF12FED49105" box="[189,371,1561,1587]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurgii">Batagur thurgii</taxonomicName>
–
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<treatmentCitation id="0ACF580CC02F1D6FFE41AF11FDD79102" author="Theobald W." box="[400,624,1561,1588]" page="12" pageId="27" pageNumber="28" year="1868">
<bibRefCitation id="EFFF03ECC02F1D6FFE41AF11FDD79102" author="Theobald W." box="[400,624,1561,1588]" pageId="27" pageNumber="28" refId="ref25847" refString="Theobald W. 1868. Catalogue of reptiles in the Museum of the Asiatic Society of Bengal. Journal of the Asiatic Society of Bengal, Extra Number. Baptist Mission Press, Kolkata. https: // doi. org / 10.5962 / bhl. title. 5477" type="book" year="1868">Theobald 1868: 12</bibRefCitation>
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<taxonomicName id="4C6E059EC02F1D6FFF6CAF49FED3916D" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[189,372,1601,1627]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurgi">Hardella thurgi</taxonomicName>
–
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<treatmentCitation id="0ACF580CC02F1D6FFE40AF49FD99916A" author="Gray J. E." box="[401,574,1601,1628]" page="58" pageId="27" pageNumber="28" year="1870">
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<taxonomicName id="4C6E059EC02F1D6FFF6CAF62FE4891B5" box="[189,495,1641,1667]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurgi" subGenus="Hardella">Batagur (Hardella) thurgi</taxonomicName>
–
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<treatmentCitation id="0ACF580CC02F1D6FFDDDAF61FCA791B5" author="Anderson J. A." box="[524,768,1641,1667]" page="764" pageId="27" pageNumber="28" year="1879">
<bibRefCitation id="EFFF03ECC02F1D6FFDDDAF61FCA791B5" author="Anderson J. A." box="[524,768,1641,1667]" pageId="27" pageNumber="28" refId="ref22354" refString="Anderson J. A. 1879. Anatomical and Zoological Researches: Comprising an Account of the Zoological Results of the two Expeditions to Western Yunnan in 1868 and 1875; and a Monograph of the two Cetacean Genera, Platanista and Orcella. Bernard Quaritch, London." type="book" year="1879">Anderson 1879: 764</bibRefCitation>
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<emphasis id="B91AA20FC02F1D6FFF6CAF99FED1919D" box="[189,374,1681,1707]" italics="true" pageId="27" pageNumber="28">Hardella thurjii</emphasis>
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–
<treatmentCitation id="0ACF580CC02F1D6FFE45AF99FD3E919A" author="Siebenrock F." box="[404,665,1681,1708]" page="456" pageId="27" pageNumber="28" year="1909">
<bibRefCitation id="EFFF03ECC02F1D6FFE45AF99FD3E919A" author="Siebenrock F." box="[404,665,1681,1708]" pageId="27" pageNumber="28" pagination="427 - 618" refId="ref25560" refString="Siebenrock F. 1909. Synopsis der rezenten Schildkroten, mit Berucksichtigung der in historischer Zeit ausgestorbenen Arten. Zoologische Jahrbucher Suppl. 10 (3): 427 - 618. https: // doi. org / 10.5962 / bhl. title. 12485" type="journal article" year="1909">Siebenrock 1909: 456</bibRefCitation>
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<emphasis id="B91AA20FC02F1D6FFF6CAFB1FED391E5" box="[189,372,1721,1747]" italics="true" pageId="27" pageNumber="28">Hardella thurgi</emphasis>
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–
<treatmentCitation id="0ACF580CC02F1D6FFE40AFB0FDED91E5" author="Smith M. A." box="[401,586,1720,1747]" page="50" pageId="27" pageNumber="28" year="1931">
<bibRefCitation id="EFFF03ECC02F1D6FFE40AFB0FDED91E5" author="Smith M. A." box="[401,586,1720,1747]" pageId="27" pageNumber="28" refId="ref25606" refString="Smith M. A. 1931. The Fauna of British India, Including Ceylon and Burma, Reptilia and Amphibia. Vol. I - Loricata, Testudines. Taylor and Francis, London." type="book" year="1931">Smith 1931: 50</bibRefCitation>
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<taxonomicName id="4C6E059EC02F1D6FFF6CAFE8FE6091CC" authority="thurjii" box="[189,455,1760,1786]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="subSpecies" species="thurjii" subSpecies="thurjii">Hardella thurjii thurjii</taxonomicName>
–
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<bibRefCitation id="EFFF03ECC02F1D6FFE34AFE8FCEE91CC" author="Wermuth H. &amp; Mertens R." box="[485,841,1760,1787]" pageId="27" pageNumber="28" refId="ref26199" refString="Wermuth H. &amp; Mertens R. 1977. Testudines, Crocodylia, Rhynchocephalia. Das Tierreich. Walter de Gruyter, Berlin." type="book" year="1977">Wermuth &amp; Mertens 1977: 40</bibRefCitation>
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<taxonomicName id="4C6E059EC02F1D6FFF6CAE00FE0B9014" baseAuthorityName="Gray" baseAuthorityYear="1870" box="[189,428,1800,1826]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="subSpecies" species="thurjii" subSpecies="indi">Hardella thurjii indi</taxonomicName>
–
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<bibRefCitation id="EFFF03ECC02F1D6FFE18AE00FC8A9014" author="Wermuth H. &amp; Mertens R." box="[457,813,1800,1827]" pageId="27" pageNumber="28" refId="ref26199" refString="Wermuth H. &amp; Mertens R. 1977. Testudines, Crocodylia, Rhynchocephalia. Das Tierreich. Walter de Gruyter, Berlin." type="book" year="1977">Wermuth &amp; Mertens 1977: 40</bibRefCitation>
