170 lines
20 KiB
XML
170 lines
20 KiB
XML
<document id="01FAE7DE18B1AF4ED88B8FCFECB7BB81" ID-CLB-Dataset="8562" ID-DOI="10.5252/geodiversitas2020v42a20" ID-GBIF-Dataset="ad4dbfb6-14ce-48e2-8fe9-99fa26a9bcf4" ID-ISSN="1638-9395" ID-Zenodo-Dep="4447563" ID-ZooBank="urn:lsid:zoobank.org:pub:8FF2A078-CE45-4BF1-A681-00136F57375E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610989940068" checkinUser="felipe" docAuthor="Ivanov, Martin, Čerňanský, Andrej, Bonilla-Salomón, Isaac & Luján, Àngel Hernández" docDate="2020" docId="03C587C7431CFFFCFEEAFEEB49BEF839" docLanguage="en" docName="Geodiversitas.42.20.343-376.pdf" docOrigin="Geodiversitas 42 (20)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Natricinae Bonaparte 1838" docType="treatment" docVersion="10" lastPageNumber="365" masterDocId="FFFCFFBF4304FFE4FFBCFF9C484CFFD2" masterDocTitle="Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications" masterLastPageNumber="376" masterPageNumber="343" pageNumber="365" updateTime="1698925529958" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="ECBDCD8A8A87D7EE895E3AA41A951F3E">Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications</mods:title>
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<mods:namePart id="A2EEC5C249C62B03C3A9AC06498A2C6C">Ivanov, Martin</mods:namePart>
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<mods:namePart id="68228924C8F448CBD1A5787EDD73727E">Čerňanský, Andrej</mods:namePart>
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<mods:namePart id="CD2A06D48F9C361B679637D563CDCFB0">Bonilla-Salomón, Isaac</mods:namePart>
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<mods:namePart id="5FBD185D7A60AB4B67C5D056F3D5EB09">Luján, Àngel Hernández</mods:namePart>
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<mods:title id="3609E7725A9064CC5CAA36B0EDC3BC09">Geodiversitas</mods:title>
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<mods:part id="F444C2E9925E43FE58A47B058C934C6C">
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<mods:date id="39A8B38817C49F9D58DDC6E1D810ED21">2020</mods:date>
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<mods:number id="07375CB56AC224B65013C6D9DF06D35F">2020-08-28</mods:number>
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<mods:number id="870E08288F85A627491AA289590A4000">42</mods:number>
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<mods:number id="50F717741872E55C7F51B5EF0B7CE496">20</mods:number>
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<mods:start id="611940CEF652FC5F2161F3078D0FA3E7">343</mods:start>
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<treatment id="03C587C7431CFFFCFEEAFEEB49BEF839" ID-DOI="http://doi.org/10.5281/zenodo.4488233" ID-GBIF-Taxon="177200722" ID-Zenodo-Dep="4488233" LSID="urn:lsid:plazi:treatment:03C587C7431CFFFCFEEAFEEB49BEF839" httpUri="http://treatment.plazi.org/id/03C587C7431CFFFCFEEAFEEB49BEF839" lastPageNumber="365" pageId="24" pageNumber="365">
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<subSubSection id="C376655A431CFFFCFEEAFEEB4A7AFE43" box="[342,566,375,401]" pageId="24" pageNumber="365" type="nomenclature">
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<paragraph id="8BD336D1431CFFFCFEEAFEEB4A7AFE43" blockId="24.[342,566,375,401]" box="[342,566,375,401]" pageId="24" pageNumber="365">
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<heading id="D09B81BD431CFFFCFEEAFEEB4A7AFE43" box="[342,566,375,401]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="365" reason="7">
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“
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<taxonomicName id="4C6C4D52431CFFFCFEDDFEEB49A9FE5D" authorityName="Bonaparte" authorityYear="1838" box="[353,485,375,401]" class="Reptilia" family="Natricinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="365" phylum="Chordata" rank="family">NATRICINAE</taxonomicName>
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” indet.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A431CFFFCFE39FE0B4A3EFE3A" pageId="24" pageNumber="365" type="description">
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||
<paragraph id="8BD336D1431CFFFCFE39FE0B4A49FE63" blockId="24.[389,517,406,433]" box="[389,517,406,433]" pageId="24" pageNumber="365">
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(
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<figureCitation id="13572A54431CFFFCFE32FE0A4997FE63" box="[398,475,406,433]" captionStart="FIG" captionStartId="23.[132,143,1530,1548]" captionTargetBox="[132,1455,215,1487]" captionTargetId="figure-224@23.[698,1267,758,1306]" captionTargetPageId="23" captionText="FIG. 12. — Natrix sp. and “Natricinae” indet. from the early Miocene (MN 4) of MWQ. Natrix sp.: trunk vertebra Pal. 1968 (2/2003 Reptile Joint) in left lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views.“Natricinae” indet.:trunk vertebra Pal. 1483 (1/2001 Turtle Joint) with preserved hypapophysis in left lateral (F), dorsal (G), ventral (H) and caudal (I) views. Abbreviations: ak, anterior keel; cd, condyle; ct, cotyle; es, epizypapophyseal spine; hy, hypapophysis; lf, lateral foramen;nc, neural canal;ns, neural spine;pctf, paracotylar foramen;po, postzygapophysis;pof, postzygapophyseal articular facet;prf, prezygapophyseal articular facet; prp, prezygapophyseal process;scf, subcentral foramen;scg, subcentral groove;scr, subcentral ridge;zy, zygosphene;zyg, zygantrum.Scale bars:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447589" httpUri="https://zenodo.org/record/4447589/files/figure.png" pageId="24" pageNumber="365">Fig. 12</figureCitation>
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F-I)
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</paragraph>
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<paragraph id="8BD336D1431CFFFCFF38FE534A3EFE3A" blockId="24.[132,626,463,488]" box="[132,626,463,488]" pageId="24" pageNumber="365">
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cf.
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<taxonomicName id="4C6C4D52431CFFFCFF1EFE4C494CFE35" authorityName="Holman" authorityYear="1973" box="[162,256,464,487]" class="Reptilia" family="Colubridae" genus="Neonatrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="365" phylum="Chordata" rank="genus">
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<emphasis id="B918EAC3431CFFFCFF1EFE4C494CFE35" box="[162,256,464,487]" italics="true" pageId="24" pageNumber="365">Neonatrix</emphasis>
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</taxonomicName>
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sp. –
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<bibRefCitation id="EFFD4B20431CFFFCFE87FE4C4A52FE3A" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[315,542,463,488]" pageId="24" pageNumber="365" pagination="223 - 239" refId="ref24955" refString="IVANOV M., MUSIL R. & BRZOBOHATY R. 2006. - Terrestrial and Marine Faunas from the Miocene Deposits of the Mokra Plateau (Drahany Upland, Czech Republic) - Impact on Palaeogeography. Beitrage zur Palaontologie 30: 223 - 239." type="journal article" year="2006">
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Ivanov
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<emphasis id="B918EAC3431CFFFCFE3FFE4C49FEFE35" box="[387,434,463,487]" italics="true" pageId="24" pageNumber="365">et al.</emphasis>
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2006: 229
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</bibRefCitation>
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, table 2.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376655A431CFFFCFF38FD9C4AC7FD9E" pageId="24" pageNumber="365" type="materials_examined">
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<paragraph id="8BD336D1431CFFFCFF38FD9C4AC7FD9E" blockId="24.[131,775,511,588]" pageId="24" pageNumber="365">
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||
MATERIAL. —
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<materialsCitation id="3B043C8C431CFFFCFE9FFE634AC7FD9E" ID-GBIF-Occurrence="3027945317" pageId="24" pageNumber="365">
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MWQ, early Miocene, Burdigalian, Orleanian,
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<collectionCode id="ED7DAE14431CFFFCFF3AFD8648FDFDE3" box="[134,177,538,561]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="24" pageNumber="365">MN</collectionCode>
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4: 1/2001 Turtle Joint: One trunk vertebra (Pal. 1483). 2/2003 Reptile Joint: Four trunk vertebrae (Pal. 1970-1973).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376655A431CFFFCFF38FDEA4A1FFA9C" pageId="24" pageNumber="365" type="description">
