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<document id="808BE48A3F943D79433545046D42A353" ID-CLB-Dataset="55973" ID-DOI="10.23885/181433262022181-3133" ID-GBIF-Dataset="5b23f969-df9f-416b-923f-4b5ce59f24e0" ID-Zenodo-Dep="6380484" ID-ZooBank="urn:lsid:zoobank.org:pub:732CCF3D-E009-4757-A83B-C755CE5F2FCE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649953843487" checkinUser="tatiana" docAuthor="Löbl, I." docDate="2022" docId="03C587B1FF8202430DA482E6FD2BCC01" docLanguage="en" docName="CaucEntomolBull.18.1.31-33.pdf" docOrigin="Caucasian Entomological Bulletin (Caucas. entomol. bull.) 18 (1)" docSource="http://dx.doi.org/10.23885/181433262022181-3133" docStyle="DocumentStyle:6A2F040C6DCB19A13F917346BF1BCB54.1:CaucEntomolBull.2017-.journal_article" docStyleId="6A2F040C6DCB19A13F917346BF1BCB54" docStyleName="CaucEntomolBull.2017-.journal_article" docStyleVersion="1" docTitle="Scaphisoma caucasicum Löbl 2022, sp. n." docType="treatment" docUuid="B94C66F5-7786-4A2C-81C7-5F3EA5478903" docUuidSource="ZooBank" docVersion="7" lastPageNumber="33" masterDocId="FFFCFFC9FF8302400E1D844CFFAFC80D" masterDocTitle="On the genus Scaphisoma Leach, 1815 (Coleoptera: Staphylinidae: Scaphidiinae) from Georgia, with description of a new species" masterLastPageNumber="33" masterPageNumber="31" pageNumber="31" updateTime="1699328934869" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F1C92A60572B322B80BBC5748FA7AF32">On the genus Scaphisoma Leach, 1815 (Coleoptera: Staphylinidae: Scaphidiinae) from Georgia, with description of a new species</mods:title>
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<mods:namePart id="90D3449CC325355DF78C0E0BDD14B93F">Löbl, I.</mods:namePart>
<mods:affiliation id="DC906A45E8B7C86775D932EB1B9D1E5C">Muséum dhistoire naturelle, case postale 6434, Genève 1211 Switzerland.</mods:affiliation>
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<mods:title id="39D454A7D553E77D544DE220E5527F93">Caucasian Entomological Bulletin</mods:title>
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<mods:date id="3E40BD2B58D9642FFBF5F8B82255785A">2022</mods:date>
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<mods:title id="FE00ED2B9CA66C2E835B9B7DE4B8F911">Caucas. entomol. bull.</mods:title>
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<mods:number id="D06D7E07E2B0C048E2205CFCEC6FF04A">2022-03-23</mods:number>
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<treatment id="03C587B1FF8202430DA482E6FD2BCC01" ID-DOI="http://doi.org/10.5281/zenodo.6461701" ID-GBIF-Taxon="194601561" ID-Zenodo-Dep="6461701" LSID="urn:lsid:zoobank.org:act:B94C66F5-7786-4A2C-81C7-5F3EA5478903" httpUri="http://treatment.plazi.org/id/03C587B1FF8202430DA482E6FD2BCC01" lastPageId="3" lastPageNumber="33" pageId="1" pageNumber="31">
<subSubSection id="C376652CFF8202410DA482E6FB45CEB2" box="[953,1258,1706,1727]" pageId="1" pageNumber="31" type="nomenclature">
<paragraph id="8BD336A7FF8202410DA482E6FB45CEB2" blockId="1.[953,1258,1706,1757]" box="[953,1258,1706,1727]" pageId="1" pageNumber="31">
<heading id="D09B81CBFF8202410DA482E6FB45CEB2" box="[953,1258,1706,1727]" centered="true" fontSize="9" level="7" pageId="1" pageNumber="31" reason="1">
<taxonomicName id="4C6C4D24FF8202410DA482E6FB03CEB2" ID-CoL="9GD3F" authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[953,1196,1706,1727]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="31" phylum="Arthropoda" rank="species" species="caucasicum" status="sp. nov.">
<emphasis id="B918EAB5FF8202410DA482E6FB03CEB2" box="[953,1196,1706,1727]" italics="true" pageId="1" pageNumber="31">Scaphisoma caucasicum</emphasis>
</taxonomicName>
<emphasis id="B918EAB5FF8202410AAF82E6FB45CEB2" bold="true" box="[1202,1258,1706,1727]" pageId="1" pageNumber="31">
<taxonomicNameLabel id="A22B57CEFF8202410AAF82E6FB45CEB2" box="[1202,1258,1706,1727]" pageId="1" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C376652CFF8202410A3D828BFB2CCED0" box="[1056,1155,1735,1757]" pageId="1" pageNumber="31" type="description">
<paragraph id="8BD336A7FF8202410A3D828BFB2CCED0" blockId="1.[953,1258,1706,1757]" box="[1056,1155,1735,1757]" pageId="1" pageNumber="31">
<heading id="D09B81CBFF8202410A3D828BFB2CCED0" box="[1056,1155,1735,1757]" centered="true" fontSize="9" level="7" pageId="1" pageNumber="31" reason="1">
(
<figureCitation id="13572A22FF8202410A3B828BFBD4CED0" box="[1062,1147,1735,1757]" captionStart="Figs 13" captionStartId="2.[237,268,1317,1334]" captionTargetBox="[195,1389,273,1302]" captionTargetId="figure-418@2.[189,1399,272,1308]" captionTargetPageId="2" captionText="Figs 13. Scaphisoma caucasicum sp. n., aedeagus. 12 aedeagus: 1 dorsal view, 2 lateral view; 3 internal sac. Scale bars 0.1 mm. Рис. 13. Scaphisoma caucasicum sp. n., эдеагус. 12 эдеагус: 1 вид сверху, 2 вид сбоку; 3 внутренний мешок. Масштабные линейки 0.1 мм." pageId="1" pageNumber="31">Figs 13</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376652CFF8202410D4182B2FB37CF31" pageId="1" pageNumber="31" type="materials_examined">
<paragraph id="8BD336A7FF8202410D4182B2FB37CF31" blockId="1.[813,1400,1789,1955]" pageId="1" pageNumber="31">
<emphasis id="B918EAB5FF8202410D4182B2FC06CF02" bold="true" box="[860,937,1790,1807]" pageId="1" pageNumber="31">Material.</emphasis>
<materialsCitation id="3B043CFAFF8202410DAC82B2FB37CF31" ID-GBIF-Occurrence="3751725314" collectingDate="2021-10-17" collectionCode="ZMB" collectorName="M. Schulke" country="Georgia" latitude="42.620277" location="Martvili" longitude="42.379166" municipality="Svaneti" pageId="1" pageNumber="31" specimenCount="511" specimenCount-male="511" typeStatus="holotype">
<typeStatus id="54D78805FF8202410DAC82B2FC54CF02" box="[945,1019,1790,1807]" pageId="1" pageNumber="31" type="holotype">Holotype</typeStatus>
,
<emphasis id="B918EAB5FF8202410A1A82B3FBBACF1C" bold="true" box="[1031,1045,1791,1809]" italics="true" pageId="1" pageNumber="31">
<specimenCount id="9D6AFD2EFF8202410A1A82B3FBBACF1C" box="[1031,1045,1791,1809]" pageId="1" pageNumber="31" type="male"></specimenCount>
</emphasis>
(
<collectionCode id="ED7DAE62FF8202410A3982B1FBFFCF02" box="[1060,1104,1789,1807]" pageId="1" pageNumber="31">ZMB</collectionCode>
): “
<collectingCountry id="F37B7637FF8202410A7A82B2FB11CF02" box="[1127,1214,1790,1807]" pageId="1" pageNumber="31">GEORGIA</collectingCountry>
[GE2021-60]:
<collectingMunicipality id="6BB7ACDDFF8202410B2482B2FADCCF02" box="[1337,1395,1790,1807]" pageId="1" pageNumber="31">Svaneti</collectingMunicipality>
, N
<location id="8EB3607CFF8202410D5F8359FC2CCF2B" LSID="urn:lsid:plazi:treatment:03C587B1FF8202430DA482E6FD2BCC01:8EB3607CFF8202410D5F8359FC2CCF2B" box="[834,899,1813,1830]" country="Georgia" municipality="Svaneti" name="Martvili" pageId="1" pageNumber="31">Martvili</location>
,
<location id="8EB3607CFF8202410D908359FBAECF2B" LSID="urn:lsid:plazi:treatment:03C587B1FF8202430DA482E6FD2BCC01:8EB3607CFF8202410D908359FBAECF2B" box="[909,1025,1813,1830]" country="Georgia" municipality="Svaneti" name="Lebarde valley" pageId="1" pageNumber="31">Lebarde valley</location>
,
<geoCoordinate id="EE585060FF8202410A168359FBCDCF28" box="[1035,1122,1812,1830]" degrees="42" direction="north" minutes="37" orientation="latitude" pageId="1" pageNumber="31" precision="15" seconds="13" value="42.620277">42°37ʹ13ʺ N</geoCoordinate>
,
<geoCoordinate id="EE585060FF8202410A708359FB6DCF28" box="[1133,1218,1812,1830]" degrees="42" direction="east" minutes="22" orientation="longitude" pageId="1" pageNumber="31" precision="15" seconds="45" value="42.379166">42°22ʹ45ʺ E</geoCoordinate>
,
<specimenCount id="9D6AFD2EFF8202410AD18359FAAECF28" box="[1228,1281,1813,1830]" pageId="1" pageNumber="31" type="male">510 m</specimenCount>
, track margin, mushroomssifted,
<collectingDate id="EF96E98FFF8202410DDF8367FBBECF31" box="[962,1041,1835,1852]" pageId="1" pageNumber="31">
<date id="FFD21067FF8202410DDF8367FBBECF31" box="[962,1041,1835,1852]" pageId="1" pageNumber="31" value="2021-10-17">17.X.2021</date>
</collectingDate>
, leg.
<collectorName id="26995371FF8202410A258367FB20CF31" box="[1080,1167,1835,1852]" pageId="1" pageNumber="31">M. Schülke</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376652CFF8202430D41830FFD7CC9E2" lastPageId="3" lastPageNumber="33" pageId="1" pageNumber="31" type="description">
<paragraph id="8BD336A7FF8202420D41830FFB0ACFA3" blockId="1.[813,1400,1789,1955]" lastBlockId="2.[813,1400,1456,2018]" lastPageId="2" lastPageNumber="32" pageId="1" pageNumber="31">
<emphasis id="B918EAB5FF8202410D41830FFC63CF5B" bold="true" box="[860,972,1859,1878]" pageId="1" pageNumber="31">Description</emphasis>
. Length 1.87 mm, width 1.25 mm. Head, pronotum and hypomere dark brown to reddish-brown. Elytra darker than pronotum on prevailing surface, becoming lighter posterior middle, and near lateral margins becoming lighter posterior basal third, light brown on entire apical fourth. Ventral side of thorax and ventrites Ito IV dark reddish-brown. Abdominal apex yellowish. Femora about as dark as pronotum, tibiae lighter, reddish, tarsi and antennomeres Iand II yellowish, following antennomeres light brown. Length/width ratios of the antennomeresas: III 16: 8, IV 27: 7, V 34: 8, VI 35: 10, VII 48: 15, VIII 38: 10, IX 48: 15, X 46: 15, XI 55: 15. Pronotumwithevenly rounded lateral margins, lateral margin carinae concealed indorsal view, disc not microsculptured, densely and very finely punctate, punctures mostly well delimited, visible at 20 times magnification, puncture intervals mostly about 3 to 5 times as large as puncture diameters; lateral striae impunctate; pubescence distinct. Exposed tip of scutellum minute, narrow, longer than wide. Elytra each with weakly rounded lateral margin, moderately narrowed apically, lateral margin carina nearly entirely concealed, exposed only near base in dorsal view; apical margin truncate, lacking crenulation at inner angle, inner angle not prominent, about at level of outer angle in dorsal view; sutural margin not raised, adsutural areaflat, narrowed apically, with fine puncture row, and additional very fine punctures forming second row in anterior third, sutural stria shallow, curved along base to form basal stria extended about to mid-width of basal margin; disc not microsculptured, with punctation mostly dense and rather coarse, puncture intervals mostly slightly larger to about twice as large as puncture diameters, punctures becoming much fine near base and along lateral stria; lateral stria distinctly punctate. Hypomera smooth. Mesepimeron about as long as interval between its tip and mesocoxa. Metaventrite not microsculptured, slightly convex between mesocoxae, flattened on apicomedian area; finely and densely punctate on median area smooth centre excepted, lacking impressions. Punctation on area between mesocoxa and metacoxa distinctly coarser than that on apicomedian area of metaventrite, anteriolateral surface of metaventrite nearly smooth. Submesocoxal area 0.05 mm, about as fourth of shortest interval to metacoxa. Submesocoxal line convex, finely punctate. Metanepisternum strongly narrowed anteriad, broadly rounded at angles, in plan with metaventrite. Tibiae straight. Exposed abdominal tergites with very miscosculpture consisting of very short striae and punctures. Exposed ventrites with strigulate microsculpture. Ventrite Iwith punctation about as fine as that on apicomedian area of metaventrite. Submetacoxal area 0.08 mm, about as fourth of interval to apical margin; submetaxocal line convex, finely punctate.
</paragraph>
<caption id="DF13662FFF8102420EF08169FBA1CD75" pageId="2" pageNumber="32" startId="2.[237,268,1317,1334]" targetBox="[195,1389,273,1302]" targetPageId="2">
<paragraph id="8BD336A7FF8102420EF08169FBA1CD75" blockId="2.[237,1038,1317,1401]" pageId="2" pageNumber="32">
Figs 13.
<taxonomicName id="4C6C4D24FF8102420F2B8169FE5CCD38" authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[310,499,1317,1334]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="caucasicum" status="sp. nov.">
<emphasis id="B918EAB5FF8102420F2B8169FE5CCD38" box="[310,499,1317,1334]" italics="true" pageId="2" pageNumber="32">Scaphisoma caucasicum</emphasis>
</taxonomicName>
<emphasis id="B918EAB5FF8102420FE58169FD8CCD38" bold="true" box="[504,547,1317,1333]" pageId="2" pageNumber="32">
<taxonomicNameLabel id="A22B57CEFF8102420FE58169FD8CCD38" box="[504,547,1317,1333]" pageId="2" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, aedeagus. 12 aedeagus: 1 dorsal view, 2 lateral view; 3 internal sac. Scale bars 0.1 mm. Рис. 13.
<taxonomicName id="4C6C4D24FF8102420F24811EFE59CD6F" authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[313,502,1362,1379]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="caucasicum" status="sp. nov.">
<emphasis id="B918EAB5FF8102420F24811EFE59CD6F" box="[313,502,1362,1379]" italics="true" pageId="2" pageNumber="32">Scaphisoma caucasicum</emphasis>
</taxonomicName>
<emphasis id="B918EAB5FF8102420FE6811EFD89CD6F" bold="true" box="[507,550,1362,1378]" pageId="2" pageNumber="32">
<taxonomicNameLabel id="A22B57CEFF8102420FE6811EFD89CD6F" box="[507,550,1362,1378]" pageId="2" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, эдеагус. 12 эдеагус: 1 вид сверху, 2 вид сбоку; 3 внутренний мешок. Масштабные линейки 0.1 мм.
</paragraph>
</caption>
<paragraph id="8BD336A7FF8102430D4183F9FD7CC9E2" blockId="2.[813,1400,1456,2018]" lastBlockId="3.[188,776,273,1036]" lastPageId="3" lastPageNumber="33" pageId="2" pageNumber="32">
Male characters. Protarsomeres Ito III slightly widened, mesotarsomeres not widened. Aedeagus (
<figureCitation id="13572A22FF8102420AA68383FAA2CFEF" box="[1211,1293,1999,2018]" captionStart="Figs 13" captionStartId="2.[237,268,1317,1334]" captionTargetBox="[195,1389,273,1302]" captionTargetId="figure-418@2.[189,1399,272,1308]" captionTargetPageId="2" captionText="Figs 13. Scaphisoma caucasicum sp. n., aedeagus. 12 aedeagus: 1 dorsal view, 2 lateral view; 3 internal sac. Scale bars 0.1 mm. Рис. 13. Scaphisoma caucasicum sp. n., эдеагус. 12 эдеагус: 1 вид сверху, 2 вид сбоку; 3 внутренний мешок. Масштабные линейки 0.1 мм." pageId="2" pageNumber="32">Figs 13</figureCitation>
) 0.56 mm long. Median lobe symmetrical, basal bulb weakly sclerotized, apical process strongly sclerotized, as long as basal bulb, strongly inflexed, with tip bent, acute in lateral view, not visible in dorsal view. Ostium subapical, overlapped by short dorsal plate. Articular process inconspicuous. Parameres slightly sinuate, nearly evenly wide in dorsal and lateral views, without lobes. Internal sac with proximal V-shaped plate-like structure, two very narrow admesal rods, very weakly sclerotized spines at each side of apical parts of rods, and two densely, well sclerotized apical spine bunches.
</paragraph>
</subSubSection>
<subSubSection id="C376652CFF8002430EF085BAFE2ECBBA" pageId="3" pageNumber="33" type="diagnosis">
<paragraph id="8BD336A7FF8002430EF085BAFE2ECBBA" blockId="3.[188,776,273,1036]" pageId="3" pageNumber="33">
<emphasis id="B918EAB5FF8002430EF085BAFE60CA01" bold="true" box="[237,463,502,524]" pageId="3" pageNumber="33">Differential diagnosis</emphasis>
.
<taxonomicName id="4C6C4D24FF8002430FC785BBFD66CA01" authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[474,713,503,524]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="caucasicum" status="sp. nov.">
<emphasis id="B918EAB5FF8002430FC785BBFD66CA01" box="[474,713,503,524]" italics="true" pageId="3" pageNumber="33">Scaphisoma caucasicum</emphasis>
</taxonomicName>
<emphasis id="B918EAB5FF8002430CD285BBFCA8CA01" bold="true" box="[719,775,503,524]" pageId="3" pageNumber="33">
<taxonomicNameLabel id="A22B57CEFF8002430CD285BBFCA8CA01" box="[719,775,503,524]" pageId="3" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
falls in the key to the Palaearctic species of
<taxonomicName id="4C6C4D24FF8002430C92865FFCA8CA25" authorityName="Leach" authorityYear="1815" box="[655,775,531,552]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAB5FF8002430C92865FFCA8CA25" box="[655,775,531,552]" italics="true" pageId="3" pageNumber="33">Scaphisoma</emphasis>
</taxonomicName>
[
<bibRefCitation id="EFFD4B56FF8002430EDE8663FE9DCA48" author="Lobl I." box="[195,306,559,581]" journalOrPublisher="Revue suisse de zoologie." pageId="3" pageNumber="33" pagination="727 - 799" part="77" refId="ref3077" refString="Lobl I. 1970. Revision der palaarktischen Arten der Gattungen Scaphisoma Leach und Caryoscapha Ganglbauer der Tribus Scaphisomini (Col. Scaphidiidae). Revue suisse de zoologie. 77: 727 - 799." title="Revision der palaarktischen Arten der Gattungen Scaphisoma Leach und Caryoscapha Ganglbauer der Tribus Scaphisomini (Col. Scaphidiidae)" type="journal article" year="1970">Löbl, 1970</bibRefCitation>
] to
<taxonomicName id="4C6C4D24FF8002430F40867CFDF8CA48" authority="Lobl, 1968" authorityName="Lobl" authorityYear="1968" box="[349,599,559,581]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="hapiroense">
<emphasis id="B918EAB5FF8002430F40867CFE4DCA49" box="[349,482,559,581]" italics="true" pageId="3" pageNumber="33">S. hapiroense</emphasis>
Löbl, 1968
</taxonomicName>
, couplet 33. This species was described and is known only from North Korea. Though the male characters of
<taxonomicName id="4C6C4D24FF8002430C258625FD14CA70" authorityName="Lobl" authorityYear="1968" box="[568,699,616,638]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="hapiroense">
<emphasis id="B918EAB5FF8002430C258625FD14CA70" box="[568,699,616,638]" italics="true" pageId="3" pageNumber="33">S. hapiroense</emphasis>
</taxonomicName>
remain unknown, the new species may be readily distinguished from it by the antennomere Vshorter than the combined length of the antennomeres III and IV, the very fine pronotal punctation, and the light apical area of the elytra weakly delimited. The aedeagal characters of
<taxonomicName id="4C6C4D24FF8002430EA0875FFEE5CB24" authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[189,330,787,809]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="caucasicum" status="sp. nov.">
<emphasis id="B918EAB5FF8002430EA0875FFEE5CB24" box="[189,330,787,809]" italics="true" pageId="3" pageNumber="33">S. caucasicum</emphasis>
</taxonomicName>
<emphasis id="B918EAB5FF8002430F4C875FFE25CB25" bold="true" box="[337,394,787,808]" pageId="3" pageNumber="33">
<taxonomicNameLabel id="A22B57CEFF8002430F4C875FFE25CB25" box="[337,394,787,808]" pageId="3" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="4C6C4D24FF8002430C9E875FFF40CB49" authority="Lobl, 2000" authorityName="Lobl" authorityYear="2000" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="B918EAB5FF8002430C9E875FFD64CB25" box="[643,715,787,808]" italics="true" pageId="3" pageNumber="33">S. latro</emphasis>
<bibRefCitation id="EFFD4B56FF8002430CC9875FFF40CB49" author="Lobl I." journalOrPublisher="Revue suisse de zoologie." pageId="3" pageNumber="33" pagination="601 - 656" part="107" refId="ref3113" refString="Lobl I. 2000. A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, II. Revue suisse de zoologie. 107: 601 - 656." title="A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, II" type="journal article" year="2000">Löbl, 2000</bibRefCitation>
</taxonomicName>
, currently known only from the Chinese provinces of Hubei and Sichuan [
<bibRefCitation id="EFFD4B56FF8002430FB78700FDB6CB6C" author="Lobl I." box="[426,537,844,866]" journalOrPublisher="Revue suisse de zoologie." pageId="3" pageNumber="33" pagination="601 - 656" part="107" refId="ref3113" refString="Lobl I. 2000. A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, II. Revue suisse de zoologie. 107: 601 - 656." title="A review of the Scaphidiinae (Coleoptera: Staphylinidae) of the People's Republic of China, II" type="journal article" year="2000">Löbl, 2000</bibRefCitation>
,
<bibRefCitation id="EFFD4B56FF8002430C388700FDF8CB6C" author="Lobl I." bookContentInfo="xvi + 418 p." box="[549,599,844,865]" journalOrPublisher="Leiden - Boston: Brill." pageId="3" pageNumber="33" refId="ref3148" refString="Lobl I. 2018. World Catalogue of Insects. Volume 16. Coleoptera: Staphylinidae: Scaphidiinae. Leiden - Boston: Brill. xvi + 418 p." title="World Catalogue of Insects. Volume 16. Coleoptera: Staphylinidae: Scaphidiinae" type="book" year="2018">2018</bibRefCitation>
,
<bibRefCitation id="EFFD4B56FF8002430C7E8707FD37CB6C" author="Lobl I." box="[611,664,843,865]" journalOrPublisher="Annales Zoologici." pageId="3" pageNumber="33" pagination="241 - 292" part="69" refId="ref3178" refString="Lobl I. 2019. New species and records of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae) from the People's Republic of China. Annales Zoologici. 69 (2): 241 - 292. DOI: 10.3161 / 00034541 ANZ 201 9.69. 2.002" title="New species and records of Scaphisoma Leach (Coleoptera: Staphylinidae: Scaphidiinae) from the People's Republic of China" type="journal article" year="2019">2019</bibRefCitation>
]. However,
<taxonomicName id="4C6C4D24FF8002430EA08725FEADCB70" authorityName="Lobl" authorityYear="2000" box="[189,258,872,894]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="33" phylum="Arthropoda" rank="species" species="latro">
<emphasis id="B918EAB5FF8002430EA08725FEADCB70" box="[189,258,872,894]" italics="true" pageId="3" pageNumber="33">S. latro</emphasis>
</taxonomicName>
is distinguished by the much shorter apical process of the median lobe and the internal sac lacking V-shaped structure and rods.
</paragraph>
</subSubSection>
<subSubSection id="C376652CFF8002430EF087F1FD87CBD9" box="[237,552,957,980]" pageId="3" pageNumber="33" type="distribution">
<paragraph id="8BD336A7FF8002430EF087F1FD87CBD9" blockId="3.[188,776,273,1036]" box="[237,552,957,980]" pageId="3" pageNumber="33">
<emphasis id="B918EAB5FF8002430EF087F1FED8CBDE" bold="true" box="[237,375,957,979]" pageId="3" pageNumber="33">Distribution.</emphasis>
Georgia: Svaneti.
</paragraph>
</subSubSection>
<subSubSection id="C376652CFF8002430EF08796FD2BCC01" pageId="3" pageNumber="33" type="etymology">
<paragraph id="8BD336A7FF8002430EF08796FD2BCC01" blockId="3.[188,776,273,1036]" pageId="3" pageNumber="33">
<emphasis id="B918EAB5FF8002430EF08796FECCCBE2" bold="true" box="[237,355,986,1007]" pageId="3" pageNumber="33">Etymology.</emphasis>
The species epithet is an adjective derive from the Caucasus, where the species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>