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</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD17E1DC02F1D6FFF6CAE38FCA7907C" blockId="27.[189,845,1442,1906]" box="[189,768,1840,1866]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CAE38FD5B907C" box="[189,764,1840,1866]" pageId="27" pageNumber="28">
<taxonomicName id="4C6E059EC02F1D6FFF6CAE38FEC9907C" box="[189,366,1840,1866]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="thurji">
<emphasis id="B91AA20FC02F1D6FFF6CAE38FEC9907C" box="[189,366,1840,1866]" italics="true" pageId="27" pageNumber="28">Hardella thurji</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACF580CC02F1D6FFE5DAE38FD5B907C" box="[396,764,1840,1866]" pageId="27" pageNumber="28">
<bibRefCitation id="EFFF03ECC02F1D6FFE5DAE38FD9D907C" author="Pritchard P. C. H." box="[396,570,1840,1866]" pageId="27" pageNumber="28" refId="ref25536" refString="Pritchard P. C. H. 1979. Encyclopedia of Turtles. T. F. H. Publications, New Jersey." type="book" year="1979">Pritchard 1979</bibRefCitation>
(ex errore): 193
</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C02F1D6FFF6CAE50FD369044" box="[189,657,1879,1906]" pageId="27" pageNumber="28" type="description">
<paragraph id="8BD17E1DC02F1D6FFF6CAE50FD369044" blockId="27.[189,845,1442,1906]" box="[189,657,1879,1906]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB7E5933C02F1D6FFF6CAE50FD2B9044" box="[189,652,1879,1906]" pageId="27" pageNumber="28">
<taxonomicName id="4C6E059EC02F1D6FFF6CAE50FE109044" authority="thurji" box="[189,439,1880,1906]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="27" pageNumber="28" phylum="Chordata" rank="subSpecies" species="thurji" subSpecies="thurji">
<emphasis id="B91AA20FC02F1D6FFF6CAE50FE109044" box="[189,439,1880,1906]" italics="true" pageId="27" pageNumber="28">Hardella thurji thurji</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ACF580CC02F1D6FFE05AE5FFD2B9044" box="[468,652,1879,1906]" pageId="27" pageNumber="28">Obst 1985: 221</treatmentCitation>
</treatmentCitationGroup>
.
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C02F1D68FF6CAE96FDA0954A" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BD17E1DC02F1D6FFF6CAE96FF5D908E" blockId="27.[189,250,1950,1976]" box="[189,250,1950,1976]" pageId="27" pageNumber="28">
<heading id="D099C971C02F1D6FFF6CAE96FF5D908E" bold="true" box="[189,250,1950,1976]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B91AA20FC02F1D6FFF6CAE96FF5D908E" bold="true" box="[189,250,1950,1976]" pageId="27" pageNumber="28">
<typeStatus id="54D5C0BFC02F1D6FFF6CAE96FF5D908E" box="[189,250,1950,1976]" pageId="27" pageNumber="28">Type</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC02F1D6FFF6CAED9FE4A90DD" blockId="27.[189,493,2001,2027]" box="[189,493,2001,2027]" pageId="27" pageNumber="28">
Unknown (
<bibRefCitation id="EFFF03ECC02F1D6FFE91AED9FE4790DD" author="Iverson J. B." box="[320,480,2001,2027]" pageId="27" pageNumber="28" refId="ref24146" refString="Iverson J. B. 1992. A Revised Checklist with Distribution Maps of the Turtles of the World. Earlham College, privately printed, Richmond." type="book" year="1992">Iverson 1992</bibRefCitation>
).
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CA803FE0D9613" blockId="28.[189,426,267,293]" box="[189,426,267,293]" pageId="28" pageNumber="29">
<heading id="D099C971C0281D68FF6CA803FE0D9613" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B91AA20FC0281D68FF6CA803FE0D9613" bold="true" box="[189,426,267,293]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CA832FB7696AD" blockId="28.[189,1399,313,411]" pageId="28" pageNumber="29">
<materialsCitation id="3B067440C0281D68FF6CA832FAD19662" ID-GBIF-Occurrence="2626092315" box="[189,1398,313,340]" collectionCode="BMNH" country="India" location="Siwalik Hills" municipality="Siwalik Hills" pageId="28" pageNumber="29" specimenCode="BMNH 39835" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F3793E8DC0281D68FF6CA832FEB79662" box="[189,272,314,340]" name="India" pageId="28" pageNumber="29">INDIA</collectingCountry>
•
<specimenCount id="9D68B594C0281D68FEFBA832FE139662" box="[298,436,314,340]" pageId="28" pageNumber="29" type="generic">1 specimen</specimenCount>
, holotype of
<taxonomicName id="4C6E059EC0281D68FD80A833FC839662" authorityName="Lydekker" authorityYear="1885" box="[593,804,314,340]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="falconeri">
<emphasis id="B91AA20FC0281D68FD80A833FC839662" box="[593,804,314,340]" italics="true" pageId="28" pageNumber="29">Batagur falconeri</emphasis>
</taxonomicName>
;
<location id="8EB128C6C0281D68FCE4A832FC739662" LSID="urn:lsid:plazi:treatment:03C7CF0BC02F1D54FDBAAA68FB2495A7:8EB128C6C0281D68FCE4A832FC739662" box="[821,980,314,340]" country="India" municipality="Siwalik Hills" name="Siwalik Hills" pageId="28" pageNumber="29">Siwalik Hills</location>
; Miocene–Pliocene;
<specimenCode id="DBC8D666C0281D68FB1DA832FAD19662" box="[1228,1398,313,340]" collectionCode="BMNH" pageId="28" pageNumber="29">BMNH 39835</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B067440C0281D68FF02A855FC1F9641" ID-GBIF-Occurrence="2626092317" box="[211,952,349,376]" collectionCode="BMNH" country="India" location="Siwalik Hills" municipality="Siwalik Hills" pageId="28" pageNumber="29" specimenCode="BMNH R.890" specimenCount="1" typeStatus="holotype">
<specimenCount id="9D68B594C0281D68FF02A855FEC79641" box="[211,352,349,375]" pageId="28" pageNumber="29" type="generic">1 specimen</specimenCount>
;
<location id="8EB128C6C0281D68FEBEA855FDB69641" LSID="urn:lsid:plazi:treatment:03C7CF0BC02F1D54FDBAAA68FB2495A7:8EB128C6C0281D68FEBEA855FDB69641" box="[367,529,349,375]" country="India" municipality="Siwalik Hills" name="Siwalik Hills" pageId="28" pageNumber="29">Siwalik Hills</location>
; Miocene–Pliocene;
<specimenCode id="DBC8D666C0281D68FCDFA856FC1F9641" box="[782,952,349,376]" pageId="28" pageNumber="29">BMNH R.890</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B067440C0281D68FC0BA855FB6996AD" ID-GBIF-Occurrence="2626092322" collectionCode="BMNH" country="India" county="Gulf of Cambay" location="Gulf of Cambay" municipality="Piram Island" pageId="28" pageNumber="29" specimenCode="BMNH R.748" specimenCount="1" stateProvince="Gujarat" typeStatus="holotype">
<specimenCount id="9D68B594C0281D68FC0BA855FBC19641" box="[986,1126,349,375]" pageId="28" pageNumber="29" type="generic">1 specimen</specimenCount>
, holotype of
<taxonomicName id="4C6E059EC0281D68FADAA855FEBF96AD" authorityName="Lydekker" authorityYear="1886" class="Reptilia" family="Emydidae" genus="Clemmys" kingdom="Animalia" order="Testudines" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="watsoni">
<emphasis id="B91AA20FC0281D68FADAA855FEBF96AD" italics="true" pageId="28" pageNumber="29">Clemmys watsoni</emphasis>
</taxonomicName>
;
<location id="8EB128C6C0281D68FEF9A889FE4F96AD" LSID="urn:lsid:plazi:treatment:03C7CF0BC02F1D54FDBAAA68FB2495A7:8EB128C6C0281D68FEF9A889FE4F96AD" box="[296,488,385,411]" country="India" county="Gulf of Cambay" municipality="Piram Island" name="Gulf of Cambay" pageId="28" pageNumber="29" stateProvince="Gujarat">Gulf of Cambay</location>
,
<collectingRegion id="49AAB0FFC0281D68FE22A889FDE996AD" box="[499,590,385,411]" country="India" name="Gujarat" pageId="28" pageNumber="29">Gujarat</collectingRegion>
,
<location id="8EB128C6C0281D68FD8BA889FD5596AD" LSID="urn:lsid:plazi:treatment:03C7CF0BC02F1D54FDBAAA68FB2495A7:8EB128C6C0281D68FD8BA889FD5596AD" box="[602,754,385,411]" country="India" county="Gulf of Cambay" municipality="Piram Island" name="Piram Island" pageId="28" pageNumber="29" stateProvince="Gujarat">Piram Island</location>
; Late Miocene–Pliocene;
<specimenCode id="DBC8D666C0281D68FBF5A889FB6996AD" box="[1060,1230,385,411]" pageId="28" pageNumber="29">BMNH R.748</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CA8CBFEFA96EA" blockId="28.[189,349,450,477]" box="[189,349,450,477]" pageId="28" pageNumber="29">
<heading id="D099C971C0281D68FF6CA8CBFEFA96EA" bold="true" box="[189,349,450,477]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B91AA20FC0281D68FF6CA8CBFEFA96EA" bold="true" box="[189,349,450,477]" pageId="28" pageNumber="29">
<typeStatus id="54D5C0BFC0281D68FF6CA8CBFF5D96EB" box="[189,250,451,477]" pageId="28" pageNumber="29">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CA8F9FE77953D" blockId="28.[189,464,497,523]" box="[189,464,497,523]" pageId="28" pageNumber="29">
“
<collectingCountry id="F3793E8DC0281D68FF16A8F9FEAF953D" box="[199,264,497,523]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
” (
<bibRefCitation id="EFFF03ECC0281D68FEF2A8F9FE64953D" author="Iverson J. B." box="[291,451,497,523]" pageId="28" pageNumber="29" refId="ref24146" refString="Iverson J. B. 1992. A Revised Checklist with Distribution Maps of the Turtles of the World. Earlham College, privately printed, Richmond." type="book" year="1992">Iverson 1992</bibRefCitation>
).
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CAB3BFEE8957B" blockId="28.[189,335,563,589]" box="[189,335,563,589]" pageId="28" pageNumber="29">
<heading id="D099C971C0281D68FF6CAB3BFEE8957B" bold="true" box="[189,335,563,589]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B91AA20FC0281D68FF6CAB3BFEE8957B" bold="true" box="[189,335,563,589]" pageId="28" pageNumber="29">Occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CAB6AFDA0954A" blockId="28.[189,519,610,636]" box="[189,519,610,636]" pageId="28" pageNumber="29">Miocene/Pliocene – Recent.</paragraph>
</subSubSection>
<subSubSection id="C3742D96C0281D68FF6CABABFCB89461" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BD17E1DC0281D68FF6CABABFCD09588" blockId="28.[189,887,675,702]" box="[189,887,675,702]" pageId="28" pageNumber="29">
<heading id="D099C971C0281D68FF6CABABFCD09588" bold="true" box="[189,887,675,702]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B91AA20FC0281D68FF6CABABFCD09588" bold="true" box="[189,887,675,702]" pageId="28" pageNumber="29">Differential osteological diagnosis using shell characters</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CABDAFCB89461" blockId="28.[189,1399,722,855]" pageId="28" pageNumber="29">
<taxonomicName id="4C6E059EC0281D68FF6CABDAFEDE95DA" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[189,377,722,748]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0281D68FF6CABDAFEDE95DA" box="[189,377,722,748]" italics="true" pageId="28" pageNumber="29">Hardella thurjii</emphasis>
</taxonomicName>
can be differentiated from other geoemydids by large carapace size (up to
<quantity id="4C96D3F8C0281D68FADBABDAFAF195DB" box="[1290,1366,722,749]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="29" unit="cm" value="60.0">60 cm</quantity>
in females), presence of hexagonal, nearly square second to fourth vertebrals, a first vertebral scute that is wider posteriorly, presence of an inflection at the margin of the gulohumeral sulcus, and an entoplastron that is not intersected by the humeropectoral sulcus.
</paragraph>
</subSubSection>
<subSubSection id="C3742D96C0281D54FF6CAA76FAF8967C" lastPageId="32" lastPageNumber="33" pageId="28" pageNumber="29" type="description">
<paragraph id="8BD17E1DC0281D68FF6CAA76FDF894AE" blockId="28.[189,607,894,920]" box="[189,607,894,920]" pageId="28" pageNumber="29">
<heading id="D099C971C0281D68FF6CAA76FDF894AE" bold="true" box="[189,607,894,920]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B91AA20FC0281D68FF6CAA76FDF894AE" bold="true" box="[189,607,894,920]" pageId="28" pageNumber="29">Description of material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0281D68FF6CAAA6FDE292A3" blockId="28.[189,1399,941,1429]" pageId="28" pageNumber="29">
BMNH 39835 (
<figureCitation id="13556298C0281D68FEA5AAA5FE6D94F1" box="[372,458,941,967]" captionStart="Fig" captionStartId="29.[189,232,1929,1955]" captionTargetBox="[199,1389,543,1875]" captionTargetId="figure@29.[189,1398,537,1879]" captionTargetPageId="29" captionText="Fig. 19. BMNH 39835, holotype of Batagur falconeri Lydekker, 1885, here identified as a synonym of Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860278" httpUri="https://zenodo.org/record/3860278/files/figure.png" pageId="28" pageNumber="29">Fig. 19</figureCitation>
),
<typeStatus id="54D5C0BFC0281D68FE0EAAA5FDE194F1" box="[479,582,941,967]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6E059EC0281D68FDBAAAA6FC9C94F1" authorityName="Lydekker" authorityYear="1885" box="[619,827,941,967]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="falconeri">
<emphasis id="B91AA20FC0281D68FDBAAAA6FC9C94F1" box="[619,827,941,967]" italics="true" pageId="28" pageNumber="29">Batagur falconeri</emphasis>
</taxonomicName>
– This is an almost complete shell, exceptionally well preserved, from the Miocene/Pliocene of the Siwalik Hills, likely of
<collectingCountry id="F3793E8DC0281D68FBFBAAD9FBCE94DD" box="[1066,1129,977,1003]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
, originally figured and described in three views by
<bibRefCitation id="EFFF03ECC0281D68FDC5AAFCFD449339" author="Lydekker R." box="[532,739,1012,1039]" pageId="28" pageNumber="29" pagination="155 - 208" refId="ref24414" refString="Lydekker R. 1885 a. Indian Tertiary and post-Tertiary Vertebrata, Part III - Siwalik and Narbada Chelonia. Memoirs of the Geological Survey of India, Palaeontologia Indica 10 (3): 155 - 208." type="journal article" year="1885">Lydekker (1885a</bibRefCitation>
: pls 23.1, 24.4). Our figures are overall comparable to those of Lydekker, although we see more details to the peripherals (
<figureCitation id="13556298C0281D68FBD1AD10FBFC9304" box="[1024,1115,1048,1074]" captionStart="Fig" captionStartId="29.[189,232,1929,1955]" captionTargetBox="[199,1389,543,1875]" captionTargetId="figure@29.[189,1398,537,1879]" captionTargetPageId="29" captionText="Fig. 19. BMNH 39835, holotype of Batagur falconeri Lydekker, 1885, here identified as a synonym of Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860278" httpUri="https://zenodo.org/record/3860278/files/figure.png" pageId="28" pageNumber="29">Fig. 19</figureCitation>
). The specimen clearly represents an adult female specimen based on overall size (carapace length greater than
<quantity id="4C96D3F8C0281D68FB2AAD33FAEB9360" box="[1275,1356,1083,1110]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="28" pageNumber="29" unit="cm" value="40.0">40 cm</quantity>
). A small keel is present on neurals IV and VI. There are no signs of growth annuli. All neural bones have short anterior sides. The third neural is half one and a half time longer than the other neural bones. The cervical scute is present, longer than wide. The first vertebral scute is broader posteriorly and narrows anteriorly. The second and third vertebrals are much broader than the other vertebral scutes. The fifth vertebral has an anterolateral constriction. Marginal scutes IV–X overlap the adjacent costal bones. The anterior and posterior plastral margins are not preserved. The pectoroabdominal sulcus has lateral notches, suggesting the former presence of longitudinal keels as a juvenile. The pectoroabdominal sulcus does not intersect the hyo-hypoplastron suture and contacts the fifth marginal scute. Axillary and inguinal scutes are likely present.
</paragraph>
<paragraph id="8BD17E1DC0281D69FF6CACB5FAEA967F" blockId="28.[189,1399,1468,2027]" lastBlockId="29.[189,1398,267,329]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
BMNH R.748 (
<figureCitation id="13556298C0281D68FEAEACB4FE7A92E0" box="[383,477,1468,1494]" captionStart="Fig" captionStartId="30.[189,232,1897,1923]" captionTargetBox="[194,1391,273,1848]" captionTargetId="figure@30.[189,1398,265,1854]" captionTargetPageId="30" captionText="Fig. 20. BMNH R.748, holotype of Clemmys watsoni Lydekker, 1886a, here identified as synonym of Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of carapace in lateral view. D. Illustration of carapace in lateral view. E. Photograph of plastron. F. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860280" httpUri="https://zenodo.org/record/3860280/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
),
<typeStatus id="54D5C0BFC0281D68FE26ACB4FDF992E0" box="[503,606,1468,1494]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6E059EC0281D68FD40ACB4FCC392E0" authorityName="Lydekker" authorityYear="1886" box="[657,868,1468,1494]" class="Reptilia" family="Emydidae" genus="Clemmys" kingdom="Animalia" order="Testudines" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="watsoni">
<emphasis id="B91AA20FC0281D68FD40ACB4FCC392E0" box="[657,868,1468,1494]" italics="true" pageId="28" pageNumber="29">Clemmys watsoni</emphasis>
</taxonomicName>
– This specimen originates from the Late Miocene–Pliocene of Piram Island, Gulf of
<collectingCountry id="F3793E8DC0281D68FD00ACE8FC9392CC" box="[721,820,1504,1530]" name="India" pageId="28" pageNumber="29">Cambay</collectingCountry>
,
<collectingRegion id="49AAB0FFC0281D68FC92ACE8FC3992CC" box="[835,926,1504,1530]" country="India" name="Gujarat" pageId="28" pageNumber="29">Gujarat</collectingRegion>
,
<collectingCountry id="F3793E8DC0281D68FC7CACE8FC4B92CC" box="[941,1004,1504,1530]" name="India" pageId="28" pageNumber="29">India</collectingCountry>
, was presented to the BMNH in 1886 by Col. J.W. Watson and figured and described in a small contribution from
<bibRefCitation id="EFFF03ECC0281D68FB75AF0BFAD4912B" author="Lydekker R." box="[1188,1395,1539,1565]" pageId="28" pageNumber="29" pagination="540 - 541" refId="ref24567" refString="Lydekker R. 1886 a. On a new Emydine chelonian from the Pliocene of India. Quarterly Journal of the Geological Society 42: 540 - 541. https: // doi. org / 10.1144 / gsl. jgs. 1886.042.01 - 04.54" type="journal article" year="1886">Lydekker (1886a</bibRefCitation>
: pl. 1). Our illustrations in three views overall confirm most of Lydekker’s observations, but we see fewer details along the neural column and the damaged portions of the costals and peripherals. The specimen is almost complete, full size, and misses some lateral peripheral bones (right and left), the anterior plastral margin, the posterior plastral lobe, and the right bridge (
<figureCitation id="13556298C0281D68FC72AF99FC5D919D" box="[931,1018,1681,1707]" captionStart="Fig" captionStartId="30.[189,232,1897,1923]" captionTargetBox="[194,1391,273,1848]" captionTargetId="figure@30.[189,1398,265,1854]" captionTargetPageId="30" captionText="Fig. 20. BMNH R.748, holotype of Clemmys watsoni Lydekker, 1886a, here identified as synonym of Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of carapace in lateral view. D. Illustration of carapace in lateral view. E. Photograph of plastron. F. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860280" httpUri="https://zenodo.org/record/3860280/files/figure.png" pageId="28" pageNumber="29">Fig. 20</figureCitation>
). The specimen is likely an adult considering its size (carapace length greater than
<quantity id="4C96D3F8C0281D68FD2EAFBDFCEE91F9" box="[767,841,1717,1744]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="cm" value="15.0">15 cm</quantity>
) and perhaps a male specimen, as modern male individuals of
<taxonomicName id="4C6E059EC0281D68FEBBAFD1FE7E91C4" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[362,473,1752,1778]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0281D68FEBBAFD1FE7E91C4" box="[362,473,1752,1778]" italics="true" pageId="28" pageNumber="29">H. thurjii</emphasis>
</taxonomicName>
reach up to
<quantity id="4C96D3F8C0281D68FDA3AFD0FD1A91C5" box="[626,701,1752,1779]" metricMagnitude="-1" metricUnit="m" metricValue="1.8" pageId="28" pageNumber="29" unit="cm" value="18.0">18 cm</quantity>
and lack intercostal fontanelles (
<bibRefCitation id="EFFF03ECC0281D68FB9BAFD0FACD91C4" author="Das I. &amp; Bhupathy S." box="[1098,1386,1752,1778]" pageId="28" pageNumber="29" pagination="5" refId="ref22842" refString="Das I. &amp; Bhupathy S. 2009 a. Hardella thurjii (Gray 1831) - Crowned River Turtle. Chelonian Research Monographs 5: 023.1 - 023.6. https: // doi. org / 10.3854 / crm. 5.023. thurjii. v 1.2009" type="journal article" year="2009">Das &amp; Bhupathy 2009a</bibRefCitation>
). Most sulci and sutures of the carapace are visible, as well as a knob on neurals IV and VIII, indicating the presence of a median keel. Growth annuli are present. The cervical scute is as wide as long and lacks a posterior notch. The first vertebral scute is wider than long, its lateral sides converge anteriorly and lack an anterolateral constriction, and it contacts the first marginal scute. The neural bones are hexagonal and anteriorly short-sided. The third vertebral has straight lateral sides and the posterior margin has an anteriorly oriented inflection that crosses the suture between neural bones III–IV. The fifth vertebral scute has an anterolateral constriction. The pygal bone is completely intersected by the twelfth intermarginal sulcus. The humeropectoral sulcus is located posterior to the entoplastron. The pectoral scute contacts the fifth marginal. For a more extensive description of this specimen, we refer to
<bibRefCitation id="EFFF03ECC0291D69FBA5A826FAEE967F" author="Lydekker R." box="[1140,1353,302,329]" pageId="29" pageNumber="30" pagination="540 - 541" refId="ref24567" refString="Lydekker R. 1886 a. On a new Emydine chelonian from the Pliocene of India. Quarterly Journal of the Geological Society 42: 540 - 541. https: // doi. org / 10.1144 / gsl. jgs. 1886.042.01 - 04.54" type="journal article" year="1886">Lydekker (1886a)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD17E1DC0291D54FF6CA878FAF8967C" blockId="29.[189,1399,368,465]" lastBlockId="32.[189,1399,267,330]" lastPageId="32" lastPageNumber="33" pageId="29" pageNumber="30">
BMNH R.890 (
<figureCitation id="13556298C0291D69FEA6A878FE6996BC" box="[375,462,368,394]" captionStart="Fig" captionStartId="31.[189,232,1930,1956]" captionTargetBox="[201,1392,512,1880]" captionTargetId="figure@31.[189,1398,506,1881]" captionTargetPageId="31" captionText="Fig. 21. BMNH R.890, referred to Hardella thurgi (Gray, 1831) by Lydekker (1889b), here identified as Hardella thurjii (Gray, 1831). A. Photograph of carapace. B. Illustration of carapace. C. Photograph of plastron. D. Illustration of plastron. Scale bar: 3 cm." figureDoi="http://doi.org/10.5281/zenodo.3860282" httpUri="https://zenodo.org/record/3860282/files/figure.png" pageId="29" pageNumber="30">Fig. 21</figureCitation>
) – This specimen is from the Miocene/Pliocene of the Siwalik Hills, likely of
<collectingCountry id="F3793E8DC0291D69FF6CA89BFF5B969B" box="[189,252,403,429]" name="India" pageId="29" pageNumber="30">India</collectingCountry>
, was purchased by P.T. Cautley in 1840, but remained unfigured to date. It is a crushed, partial specimen, that probably represents an adult female considering its large size (carapace length greater than
<quantity id="4C96D3F8C02B1D6BFF24A803FEE79610" box="[245,320,267,294]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="cm" value="20.0">20 cm</quantity>
). Both the anterior and posterior margins of the carapace are missing. Neural bones III and IV have a knob, indicating the presence of a median carapacial keel. Neural bones II–IV are hexagonal with anterior short sides. The first vertebral scute has anteriorly converging lateral sides. The third vertebral scute has straight lateral sulci. The posterior margins of the first and second pleurals are straight, placed over costals II and IV, respectively, and lack an anterior projection. Both the anterior and posterior plastral margins are not preserved. The plastron has two lateral longitudinal keels, which cross the lateral sides of the hyo- and hyoplastra. The humeropectoral sulcus is apparently located posterior to the entoplastron, which is not preserved. Inguinal scutes are likely present that contact the femoral scute.
</paragraph>
</subSubSection>
<caption id="DF112E95C0291D69FF6CAE81FC8790DC" ID-DOI="http://doi.org/10.5281/zenodo.3860278" ID-Zenodo-Dep="3860278" httpUri="https://zenodo.org/record/3860278/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,232,1929,1955]" targetBox="[199,1389,543,1875]" targetPageId="29">
<paragraph id="8BD17E1DC0291D69FF6CAE81FC8790DC" blockId="29.[189,1398,1928,2026]" pageId="29" pageNumber="30">
<emphasis id="B91AA20FC0291D69FF6CAE81FEBE9095" bold="true" box="[189,281,1929,1955]" pageId="29" pageNumber="30">Fig. 19.</emphasis>
BMNH 39835, holotype of
<taxonomicName id="4C6E059EC0291D69FDB6AE82FC5E9095" authority="Lydekker, 1885" authorityName="Lydekker" authorityYear="1885" box="[615,1017,1928,1955]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="falconeri">
<emphasis id="B91AA20FC0291D69FDB6AE82FC9F9095" box="[615,824,1929,1955]" italics="true" pageId="29" pageNumber="30">Batagur falconeri</emphasis>
Lydekker, 1885
</taxonomicName>
, here identified as a synonym of
<taxonomicName id="4C6E059EC0291D69FF6CAEA4FDB390F0" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[189,532,1964,1990]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0291D69FF6CAEA4FED190F0" box="[189,374,1964,1990]" italics="true" pageId="29" pageNumber="30">Hardella thurjii</emphasis>
(Gray, 1831)
</taxonomicName>
.
<emphasis id="B91AA20FC0291D69FDF2AEA4FD9F90F0" bold="true" box="[547,568,1964,1990]" pageId="29" pageNumber="30">A</emphasis>
. Photograph of carapace.
<emphasis id="B91AA20FC0291D69FCBAAEA4FCD890F0" bold="true" box="[875,895,1964,1990]" pageId="29" pageNumber="30">B</emphasis>
. Illustration of carapace.
<emphasis id="B91AA20FC0291D69FB7AAEA4FB6790F0" bold="true" box="[1195,1216,1964,1990]" pageId="29" pageNumber="30">C</emphasis>
. Photograph of plastron.
<emphasis id="B91AA20FC0291D69FEFBAEC7FE9890DF" bold="true" box="[298,319,1999,2025]" pageId="29" pageNumber="30">D</emphasis>
. Illustration of plastron. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="DF112E95C02A1D6AFF6CAE61FD1790D8" ID-DOI="http://doi.org/10.5281/zenodo.3860280" ID-Zenodo-Dep="3860280" httpUri="https://zenodo.org/record/3860280/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1897,1923]" targetBox="[194,1391,273,1848]" targetPageId="30">
<paragraph id="8BD17E1DC02A1D6AFF6CAE61FD1790D8" blockId="30.[189,1399,1897,2030]" pageId="30" pageNumber="31">
<emphasis id="B91AA20FC02A1D6AFF6CAE61FEBD90B5" bold="true" box="[189,282,1897,1923]" pageId="30" pageNumber="31">Fig. 20.</emphasis>
BMNH R.748, holotype of
<taxonomicName id="4C6E059EC02A1D6AFDBBAE62FBAE90B5" authority="Lydekker, 1886 a" authorityName="Lydekker" authorityYear="1886" box="[618,1033,1897,1924]" class="Reptilia" family="Emydidae" genus="Clemmys" kingdom="Animalia" order="Testudines" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="watsoni">
<emphasis id="B91AA20FC02A1D6AFDBBAE62FC9E90B5" box="[618,825,1898,1924]" italics="true" pageId="30" pageNumber="31">Clemmys watsoni</emphasis>
<bibRefCitation id="EFFF03ECC02A1D6AFC93AE61FBAE90B5" author="Lydekker R." box="[834,1033,1897,1923]" pageId="30" pageNumber="31" pagination="540 - 541" refId="ref24567" refString="Lydekker R. 1886 a. On a new Emydine chelonian from the Pliocene of India. Quarterly Journal of the Geological Society 42: 540 - 541. https: // doi. org / 10.1144 / gsl. jgs. 1886.042.01 - 04.54" type="journal article" year="1886">Lydekker, 1886a</bibRefCitation>
</taxonomicName>
, here identified as synonym of
<taxonomicName id="4C6E059EC02A1D6AFF6CAE85FDB89091" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[189,543,1933,1959]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC02A1D6AFF6CAE85FEDE9091" box="[189,377,1933,1959]" italics="true" pageId="30" pageNumber="31">Hardella thurjii</emphasis>
(Gray, 1831)
</taxonomicName>
.
<emphasis id="B91AA20FC02A1D6AFDFCAE85FDE59091" bold="true" box="[557,578,1933,1959]" pageId="30" pageNumber="31">A</emphasis>
. Photograph of carapace.
<emphasis id="B91AA20FC02A1D6AFCAEAE85FC349091" bold="true" box="[895,915,1933,1959]" pageId="30" pageNumber="31">B</emphasis>
. Illustration of carapace.
<emphasis id="B91AA20FC02A1D6AFB18AE85FB799091" bold="true" box="[1225,1246,1933,1959]" pageId="30" pageNumber="31">C</emphasis>
. Photograph of carapace in lateral view.
<emphasis id="B91AA20FC02A1D6AFDCAAEB8FD9790FC" bold="true" box="[539,560,1968,1994]" pageId="30" pageNumber="31">D</emphasis>
. Illustration of carapace in lateral view.
<emphasis id="B91AA20FC02A1D6AFBFEAEB8FBE490FC" bold="true" box="[1071,1091,1968,1994]" pageId="30" pageNumber="31">E</emphasis>
. Photograph of plastron.
<emphasis id="B91AA20FC02A1D6AFF6CAEDBFF6890DB" bold="true" box="[189,207,2003,2029]" pageId="30" pageNumber="31">F</emphasis>
. Illustration of plastron. Scale bar: 3 cm.
</paragraph>
</caption>
<caption id="DF112E95C02B1D6BFF6CAE82FC8790DD" ID-DOI="http://doi.org/10.5281/zenodo.3860282" ID-Zenodo-Dep="3860282" httpUri="https://zenodo.org/record/3860282/files/figure.png" pageId="31" pageNumber="32" startId="31.[189,232,1930,1956]" targetBox="[201,1392,512,1880]" targetPageId="31">
<paragraph id="8BD17E1DC02B1D6BFF6CAE82FC8790DD" blockId="31.[189,1398,1930,2027]" pageId="31" pageNumber="32">
<emphasis id="B91AA20FC02B1D6BFF6CAE82FEBE9092" bold="true" box="[189,281,1930,1956]" pageId="31" pageNumber="32">Fig. 21.</emphasis>
BMNH R.890, referred to
<taxonomicName id="4C6E059EC02B1D6BFD82AE82FC029092" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[595,933,1930,1956]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="thurgi">
<emphasis id="B91AA20FC02B1D6BFD82AE82FCAD9092" box="[595,778,1930,1956]" italics="true" pageId="31" pageNumber="32">Hardella thurgi</emphasis>
(Gray, 1831)
</taxonomicName>
by
<bibRefCitation id="EFFF03ECC02B1D6BFC1EAE82FB029093" author="Lydekker R." box="[975,1189,1930,1957]" pageId="31" pageNumber="32" refId="ref24792" refString="Lydekker R. 1889 b. Catalogue of the Fossil Reptilia and Amphibia in the British Museum (Natural History). Part III. Chelonia. Order of the Trustees, London." type="book" year="1889">Lydekker (1889b)</bibRefCitation>
, here identified as
<taxonomicName id="4C6E059EC02B1D6BFF6CAEA6FDB090FE" authority="(Gray, 1831)" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[189,535,1966,1992]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC02B1D6BFF6CAEA6FED190FE" box="[189,374,1966,1992]" italics="true" pageId="31" pageNumber="32">Hardella thurjii</emphasis>
(Gray, 1831)
</taxonomicName>
.
<emphasis id="B91AA20FC02B1D6BFDF2AEA5FD9F90F1" bold="true" box="[547,568,1965,1991]" pageId="31" pageNumber="32">A</emphasis>
. Photograph of carapace.
<emphasis id="B91AA20FC02B1D6BFCBAAEA6FCD890FE" bold="true" box="[875,895,1966,1992]" pageId="31" pageNumber="32">B</emphasis>
. Illustration of carapace.
<emphasis id="B91AA20FC02B1D6BFB7AAEA5FB6790F1" bold="true" box="[1195,1216,1965,1991]" pageId="31" pageNumber="32">C</emphasis>
. Photograph of plastron.
<emphasis id="B91AA20FC02B1D6BFEFBAED9FE9890DD" bold="true" box="[298,319,2001,2027]" pageId="31" pageNumber="32">D</emphasis>
. Illustration of plastron. Scale bar: 3 cm.
</paragraph>
</caption>
<subSubSection id="C3742D96C0141D54FF6CA879FB2495A7" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BD17E1DC0141D54FF6CA879FEE396BD" blockId="32.[189,324,369,395]" box="[189,324,369,395]" pageId="32" pageNumber="33">
<heading id="D099C971C0141D54FF6CA879FEE396BD" bold="true" box="[189,324,369,395]" fontSize="11" level="3" pageId="32" pageNumber="33" reason="3">
<emphasis id="B91AA20FC0141D54FF6CA879FEE396BD" bold="true" box="[189,324,369,395]" pageId="32" pageNumber="33">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BD17E1DC0141D54FF6CA8A9FB2495A7" blockId="32.[189,1399,416,657]" pageId="32" pageNumber="33">
We here attribute these specimens to
<taxonomicName id="4C6E059EC0141D54FDBAA8A8FC84968C" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[619,803,416,442]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0141D54FDBAA8A8FC84968C" box="[619,803,416,442]" italics="true" pageId="32" pageNumber="33">Hardella thurjii</emphasis>
</taxonomicName>
based on the presence of a short median keel, large and square second and third vertebrals that are about the same width as the fourth vertebral scute, large inguinal scute, and an entoplastron that is not intersected by the humeropectoral sulcus. This confirms the synonymy of
<taxonomicName id="4C6E059EC0141D54FE59AB04FDB59513" authorityName="Lydekker" authorityYear="1885" box="[392,530,523,549]" class="Reptilia" family="Geoemydidae" genus="Batagur" kingdom="Animalia" order="Testudines" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="falconeri">
<emphasis id="B91AA20FC0141D54FE59AB04FDB59513" box="[392,530,523,549]" italics="true" pageId="32" pageNumber="33">B. falconeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E059EC0141D54FD9BAB04FD609513" authorityName="Lydekker" authorityYear="1886" box="[586,711,524,550]" class="Reptilia" family="Emydidae" genus="Clemmys" kingdom="Animalia" order="Testudines" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="watsoni">
<emphasis id="B91AA20FC0141D54FD9BAB04FD609513" box="[586,711,524,550]" italics="true" pageId="32" pageNumber="33">C. watsoni</emphasis>
</taxonomicName>
with
<taxonomicName id="4C6E059EC0141D54FCD8AB04FCD29513" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[777,885,523,549]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0141D54FCD8AB04FCD29513" box="[777,885,523,549]" italics="true" pageId="32" pageNumber="33">H. thurjii</emphasis>
</taxonomicName>
, as first suggested by
<bibRefCitation id="EFFF03ECC0141D54FBA7AB03FAE29510" author="Boulenger G. A." box="[1142,1349,523,550]" pageId="32" pageNumber="33" refId="ref22539" refString="Boulenger G. A. 1889. Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum (Natural History). Trustees of the British Museum (Natural History), London." type="book" year="1889">Boulenger (1889)</bibRefCitation>
and later supported by
<bibRefCitation id="EFFF03ECC0141D54FE71AB27FDD0957F" author="Lydekker R." box="[416,631,559,585]" pageId="32" pageNumber="33" pagination="56 - 59" refId="ref24763" refString="Lydekker R. 1889 a. Notes on Siwalik and Narbada Chelonia. Records of the Geological Survey of India 22 (1): 56 - 59." type="journal article" year="1889">Lydekker (1889a)</bibRefCitation>
, TEWG (2015) and TTWG (2017). We furthermore follow
<bibRefCitation id="EFFF03ECC0141D54FA98AB38FEAC955B" author="Das I." pageId="32" pageNumber="33" pagination="41 - 46" refId="ref22808" refString="Das I. 1994. The identity of the Plio-Pleistocene turtle, Geoemyda pilgrimi Prasad and Satsangi, 1967 (Testudines: Cryptodira: Bataguridae). Hamadryad 19: 41 - 46." type="journal article" year="1994">Das (1994)</bibRefCitation>
by recognizing the synonymy of
<taxonomicName id="4C6E059EC0141D54FD4CAB5BFC25955B" authorityName="Prasad &amp; Satsangi" authorityYear="1967" box="[669,898,595,621]" class="Reptilia" family="Geoemydidae" genus="Geoemyda" kingdom="Animalia" order="Testudines" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="pilgrimi">
<emphasis id="B91AA20FC0141D54FD4CAB5BFC25955B" box="[669,898,595,621]" italics="true" pageId="32" pageNumber="33">Geoemyda pilgrimi</emphasis>
</taxonomicName>
with
<taxonomicName id="4C6E059EC0141D54FC16AB5CFB92955B" baseAuthorityName="Gray" baseAuthorityYear="1831" box="[967,1077,595,621]" class="Reptilia" family="Geoemydidae" genus="Hardella" kingdom="Animalia" order="Testudines" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="thurjii">
<emphasis id="B91AA20FC0141D54FC16AB5CFB92955B" box="[967,1077,595,621]" italics="true" pageId="32" pageNumber="33">H. thurjii</emphasis>
</taxonomicName>
(followed by TEWG 2015 and TTWG 2017) although we did not study the
<typeStatus id="54D5C0BFC0141D54FD28AB7FFCC795A7" box="[761,864,631,657]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
of this species firsthand.
</paragraph>
</subSubSection>
</treatment>
</document>
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