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||
<paragraph id="8BD336D1431CFFFCFF38FDEA4959FD5C" blockId="24.[130,777,630,1358]" box="[132,277,630,656]" pageId="24" pageNumber="365">DESCRIPTION</paragraph>
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<paragraph id="8BD336D1431CFFFCFF38FD0A4965FD62" blockId="24.[130,777,630,1358]" box="[132,297,662,688]" pageId="24" pageNumber="365">
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<emphasis id="B918EAC3431CFFFCFF38FD0A4965FD62" box="[132,297,662,688]" italics="true" pageId="24" pageNumber="365">Trunk vertebrae</emphasis>
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</paragraph>
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<paragraph id="8BD336D1431CFFFCFF38FD294A1FFA9C" blockId="24.[130,777,630,1358]" pageId="24" pageNumber="365">
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The most complete vertebra (Pal. 1483) has strongly damaged prezygapophyses and the zygosphene, and the neural spine is broken-off close to its base.In lateral view,the interzygapophyseal ridges are moderately developed and lateral foramina are situated within deep depressions.The subcentral ridges are nearly straight. The distal termination of sigmoid hypapophysis does not reach posterior to the caudal border of condyle. In dorsal view, the vertebra is cylindrical, the damaged zygosphene was wide relative to the neural arch width. Epizygapophyseal ridges are absent. In ventral view, the subcentral grooves are very shallow and short and the subcentral foramina are rather small and situated just posterior to the base of the hypapophysis. The subcotylar tubercles are missing.The damaged postzygapophyseal articular facets have a subrectangular outline and are not elongated laterally. In cranial view, the neural arch is slightly vaulted, and the large neural canal has developed conspicuously large and wide lateral sinuses.The cranial margin of the zygosphene was arched dorsally.The small paracotylar foramina are situated within wide depressions on either side of the rounded cotyle. The vertebral dimensions of the best-preserved specimen (Pal. 1483) are: cl =
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<quantity id="4C949B34431CFFFCFF25FAAF48B1FA9C" box="[153,253,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.51" pageId="24" pageNumber="365" unit="mm" value="3.51">3.51 mm</quantity>
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; naw =
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<quantity id="4C949B34431CFFFCFEF1FAAF49FDFA9C" box="[333,433,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="24" pageNumber="365" unit="mm" value="2.24">2.24 mm</quantity>
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; cl/naw = 1.57.
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A431CFFFCFF38FAE849BEF839" pageId="24" pageNumber="365" type="discussion">
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<paragraph id="8BD336D1431CFFFCFF38FAE848A4FA5E" blockId="24.[130,776,1396,2028]" box="[132,232,1396,1422]" pageId="24" pageNumber="365">REMARKS</paragraph>
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<paragraph id="8BD336D1431CFFFCFF38FA0F49BEF839" blockId="24.[130,776,1396,2028]" pageId="24" pageNumber="365">
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The sigmoid shape of the hypapophysis, with an anterior keel sloping towards the cotylar rim is reminiscent of precaudal vertebrae of the genus
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<taxonomicName id="4C6C4D52431CFFFCFEF5FA4F4A26FA3F" authority="Szyndlar, 1982" authorityName="Szyndlar" authorityYear="1982" box="[329,618,1491,1517]" class="Reptilia" family="Colubridae" genus="Palaeonatrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="365" phylum="Chordata" rank="genus">
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<emphasis id="B918EAC3431CFFFCFEF5FA4F4984FA3F" box="[329,456,1491,1517]" italics="true" pageId="24" pageNumber="365">Palaeonatrix</emphasis>
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Szyndlar, 1982
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</taxonomicName>
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(seeMłynarski
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<emphasis id="B918EAC3431CFFFCFF38FA6F48FAF9DF" box="[132,182,1523,1549]" italics="true" pageId="24" pageNumber="365">et al.</emphasis>
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1982) reported from the early Miocene (MN 4) of Dolnice,
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<collectingCountry id="F37B7641431CFFFCFF02F98E492BF9FE" box="[190,359,1554,1580]" name="Czech Republic" pageId="24" pageNumber="365">Czech Republic</collectingCountry>
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(
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<bibRefCitation id="EFFD4B20431CFFFCFEC9F98F4A03F9FF" author="RAGE J. - C. & ROCEK Z." box="[373,591,1554,1581]" pageId="24" pageNumber="365" pagination="17 - 21" refId="ref26481" refString="RAGE J. - C. & ROCEK Z. 1983. - Dolniceophis lehmani (Serpentes, Colubridae), a new fossil snake from the Lower Miocene of Czechoslovakia. Casopis pro mineralalogii a geologii 28: 17 - 21." type="journal article" year="1983">Rage & Roček 1983</bibRefCitation>
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;
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<bibRefCitation id="EFFD4B20431CFFFCFDE0F98E4ABBF9FF" author="SZYNDLAR Z." box="[604,759,1554,1581]" pageId="24" pageNumber="365" pagination="55 - 71" refId="ref27331" refString="SZYNDLAR Z. 1987. - Snakes from the Lower Miocene locality of Dolnice (Czechoslovakia). Journal of Vertebrate Paleontology 7 (1): 55 - 71. https: // doi. org / 10.1080 / 02724634.1987.10011637" type="journal article" year="1987">Szyndlar 1987</bibRefCitation>
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), Petersbuch 2 and Langenau,
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<collectingCountry id="F37B7641431CFFFCFE04F9AF4A51F99F" box="[440,541,1587,1613]" name="Germany" pageId="24" pageNumber="365">Germany</collectingCountry>
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(
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<bibRefCitation id="EFFD4B20431CFFFCFD91F9AE48F7F9BE" author="SZYNDLAR Z. & SCHLEICH H. H." pageId="24" pageNumber="365" pagination="1 - 47" refId="ref27799" refString="SZYNDLAR Z. & SCHLEICH H. H. 1993. - Description of Miocene snakes from Petersbuch 2 with comments on the Lower and Middle Miocene ophidian faunas of Southern Germany. Stuttgarter Beitrage zur Naturkunde Serie B (Geologie und Palaontologie) 192: 1 - 47." type="journal article" year="1993">Szyndlar & Schleich 1993</bibRefCitation>
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), Oberdorf,
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<collectingCountry id="F37B7641431CFFFCFEF0F9CE49D6F9BE" box="[332,410,1618,1644]" name="Austria" pageId="24" pageNumber="365">Austria</collectingCountry>
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(
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<bibRefCitation id="EFFD4B20431CFFFCFE10F9CE4A07F9BE" author="SZYNDLAR Z." box="[428,587,1618,1644]" pageId="24" pageNumber="365" pagination="31 - 38" refId="ref27605" refString="SZYNDLAR Z. 1998. - Vertebrates from the Early Miocene lignite deposits of the opencast mine Oberdorf (Western Styrian Basin, Austria). Annalen des Naturhistorischen Museums inWien A 99: 31 - 38." type="journal article" year="1998">Szyndlar 1998</bibRefCitation>
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) and the middle Miocene of Sansan,
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<collectingCountry id="F37B7641431CFFFCFED2F9EF49F4F95F" box="[366,440,1651,1677]" name="France" pageId="24" pageNumber="365">France</collectingCountry>
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(MN 6;
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<bibRefCitation id="EFFD4B20431CFFFCFD98F9EE4AB2F95E" author="AUGE M. & RAGE J. - C." box="[548,766,1650,1677]" pageId="24" pageNumber="365" refId="ref22449" refString="AUGE M. & RAGE J. - C. 2000. - Les Squamates (Reptilia) du Miocene moyen de Sansan (Gers, France), in GINSBURG L. (ed.), La faune miocene de Sansan et son environnement. Museum national d'Histoire naturelle, Paris: 263 - 313 (Memoires du Museum national d'Histoire naturelle; 183)." type="journal volume" year="2000">Augé & Rage 2000</bibRefCitation>
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) and
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<collectingRegion id="49A8F833431CFFFCFF0FF90E48B6F97E" box="[179,250,1682,1708]" country="Poland" name="Opolskie" pageId="24" pageNumber="365">Opole</collectingRegion>
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,
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<collectingCountry id="F37B7641431CFFFCFEB8F90E4903F97E" box="[260,335,1682,1708]" name="Poland" pageId="24" pageNumber="365">Poland</collectingCountry>
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(MN 7+8;
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<bibRefCitation id="EFFD4B20431CFFFCFE77F90E4AFEF97E" author="MLYNARSKI M. & SZYNDLAR Z. & ESTES R. & SANCHIZ B." box="[459,690,1682,1708]" pageId="24" pageNumber="365" pagination="103 - 119" refId="ref25858" refString="MLYNARSKI M., SZYNDLAR Z., ESTES R. & SANCHIZ B. 1982. - Lower vertebrate fauna from the Miocene of Opole (Poland). Estudios geologicos 38 (1 - 2): 103 - 119." type="journal article" year="1982">
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Młynarski
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<emphasis id="B918EAC3431CFFFCFDFDF90F4A38F97E" box="[577,628,1682,1708]" italics="true" pageId="24" pageNumber="365">et al.</emphasis>
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1982
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</bibRefCitation>
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). However, a generic allocation is impossible because the relatively low neural spine which characterizes the genus
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<taxonomicName id="4C6C4D52431CFFFCFD39F94E4B4BF93E" authorityName="Szyndlar" authorityYear="1982" box="[645,775,1746,1772]" class="Reptilia" family="Colubridae" genus="Palaeonatrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="365" phylum="Chordata" rank="genus">
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<emphasis id="B918EAC3431CFFFCFD39F94E4B4BF93E" box="[645,775,1746,1772]" italics="true" pageId="24" pageNumber="365">Palaeonatrix</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFFD4B20431CFFFCFF31F96E4969F8DE" author="SZYNDLAR Z." box="[141,293,1778,1804]" pageId="24" pageNumber="365" pagination="55 - 71" refId="ref27331" refString="SZYNDLAR Z. 1987. - Snakes from the Lower Miocene locality of Dolnice (Czechoslovakia). Journal of Vertebrate Paleontology 7 (1): 55 - 71. https: // doi. org / 10.1080 / 02724634.1987.10011637" type="journal article" year="1987">Szyndlar 1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFD4B20431CFFFCFE8DF96E493AF8DE" author="SZYNDLAR Z." box="[305,374,1778,1804]" pageId="24" pageNumber="365" pagination="237 - 266" refId="ref27471" refString="SZYNDLAR Z. 1991 b. - A rewiew of Neogene and Quaternary snakes of Central and Eastern Europe. Part II: Natricinae, Elapidae, Viperidae. Estudios geologicos 47 (3 - 4): 237 - 266. https: // doi. org / 10.3989 / egeol. 91473 - 4422" type="journal article" year="1991">1991b</bibRefCitation>
|
||
) is not preserved in studied specimens from MWQ. Although the very strongly developed subcentral ridges are typical for the genus
|
||
<taxonomicName id="4C6C4D52431CFFFCFE6CF8AD4B4BF89E" authority="(Szyndlar 1987)" baseAuthorityName="Szyndlar" baseAuthorityYear="1987" box="[464,775,1841,1868]" class="Reptilia" family="Colubridae" genus="Palaeonatrix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="365" phylum="Chordata" rank="genus">
|
||
<emphasis id="B918EAC3431CFFFCFE6CF8AD4A1EF899" box="[464,594,1841,1867]" italics="true" pageId="24" pageNumber="365">Palaeonatrix</emphasis>
|
||
(
|
||
<bibRefCitation id="EFFD4B20431CFFFCFDDEF8AD4AB2F89E" author="SZYNDLAR Z." box="[610,766,1841,1868]" pageId="24" pageNumber="365" pagination="55 - 71" refId="ref27331" refString="SZYNDLAR Z. 1987. - Snakes from the Lower Miocene locality of Dolnice (Czechoslovakia). Journal of Vertebrate Paleontology 7 (1): 55 - 71. https: // doi. org / 10.1080 / 02724634.1987.10011637" type="journal article" year="1987">Szyndlar 1987</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
we cannot exclude the possibility that the subcentral ridges were shorter and less distinct in vertebrae situated anterior to the preserved vertebra. Because of the limited fossil material and lack of information on intracolumnar variability, a more precise assignment is not possible.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |