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<document id="28E32DE4E478B95C479EDCA2D8E7C71C" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629613271721" checkinUser="felipe" docAuthor="Paquin, Pierre &amp; Dupérré, Nadine" docDate="2009" docId="03C55737FFD9FFEFFF50F91AD32FFD68" docLanguage="en" docName="zt02002p067.pdf" docOrigin="Zootaxa 2002" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cicurina (Cicurella) brunsi Cokendolpher 2004" docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="FFFC2F4FFFD6FFFFFFC7FFA4D12FFFC0" masterDocTitle="of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution" masterLastPageNumber="67" masterPageNumber="1" pageNumber="16" updateTime="1699083252512" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="661723B923C5FD9A6D3582A9E079D0DD">of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution</mods:title>
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<mods:namePart id="2D653C8BDDD5469844AD5DCF646E3B35">Paquin, Pierre</mods:namePart>
<mods:affiliation id="611CFA52A5E3433D6EA45FBF0AF7E349">Cave and Endangered Invertebrate Research Laboratory, SWCA Environmental Consultants, 4407 Monterey Oaks Boulevard, Building 1, Suite 110, Austin, Texas, 78749, USA. E-mail: ppaquin @ swca. com &amp; This is publication no. 15 of the Karst Biosciences and Environmental Geophysics Research Laboratories, SWCA Environmental Consultants</mods:affiliation>
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<mods:namePart id="C9EBC722A5EF16BB4BBC8124756D4EC3">Dupérré, Nadine</mods:namePart>
<mods:affiliation id="BA1FD6E65C7C37C9869A230AB5430B43">American Museum of Natural History, Division of Invertebrate Zoology, Central Park West at 79 Street, New York, New York, 10024 USA. E-mail: nduperre @ amnh. org</mods:affiliation>
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<mods:date id="582DB2E004B90AE28B54739733815B89">2009</mods:date>
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<treatment id="03C55737FFD9FFEFFF50F91AD32FFD68" ID-DOI="http://doi.org/10.5281/zenodo.5243010" ID-Zenodo-Dep="5243010" LSID="urn:lsid:plazi:treatment:03C55737FFD9FFEFFF50F91AD32FFD68" httpUri="http://treatment.plazi.org/id/03C55737FFD9FFEFFF50F91AD32FFD68" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<subSubSection id="C376B5AAFFD9FFF0FF50F91AD3F0F918" box="[151,735,1726,1752]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BD3E621FFD9FFF0FF50F91AD3F0F918" blockId="15.[151,735,1726,1787]" box="[151,735,1726,1752]" pageId="15" pageNumber="16">
<heading id="D09B514DFFD9FFF0FF50F91AD3F0F918" bold="true" box="[151,735,1726,1752]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C6C9DA2FFD9FFF0FF50F91AD3F0F918" ID-CoL="5YTTR" authority="Cokendolpher 2004" authorityName="Cokendolpher" authorityYear="2004" box="[151,735,1726,1752]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="brunsi" subGenus="Cicurella">
<emphasis id="B9183A33FFD9FFF0FF50F91AD3F0F918" bold="true" box="[151,735,1726,1752]" pageId="15" pageNumber="16">
<emphasis id="B9183A33FFD9FFF0FF50F91AD0CCF918" bold="true" box="[151,483,1726,1752]" italics="true" pageId="15" pageNumber="16">Cicurina (Cicurella) brunsi</emphasis>
Cokendolpher 2004
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFD9FFF0FF50F945D048F93B" box="[151,359,1761,1787]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BD3E621FFD9FFF0FF50F945D048F93B" blockId="15.[151,735,1726,1787]" box="[151,359,1761,1787]" pageId="15" pageNumber="16">
(
<figureCitation id="1357FAA4FFD9FFF0FF58F945D00AF93B" box="[159,293,1761,1787]" captionStart="FIGURES 827" captionStartId="19.[151,272,1804,1828]" captionTargetBox="[275,1316,207,1785]" captionTargetId="figure-119@19.[272,1329,197,1798]" captionTargetPageId="19" captionText="FIGURES 827. Cleared epigynum of troglobitic Cicurina (Cicurella) holotypes. For each species, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. 89 Cicurina bandera Gertsch 1992. 1011 Cicurina bandida Gertsch 1992. 1213 Cicurina baronia Gertsch 1992. 1415 Cicurina barri Gertsch 1992. 1617 Cicurina browni Gertsch 1992. 1819 Cicurina brunsi" pageId="15" pageNumber="16">Figs 1819</figureCitation>
,
<figureCitation id="1357FAA4FFD9FFF0FEF5F945D070F93B" box="[306,351,1761,1787]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin &amp; Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="15" pageNumber="16">134</figureCitation>
)
</paragraph>
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<subSubSection id="C376B5AAFFD9FFF0FF50F88BD594F8A5" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8BD3E621FFD9FFF0FF50F88BD594F8A5" blockId="15.[151,1436,1838,1893]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB7CC10FFFD9FFF0FF50F88BD594F8A5" pageId="15" pageNumber="16">
<treatmentCitation id="0ACDC030FFD9FFF0FF50F88BD3AAF885" author="Cokendolpher, J. C." box="[151,645,1838,1862]" page="20" pageId="15" pageNumber="16" year="2004">
<taxonomicName id="4C6C9DA2FFD9FFF0FF50F88BD3AAF885" ID-CoL="5YTTR" authority="Cokendolpher 2004 a: 19 - 20" authorityName="Cokendolpher" authorityPageNumber="19 - 20" authorityYear="2004" box="[151,645,1838,1862]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="brunsi">
<emphasis id="B9183A33FFD9FFF0FF50F88BD06CF886" box="[151,323,1839,1862]" italics="true" pageId="15" pageNumber="16">Cicurina brunsi</emphasis>
<bibRefCitation id="EFFD9BD0FFD9FFF0FE8AF88AD3AAF885" author="Cokendolpher, J. C." box="[333,645,1838,1861]" pageId="15" pageNumber="16" pagination="13 - 58" refId="ref39625" refString="Cokendolpher, J. C. (2004 a) Cicurina spiders from caves in Bexar County, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, 4, 13 - 58." type="journal article" year="2004">Cokendolpher 2004a: 1920</bibRefCitation>
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, 22, 24, 27, 34,
<emphasis id="B9183A33FFD9FFF0FCF8F88AD275F886" bold="true" box="[831,858,1838,1862]" pageId="15" pageNumber="16">38</emphasis>
40, figs 3233 (description of female).
<emphasis id="B9183A33FFD9FFF0FADDF88AD41AF886" bold="true" box="[1306,1333,1838,1862]" pageId="15" pageNumber="16"></emphasis>
<treatmentCitation id="0ACDC030FFD9FFF0FAF8F88AD01AF8A5" author="Jackman, J. A." page="162" pageId="15" pageNumber="16" year="1997">
<bibRefCitation id="EFFD9BD0FFD9FFF0FAF8F88AD01AF8A5" author="Jackman, J. A." pageId="15" pageNumber="16" refId="ref40113" refString="Jackman, J. A. (1997) A Field Guide to Spiders and Scorpions of Texas. Houston, Texas: Gulf Publishing Company. xiv + 201 pp. + 32 plates." type="book" year="1997">Jackman 1997: 162</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACDC030FFD9FFF0FE87F8EAD39CF8A5" author="Reddell, J. R. &amp; Cokendolpher, J. C." box="[320,691,1870,1893]" page="79" pageId="15" pageNumber="16" year="2004">
<bibRefCitation id="EFFD9BD0FFD9FFF0FE87F8EAD39CF8A5" author="Reddell, J. R. &amp; Cokendolpher, J. C." box="[320,691,1870,1893]" pageId="15" pageNumber="16" pagination="75 - 94" refId="ref41088" refString="Reddell, J. R. &amp; Cokendolpher, J. C. (2004) The cave spiders (Araneae) of Bexar and Comal counties, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, IV, 75 - 94." type="journal article" year="2004">Reddell &amp; Cokendolpher 2004: 79</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACDC030FFD9FFF0FD78F8EAD51FF8A5" author="Paquin, P. &amp; Hedin, M." box="[703,1072,1870,1893]" page="3243" pageId="15" pageNumber="16" year="2004">
<bibRefCitation id="EFFD9BD0FFD9FFF0FD78F8EAD51FF8A5" author="Paquin, P. &amp; Hedin, M." box="[703,1072,1870,1893]" pageId="15" pageNumber="16" pagination="1 - 40" refId="ref40436" refString="Paquin, P., &amp; Hedin, M. (2004) The power and perils of ' molecular taxonomy': a case study of eyeless and endangered relatives. Report to Fish and Wildlife Services (Austin, Texas, U. S. A). 1 - 40 p." type="book chapter" year="2004">Paquin &amp; Hedin 2004: 32433245</bibRefCitation>
</treatmentCitation>
, 3249, 3253.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFD9FFF0FF50F831D260F816" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BD3E621FFD9FFF0FF50F831D260F816" blockId="15.[151,1435,1941,2006]" pageId="15" pageNumber="16">
<materialsCitation id="3B04EC7CFFD9FFF0FF50F831D260F816" collectingDate="2001-11-01" collectionCode="AMNH" collectorName="J. R. Reddell &amp; M. Reyes" county="Bexar Co." location="Stahl Cave" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="B9183A33FFD9FFF0FF50F831D08EF86F" bold="true" box="[151,417,1941,1967]" pageId="15" pageNumber="16">
<typeStatus id="54D75883FFD9FFF0FF50F831D012F86F" box="[151,317,1941,1967]" pageId="15" pageNumber="16" type="holotype">HOLOTYPE</typeStatus>
female:
</emphasis>
<collectingRegion id="49A828C3FFD9FFF0FE74F831D0D1F86F" box="[435,510,1941,1967]" country="United States of America" name="Texas" pageId="15" pageNumber="16">Texas</collectingRegion>
:
<collectingCounty id="62B29EADFFD9FFF0FDCDF831D3ACF86F" box="[522,643,1941,1967]" pageId="15" pageNumber="16">Bexar Co.</collectingCounty>
, Camp Bullis,
<location id="8EB3B0FAFFD9FFF0FCF4F831D29AF86F" LSID="urn:lsid:plazi:treatment:03C55737FFD9FFEFFF50F91AD32FFD68:8EB3B0FAFFD9FFF0FCF4F831D29AF86F" box="[819,949,1941,1967]" county="Bexar Co." name="Stahl Cave" pageId="15" pageNumber="16" stateProvince="Texas">Stahl Cave</location>
,
<date id="FFD2C0E1FFD9FFF0FC06F831D57BF86F" box="[961,1108,1941,1967]" pageId="15" pageNumber="16" value="2001-11-01">
<collectingDate id="EF963909FFD9FFF0FC06F831D57BF86F" box="[961,1108,1941,1967]" pageId="15" pageNumber="16" value="2001-11-01">1 Nov. 2001</collectingDate>
</date>
,
<collectorName id="269983F7FFD9FFF0FBA6F831D5DEF86F" box="[1121,1265,1941,1967]" pageId="15" pageNumber="16">J.R. Reddell</collectorName>
and
<collectorName id="269983F7FFD9FFF0FAECF831D4B4F86F" box="[1323,1435,1941,1967]" pageId="15" pageNumber="16">M. Reyes</collectorName>
(molted
<collectingDate id="EF963909FFD9FFF0FF3EF818D312F816" box="[249,573,1980,2006]" isEnumeration="true" pageId="15" pageNumber="16">
<date id="FFD2C0E1FFD9FFF0FF3EF818D0B6F816" box="[249,409,1980,2006]" pageId="15" pageNumber="16" value="2001-12-14">14 Dec. 2001</date>
,
<date id="FFD2C0E1FFD9FFF0FE62F818D312F816" box="[421,573,1980,2006]" pageId="15" pageNumber="16" value="2002-08-02">2 Aug. 2002</date>
</collectingDate>
)”. Examined (
<collectionCode id="ED7D7EE4FFD9FFF0FD25F818D26DF816" box="[738,834,1980,2006]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="15" pageNumber="16">AMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFC6FFEFFF00FF33D4BBFD9B" pageId="16" pageNumber="17" type="description">
<paragraph id="8BD3E621FFC6FFEFFF00FF33D305FF71" blockId="16.[151,1437,151,680]" box="[199,554,151,177]" pageId="16" pageNumber="17">
<emphasis id="B9183A33FFC6FFEFFF00FF33D070FF71" bold="true" box="[199,351,151,177]" pageId="16" pageNumber="17">Description.</emphasis>
<emphasis id="B9183A33FFC6FFEFFEA1FF3CD305FF71" box="[358,554,152,177]" italics="true" pageId="16" pageNumber="17">
Female
<typeStatus id="54D75883FFC6FFEFFE02FF3CD305FF71" box="[453,554,152,177]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BD3E621FFC6FFEFFF00FF1AD4BBFD9B" blockId="16.[151,1437,151,680]" pageId="16" pageNumber="17">
Total length: 3.51; carapace length: 1.53; carapace width: 1.08; opisthosoma length: 1.98. Eyes absent. Cheliceral promargin right side with 3 teeth (1 large, 2 small), left side likewise; retromargin right side with 7 denticles (3 large, 4 small), left side with 6 denticles (3 large, 3 small). Leg I: total length: 4.30; femur: 1.25; patella: 0.45; tibia: 1.05; metatarsus: 0.95; tarsus: 0.60; leg IV: total length: 4.75; femur: 1.15; patella: 0.50; tibia: 1.10; metatarsus: 1.25; tarsus: 0.75. Tarsal claw IV: 0.15. Epigynum:
<quantity id="4C944BC4FFC6FFEFFBD6FEFDD5A3FEB3" box="[1041,1164,345,371]" metricMagnitude="-4" metricUnit="m" metricValue="2.94" pageId="16" pageNumber="17" unit="mm" value="0.294">0.294 mm</quantity>
. Epigynal ventral plate with long transverse slit, bearing medial, shallow, inverted V-shaped indentation; atrium rectangular, half intruded by dorsal plate, atrial aperture moderate; bursa enlarged, not fused; copulatory ducts scarcely separated at midpoint, reaching top of primary lobe apex; copulatory ducts constriction located basally; primary pores not visible; stalk, short, straight, horizontal, joining the primary lobe at the aperture of the dictynoid pore; dictynoid pore rounded; primary lobe ear-shaped; secondary lobe absent; fertilization canal long, externally positioned from dictynoid pore; fertilization duct medially attached to the primary lobe base.
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFC6FFEFFF00FDC3D403FD42" box="[199,1324,615,642]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BD3E621FFC6FFEFFF00FDC3D403FD42" blockId="16.[151,1437,151,680]" box="[199,1324,615,642]" pageId="16" pageNumber="17">
<emphasis id="B9183A33FFC6FFEFFF00FDC3D04FFD41" bold="true" box="[199,352,615,641]" pageId="16" pageNumber="17">Distribution</emphasis>
. Only reported from the
<typeStatus id="54D75883FFC6FFEFFD42FDCCD398FD42" box="[645,695,616,642]" pageId="16" pageNumber="17">type</typeStatus>
locality: Stahl Cave (Bexar County,
<collectingRegion id="49A828C3FFC6FFEFFBA1FDCCD582FD42" box="[1126,1197,616,642]" country="United States of America" name="Texas" pageId="16" pageNumber="17">Texas</collectingRegion>
,
<figureCitation id="1357FAA4FFC6FFEFFB7EFDCCD431FD42" box="[1209,1310,616,642]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin &amp; Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="16" pageNumber="17">Fig. 134</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C376B5AAFFC6FFEFFF00FD2AD32FFD68" box="[199,512,654,680]" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BD3E621FFC6FFEFFF00FD2AD32FFD68" blockId="16.[151,1437,151,680]" box="[199,512,654,680]" pageId="16" pageNumber="17">
<emphasis id="B9183A33FFC6FFEFFF00FD2AD06FFD68" bold="true" box="[199,320,654,680]" pageId="16" pageNumber="17">Remarks.</emphasis>
Male unknown.
</paragraph>
</subSubSection>
</treatment>
</document>