140 lines
21 KiB
XML
140 lines
21 KiB
XML
<document id="CBC52E35BA183F7CD9A154BF91AE68B2" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629613271721" checkinUser="felipe" docAuthor="Paquin, Pierre & Dupérré, Nadine" docDate="2009" docId="03C55737FFF6FFDEFF50FA52D540FD68" docLanguage="en" docName="zt02002p067.pdf" docOrigin="Zootaxa 2002" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cicurina (Cicurella) neovespera Cokendolpher 2004" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="FFFC2F4FFFD6FFFFFFC7FFA4D12FFFC0" masterDocTitle="of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution" masterLastPageNumber="67" masterPageNumber="1" pageNumber="33" updateTime="1699083252512" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2526D226A2BE8BE4BBF8792E1A1B9077">of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution</mods:title>
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<mods:name id="082981B335D2A3C6099E424584D15EB5" type="personal">
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<mods:roleTerm id="148DCB6926FB2C7BE2C5C3BA54694092">Author</mods:roleTerm>
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<mods:namePart id="6B97D438FD308B2B16ACD64F224564CE">Paquin, Pierre</mods:namePart>
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<mods:affiliation id="7E9B36ADC4FC4083335B73206C0A31F2">Cave and Endangered Invertebrate Research Laboratory, SWCA Environmental Consultants, 4407 Monterey Oaks Boulevard, Building 1, Suite 110, Austin, Texas, 78749, USA. E-mail: ppaquin @ swca. com & This is publication no. 15 of the Karst Biosciences and Environmental Geophysics Research Laboratories, SWCA Environmental Consultants</mods:affiliation>
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<mods:name id="AFD9573C5C7D0FE5ADCACCF52279737A" type="personal">
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<mods:namePart id="AC5EBC5280F4B6C30C888195C1EC3C2C">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="85465D4594E7F00FCC94F46919D311D5">American Museum of Natural History, Division of Invertebrate Zoology, Central Park West at 79 Street, New York, New York, 10024 USA. E-mail: nduperre @ amnh. org</mods:affiliation>
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<mods:typeOfResource id="ACB6394CBA3DAB6C8B60285D23603E04">text</mods:typeOfResource>
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<mods:title id="5ADC00A6A2AF1B9BDA72DC45B4B8B286">Zootaxa</mods:title>
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<mods:part id="F726A1B493C7B0862D27FC8E45E094A1">
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<mods:date id="A6EA26F581AEB51B2B203AFC8BAE50B4">2009</mods:date>
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<mods:number id="749CFF256AC6AFB6B5F696BBE5547C58">2009-02-04</mods:number>
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<mods:detail id="109F607023510CCC4EF82CD7590D794D" type="volume">
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<mods:number id="BA5EE03569BAB2F553B4A77DC5C515C8">2002</mods:number>
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<mods:end id="970C51A0BA9CB62B119C0219F14F40E5">67</mods:end>
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<mods:classification id="CD7DA6798413C65ADEEE6DFBB1146F80">journal article</mods:classification>
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<mods:identifier id="DEEA7B7328B2941A3B31FE63E4210B63" type="ISSN">11755334</mods:identifier>
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<treatment id="03C55737FFF6FFDEFF50FA52D540FD68" ID-DOI="http://doi.org/10.5281/zenodo.5243049" ID-Zenodo-Dep="5243049" LSID="urn:lsid:plazi:treatment:03C55737FFF6FFDEFF50FA52D540FD68" httpUri="http://treatment.plazi.org/id/03C55737FFF6FFDEFF50FA52D540FD68" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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<subSubSection id="C376B5AAFFF6FFDFFF50FA52D237F9D0" box="[151,792,1526,1552]" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="8BD3E621FFF6FFDFFF50FA52D237F9D0" blockId="32.[151,792,1526,1587]" box="[151,792,1526,1552]" pageId="32" pageNumber="33">
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<heading id="D09B514DFFF6FFDFFF50FA52D237F9D0" bold="true" box="[151,792,1526,1552]" fontSize="11" level="1" pageId="32" pageNumber="33" reason="1">
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<taxonomicName id="4C6C9DA2FFF6FFDFFF50FA52D237F9D0" ID-CoL="VB24" authority="Cokendolpher 2004" authorityName="Cokendolpher" authorityYear="2004" box="[151,792,1526,1552]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="neovespera" subGenus="Cicurella">
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<emphasis id="B9183A33FFF6FFDFFF50FA52D237F9D0" bold="true" box="[151,792,1526,1552]" pageId="32" pageNumber="33">
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<emphasis id="B9183A33FFF6FFDFFF50FA52D333F9D0" bold="true" box="[151,540,1526,1552]" italics="true" pageId="32" pageNumber="33">Cicurina (Cicurella) neovespera</emphasis>
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Cokendolpher 2004
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFF6FFDFFF50F9BDD048F9F3" box="[151,359,1561,1587]" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BD3E621FFF6FFDFFF50F9BDD048F9F3" blockId="32.[151,792,1526,1587]" box="[151,359,1561,1587]" pageId="32" pageNumber="33">
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(
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<figureCitation id="1357FAA4FFF6FFDFFF58F9BDD009F9F3" box="[159,294,1561,1587]" captionStart="FIGURES 48–67" captionStartId="30.[151,269,1797,1821]" captionTargetBox="[272,1310,209,1785]" captionTargetId="figure-135@30.[272,1315,194,1788]" captionTargetPageId="30" captionText="FIGURES 48–67. Genitalic structure of troglobitic Cicurina (Cicurella) holotypes. For each female holotype, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. For the male holotype, a ventral and a lateral view of the left palp are presented, respectively. 48–49 Cicurina machete Gertsch 1992. 50–51 Cicurina madla Gertsch 1992. 52–53 Cicurina maya Gertsch 1977. 54–55" pageId="32" pageNumber="33">Figs 64–65</figureCitation>
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,
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<figureCitation id="1357FAA4FFF6FFDFFEF5F9BDD070F9F3" box="[306,351,1561,1587]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin & Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="32" pageNumber="33">134</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFF6FFDFFF00F9C3D3EFF967" pageId="32" pageNumber="33" type="reference_group">
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<paragraph id="8BD3E621FFF6FFDFFF00F9C3D3EFF967" blockId="32.[151,1435,1638,1703]" pageId="32" pageNumber="33">
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<treatmentCitationGroup id="AB7CC10FFFF6FFDFFF00F9C3D3EFF967" pageId="32" pageNumber="33">
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<treatmentCitation id="0ACDC030FFF6FFDFFF00F9C3D235F940" author="Cokendolpher, J. C." box="[199,794,1638,1664]" page="20" pageId="32" pageNumber="33" year="2004">
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<taxonomicName id="4C6C9DA2FFF6FFDFFF00F9C3D235F940" ID-CoL="VB24" authority="Cokendolpher 2004 a: 19 - 20" authorityName="Cokendolpher" authorityPageNumber="19 - 20" authorityYear="2004" box="[199,794,1638,1664]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="neovespera">
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<emphasis id="B9183A33FFF6FFDFFF00F9C3D091F940" box="[199,446,1639,1664]" italics="true" pageId="32" pageNumber="33">Cicurina neovespera</emphasis>
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<bibRefCitation id="EFFD9BD0FFF6FFDFFE00F9C2D235F940" author="Cokendolpher, J. C." box="[455,794,1638,1664]" pageId="32" pageNumber="33" pagination="13 - 58" refId="ref39625" refString="Cokendolpher, J. C. (2004 a) Cicurina spiders from caves in Bexar County, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, 4, 13 - 58." type="journal article" year="2004">Cokendolpher 2004a: 19–20</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, 22, 24, 27, 34, 41,
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<emphasis id="B9183A33FFF6FFDFFBCAF9C2D516F940" bold="true" box="[1037,1081,1638,1664]" pageId="32" pageNumber="33">47–</emphasis>
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49, figs 52–53 (description of female).
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<emphasis id="B9183A33FFF6FFDFFF38F929D033F967" bold="true" box="[255,284,1677,1703]" pageId="32" pageNumber="33">—</emphasis>
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<treatmentCitation id="0ACDC030FFF6FFDFFEE3F929D393F967" author="Reddell, J. R. & Cokendolpher, J. C." box="[292,700,1677,1703]" page="80" pageId="32" pageNumber="33" year="2004">
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<bibRefCitation id="EFFD9BD0FFF6FFDFFEE3F929D393F967" author="Reddell, J. R. & Cokendolpher, J. C." box="[292,700,1677,1703]" pageId="32" pageNumber="33" pagination="75 - 94" refId="ref41088" refString="Reddell, J. R. & Cokendolpher, J. C. (2004) The cave spiders (Araneae) of Bexar and Comal counties, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, IV, 75 - 94." type="journal article" year="2004">Reddell & Cokendolpher 2004: 80</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFF6FFDFFF50F97ED447F934" box="[151,1384,1754,1780]" pageId="32" pageNumber="33" type="materials_examined">
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<paragraph id="8BD3E621FFF6FFDFFF50F97ED447F934" blockId="32.[151,1436,1754,1935]" box="[151,1384,1754,1780]" pageId="32" pageNumber="33">
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<materialsCitation id="3B04EC7CFFF6FFDFFF50F97ED447F934" box="[151,1384,1754,1780]" collectionCode="AMNH" collectorName="A. G. Grubbs" county="Bexar Co." location="Elm Springs Cave" pageId="32" pageNumber="33" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
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<emphasis id="B9183A33FFF6FFDFFF50F97ED08EF934" bold="true" box="[151,417,1754,1780]" pageId="32" pageNumber="33">
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||
<typeStatus id="54D75883FFF6FFDFFF50F97ED012F934" box="[151,317,1754,1780]" pageId="32" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
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female:
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</emphasis>
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“
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<collectingRegion id="49A828C3FFF6FFDFFE74F97ED0D1F934" box="[435,510,1754,1780]" country="United States of America" name="Texas" pageId="32" pageNumber="33">Texas</collectingRegion>
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:
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<collectingCounty id="62B29EADFFF6FFDFFDCFF97ED3ACF934" box="[520,643,1754,1780]" pageId="32" pageNumber="33">Bexar Co.</collectingCounty>
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:
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<location id="8EB3B0FAFFF6FFDFFD48F97ED248F934" LSID="urn:lsid:plazi:treatment:03C55737FFF6FFDEFF50FA52D540FD68:8EB3B0FAFFF6FFDFFD48F97ED248F934" box="[655,871,1754,1780]" county="Bexar Co." name="Elm Springs Cave" pageId="32" pageNumber="33" stateProvince="Texas">Elm Springs Cave</location>
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,
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<collectorName id="269983F7FFF6FFDFFCB4F97ED526F934" box="[883,1033,1754,1780]" pageId="32" pageNumber="33">A.G. Grubbs</collectorName>
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, date?”. Examined (
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<collectionCode id="ED7D7EE4FFF6FFDFFB3BF97ED473F934" box="[1276,1372,1754,1780]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="32" pageNumber="33">AMNH</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFF6FFDEFF00F8A4D5FAFEB3" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
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<paragraph id="8BD3E621FFF6FFDFFF00F8A4D305F8DA" blockId="32.[151,1436,1754,1935]" box="[199,554,1792,1818]" pageId="32" pageNumber="33">
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||
<emphasis id="B9183A33FFF6FFDFFF00F8A4D070F8DA" bold="true" box="[199,351,1792,1818]" pageId="32" pageNumber="33">Description.</emphasis>
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<emphasis id="B9183A33FFF6FFDFFEA1F8A5D305F8DA" box="[358,554,1793,1818]" italics="true" pageId="32" pageNumber="33">
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Female
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<typeStatus id="54D75883FFF6FFDFFE02F8A5D305F8DA" box="[453,554,1793,1818]" pageId="32" pageNumber="33" type="holotype">holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8BD3E621FFF6FFDEFF00F88CD5FAFEB3" blockId="32.[151,1436,1754,1935]" lastBlockId="33.[151,1436,152,680]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">Total length: 4.23; carapace length: 1.98; carapace width: 1.35; opisthosoma length: 2.25. Eyes absent. Cheliceral promargin right side with 3 teeth (2 large, 1 small), left side likewise; retromargin right side with 4 denticles (3 large, 1 small), left side with 6 denticles (2 large, 4 small). Leg I: total length: 5.65; femur: 1.60; with long transverse slit, bearing medial, shallow indentation; atrium rectangular, almost intruded by slightly rounded dorsal plate, atrial aperture small; bursa enlarged, not fused; copulatory ducts touching at midpoint, reaching slightly over primary lobe apex; copulatory ducts constriction located medially; primary pores visible (1 seen); stalk short, recurved, joining the primary lobe at the aperture of the dictynoid pore; dictynoid pore large, rounded; primary lobe ear-shaped; secondary lobe absent; fertilization canal short, externally positioned from dictynoid pore; fertilization duct medially attached to the primary lobe base.</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFF7FFDEFF00FEDBD3B5FE00" pageId="33" pageNumber="34" type="distribution">
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<paragraph id="8BD3E621FFF7FFDEFF00FEDBD3B5FE00" blockId="33.[151,1436,152,680]" pageId="33" pageNumber="34">
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<emphasis id="B9183A33FFF7FFDEFF00FEDBD04FFE59" bold="true" box="[199,352,383,409]" pageId="33" pageNumber="34">Distribution</emphasis>
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. Known from two caves in Bexar County (
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<figureCitation id="1357FAA4FFF7FFDEFC9AFE24D2EBFE5A" box="[861,964,384,410]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin & Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="33" pageNumber="34">Fig. 134</figureCitation>
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): Elm Spring Cave (=Grubbs Cave ES), and Grubbs Cave No. 23, 8 km NE Helotes.
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</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFF7FFDEFF00FE69D540FD68" pageId="33" pageNumber="34" type="discussion">
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||
<paragraph id="8BD3E621FFF7FFDEFF00FE69D540FD68" blockId="33.[151,1436,152,680]" pageId="33" pageNumber="34">
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<emphasis id="B9183A33FFF7FFDEFF00FE69D06FFE27" bold="true" box="[199,320,461,487]" pageId="33" pageNumber="34">Remarks.</emphasis>
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Male unknown. The exact location of cave no 23 could not be verified and is only approximate (
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<figureCitation id="1357FAA4FFF7FFDEFF58FE50D02AFDCE" box="[159,261,500,526]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin & Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="33" pageNumber="34">Fig. 134</figureCitation>
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), while Elm Spring Cave is now partially sealed with concrete for safety, and is not accessible. This makes the reassessment of the validity of that species by recollection impossible in the actual conditions. In addition, the specimen from Grubbs Cave No. 23, 8 km NE Helotes could not be located and examined to confirm the occurrence of
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<taxonomicName id="4C6C9DA2FFF7FFDEFE14FDCCD27DFD41" authorityName="Cokendolpher" authorityYear="2004" box="[467,850,616,641]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="neovespera" subGenus="Cicurella">
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<emphasis id="B9183A33FFF7FFDEFE14FDCCD27DFD41" box="[467,850,616,641]" italics="true" pageId="33" pageNumber="34">Cicurina (Cicurella) neovespera</emphasis>
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</taxonomicName>
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in that cave. However, the locality is included in
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<figureCitation id="1357FAA4FFF7FFDEFF50FD2AD1D7FD68" box="[151,248,654,680]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin & Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="33" pageNumber="34">Fig. 134</figureCitation>
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based on
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<bibRefCitation id="EFFD9BD0FFF7FFDEFEA9FD2AD3A4FD68" author="Cokendolpher, J. C." box="[366,651,654,680]" pageId="33" pageNumber="34" pagination="13 - 58" refId="ref39625" refString="Cokendolpher, J. C. (2004 a) Cicurina spiders from caves in Bexar County, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, 4, 13 - 58." type="journal article" year="2004">Cokendolpher’s (2004a)</bibRefCitation>
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record and indicated by an open symbol.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |