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<mods:title id="A0835BAAB7D6E8579208E4C9C7DFE412">Diversity and morphology of Eocene and Oligocene Mordellidae (Coleoptera)</mods:title>
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<subSubSection id="C3299CC5F26A8C60FCC3FB65C799FA84" pageId="3" pageNumber="454" type="nomenclature">
<paragraph id="8B8CCF4EF26A8C60FCC3FB65C799FA84" blockId="3.[836,1428,1237,1293]" pageId="3" pageNumber="454">
<heading id="D0C47822F26A8C60FCC3FB65C799FA84" centered="true" fontSize="10" level="2" pageId="3" pageNumber="454" reason="3">
<emphasis id="B947135CF26A8C60FCC3FB65C799FA84" bold="true" pageId="3" pageNumber="454">
<taxonomicName id="4C33B4CDF26A8C60FCC3FB65C6A4FB64" authority="(Perkovsky &amp; Odnosum, 2013)" authorityName="Batelka &amp; Rosová &amp; Prokop" authorityYear="2023" baseAuthorityName="Perkovsky &amp; Odnosum" baseAuthorityYear="2013" box="[836,1428,1237,1261]" class="Insecta" family="Mordellidae" genus="Baltimorda" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="friedrichi" status="comb. nov.">
<emphasis id="B947135CF26A8C60FCC3FB65C71FFB64" bold="true" box="[836,1071,1237,1261]" italics="true" pageId="3" pageNumber="454">Baltimorda friedrichi</emphasis>
(
<bibRefCitation id="EFA2B2BFF26A8C60FBBBFB66C6BDFB64" author="PERKOVSKY E. E. &amp; ODNOSUM V. K." box="[1084,1421,1238,1261]" pageId="3" pageNumber="454" pagination="177 - 179" refId="ref21948" refString="PERKOVSKY E. E. &amp; ODNOSUM V. K. 2013: A new species of tumbling flower beetles of the genus Mordellaria (Insecta: Coleoptera: Mordellidae) from the Baltic amber. Paleontological Journal 47: 177 - 179." type="journal article" year="2013">Perkovsky &amp; Odnosum, 2013</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A274AE27F26A8C60FBA8FB46C799FA84" box="[1071,1193,1270,1293]" pageId="3" pageNumber="454" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3299CC5F26A8C60FCBBFAACC0FDFAC0" pageId="3" pageNumber="454" type="reference_group">
<paragraph id="8B8CCF4EF26A8C60FCBBFAACC0FDFAC0" blockId="3.[828,1438,1308,1353]" pageId="3" pageNumber="454">
<taxonomicName id="4C33B4CDF26A8C60FCBBFAACC60CFAA6" authority="Perkovsky &amp; Odnosum. 2013: 177" authorityName="Perkovsky &amp; Odnosum." authorityPageNumber="177" authorityYear="2013" box="[828,1340,1308,1327]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="friedrichi">
<emphasis id="B947135CF26A8C60FCBBFAACC0CDFAA6" box="[828,1021,1308,1327]" italics="true" pageId="3" pageNumber="454">Mordellaria friedrichi</emphasis>
Perkovsky &amp; Odnosum. 2013: 177
</taxonomicName>
.
<typeStatus id="548871ECF26A8C60FAC0FAACC6AAFAA6" box="[1351,1434,1308,1327]" pageId="3" pageNumber="454" type="holotype">Holotype</typeStatus>
: Baltic amber.
</paragraph>
</subSubSection>
<subSubSection id="C3299CC5F26A8C67FCBBFAD2C1C5FC5C" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454" type="discussion">
<paragraph id="8B8CCF4EF26A8C67FCBBFAD2C299FCFC" blockId="3.[828,1437,1378,2041]" lastBlockId="4.[151,760,190,981]" lastPageId="4" lastPageNumber="455" pageId="3" pageNumber="454">
<emphasis id="B947135CF26A8C60FCBBFAD2C7B6FAF0" bold="true" box="[828,1158,1378,1401]" pageId="3" pageNumber="454">Comments on identification.</emphasis>
A casual examination of pictures of the habitus of
<taxonomicName id="4C33B4CDF26A8C60FBCDFA32C607FA10" box="[1098,1335,1410,1433]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="friedrichi">
<emphasis id="B947135CF26A8C60FBCDFA32C607FA10" box="[1098,1335,1410,1433]" italics="true" pageId="3" pageNumber="454">Mordellaria friedrichi</emphasis>
</taxonomicName>
indicates marked differences from those of the West Palaearctic
<taxonomicName id="4C33B4CDF26A8C60FAF8FA12C75CFA50" authority="Comolli, 1837" authorityName="Comolli" authorityYear="1837" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="aurofasciata">
<emphasis id="B947135CF26A8C60FAF8FA12C6A5FA30" box="[1407,1429,1442,1465]" italics="true" pageId="3" pageNumber="454">M</emphasis>
.
<emphasis id="B947135CF26A8C60FCBBFA72C0F2FA50" box="[828,962,1474,1497]" italics="true" pageId="3" pageNumber="454">aurofasciata</emphasis>
Comolli, 1837
</taxonomicName>
. The explanation for this is an incorrect generic diagnosis of
<taxonomicName id="4C33B4CDF26A8C60FB10FA52C628FA70" authorityName="Ermisch" authorityYear="1950" box="[1175,1304,1506,1529]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF26A8C60FB10FA52C628FA70" box="[1175,1304,1506,1529]" italics="true" pageId="3" pageNumber="454">Mordellaria</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFA2B2BFF26A8C60FCBBF9B2C7A3F990" author="PERKOVSKY E. E. &amp; ODNOSUM V. K." box="[828,1171,1538,1561]" pageId="3" pageNumber="454" pagination="177 - 179" refId="ref21948" refString="PERKOVSKY E. E. &amp; ODNOSUM V. K. 2013: A new species of tumbling flower beetles of the genus Mordellaria (Insecta: Coleoptera: Mordellidae) from the Baltic amber. Paleontological Journal 47: 177 - 179." type="journal article" year="2013">PERKOVSKY &amp; ODNOSUM (2013)</bibRefCitation>
in which they state that: ‘
<taxonomicName id="4C33B4CDF26A8C60FCC2F992C0F9F9B0" authorityName="Ermisch" authorityYear="1950" box="[837,969,1570,1593]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF26A8C60FCC2F992C0F9F9B0" box="[837,969,1570,1593]" italics="true" pageId="3" pageNumber="454">Mordellaria</emphasis>
</taxonomicName>
is distinguished from the other tumbling flower beetles by the presence of fine and straight dorsal striae on metatibia that stretch from its base to its apical margin. Metatarsomeres 1 and 2 also bear fine and straight dorsal striae.’ The authors also compared their species with
<taxonomicName id="4C33B4CDF26A8C60FCF6F972C72AF950" authorityName="Comolli" authorityYear="1837" box="[881,1050,1730,1753]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="aurofasciata">
<emphasis id="B947135CF26A8C60FCF6F972C0B7F950" box="[881,903,1730,1753]" italics="true" pageId="3" pageNumber="454">M</emphasis>
.
<emphasis id="B947135CF26A8C60FC13F972C72AF950" box="[916,1050,1730,1753]" italics="true" pageId="3" pageNumber="454">aurofasciata</emphasis>
</taxonomicName>
in which only a dorsolateral ridge is present on the metatibia and metatarsomere I. Although it looks like a simple mistake, it had a serious effect on the taxonomic conclusions of the authors. In addition, the situation is even more complex. In the generic assignment of his
<taxonomicName id="4C33B4CDF26A8C60FCE7F8D2C633F8F0" authority="Franciscolo, 1956" authorityName="Franciscolo" authorityYear="1956" box="[864,1283,1890,1913]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B947135CF26A8C60FCE7F8D2C70FF8F0" box="[864,1087,1890,1913]" italics="true" pageId="3" pageNumber="454">Mordellaria africana</emphasis>
Franciscolo, 1956
</taxonomicName>
, FRANCISCOLO (1956) points out that the delimitation of this genus was incomplete at that time.
<bibRefCitation id="EFA2B2BFF26A8C60FBC7F812C7C7F830" author="ERMISCH K." box="[1088,1271,1954,1977]" pageId="3" pageNumber="454" pagination="34 - 92" refId="ref20156" refString="ERMISCH K. 1950 a: Die Gattungen der Mordelliden der Welt. Entomologische Blatter 45 - 46: 34 - 92." type="journal article" year="1950">ERMISCH (1950a)</bibRefCitation>
established the genus
<taxonomicName id="4C33B4CDF26A8C60FC04F872C734F850" authorityName="Ermisch" authorityYear="1950" box="[899,1028,1986,2009]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF26A8C60FC04F872C734F850" box="[899,1028,1986,2009]" italics="true" pageId="3" pageNumber="454">Mordellaria</emphasis>
</taxonomicName>
based on
<taxonomicName id="4C33B4CDF26A8C60FBF1F872C0C5F870" authority="Fairmaire &amp; Germain, 1863" authorityName="Fairmaire &amp; Germain" authorityYear="1863" class="Insecta" family="Mordellidae" genus="Mordella" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="scripta">
<emphasis id="B947135CF26A8C60FBF1F872C61CF850" box="[1142,1324,1986,2009]" italics="true" pageId="3" pageNumber="454">Mordella scripta</emphasis>
Fairmaire &amp; Germain, 1863
</taxonomicName>
from
<collectingCountry id="F3248FDEF26A8C60FBB3F852C740F870" box="[1076,1136,2018,2041]" name="Chile" pageId="3" pageNumber="454">Chile</collectingCountry>
, but as FRANCISCOLO (1956) correctly states ‘Ermisch indicated that the hind tibiae bear a very distinct dorsal ridge, but does not specify whether this
<typeStatus id="548871ECF26D8C67FF41FF4EC3C4FE9C" box="[198,244,254,277]" pageId="4" pageNumber="455">type</typeStatus>
of ridge is present also on the first and second hind tarsal segments, as it is the case of
<taxonomicName id="4C33B4CDF26D8C67FDAAFEAEC270FEDC" authority="Erm." authorityName="Erm." class="Insecta" family="Mordellidae" genus="Pseudomordellaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="bifasciata">
<emphasis id="B947135CF26D8C67FDAAFEAEC230FEDC" italics="true" pageId="4" pageNumber="455">Pseudomordellaria bifasciata</emphasis>
Erm.
</taxonomicName>
from
<collectingCountry id="F3248FDEF26D8C67FE00FE8EC2FEFEDC" box="[391,462,318,341]" name="Brazil" pageId="4" pageNumber="455">Brazil</collectingCountry>
.’ As a consequence, FRAN- CISCOLO (1956) attributed his
<taxonomicName id="4C33B4CDF26D8C67FE4DFEEEC113FEFC" authorityName="Franciscolo" authorityYear="1956" box="[458,547,350,373]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="B947135CF26D8C67FE4DFEEEC113FEFC" box="[458,547,350,373]" italics="true" pageId="4" pageNumber="455">africana</emphasis>
</taxonomicName>
to
<taxonomicName id="4C33B4CDF26D8C67FDC6FEEEC1F0FEFC" authorityName="Ermisch" authorityYear="1950" box="[577,704,350,373]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF26D8C67FDC6FEEEC1F0FEFC" box="[577,704,350,373]" italics="true" pageId="4" pageNumber="455">Mordellaria</emphasis>
</taxonomicName>
, providing that in the
<typeStatus id="548871ECF26D8C67FEC8FECEC24DFE1C" box="[335,381,382,405]" pageId="4" pageNumber="455">type</typeStatus>
species of the genus ‘(a) the middle tibiae are exactly as long as the middle tarsi (shorter in
<taxonomicName id="4C33B4CDF26D8C67FD5CFE2EC2ACFE5C" authority="Fairm. &amp; Germ." authorityName="Fairmaire &amp; Germain" authorityYear="1863" class="Insecta" family="Mordellidae" genus="Mordella" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="scripta">
<emphasis id="B947135CF26D8C67FD5CFE2EC1C1FE3C" box="[731,753,414,437]" italics="true" pageId="4" pageNumber="455">M</emphasis>
.
<emphasis id="B947135CF26D8C67FF10FE0EC3D1FE5C" box="[151,225,446,469]" italics="true" pageId="4" pageNumber="455">scripta</emphasis>
Fairm. &amp; Germ.
</taxonomicName>
); and (b) the first tarsal segment of the hind legs bears a distinct dorso-lateral ridge.’ But in the description, he added ‘the second [metatarsomere] bears a rudimentary dorso-lateral ridge (see FRANCISCOLO 1956: fig. L-4)’. This uncertainty about the key characters attributed to putative
<taxonomicName id="4C33B4CDF26D8C67FE02FDEEC16DFDFC" box="[389,605,606,629]" pageId="4" pageNumber="455">
<emphasis id="B947135CF26D8C67FE02FDEEC136FDFC" box="[389,518,606,629]" italics="true" pageId="4" pageNumber="455">Mordellaria</emphasis>
species
</taxonomicName>
distributed in South American, Euro-Asian and African landmasses raises doubts about their congeneric relationship. Examination of the
<typeStatus id="548871ECF26D8C67FF58FD0EC23DFD5C" box="[223,269,702,725]" pageId="4" pageNumber="455">type</typeStatus>
of
<taxonomicName id="4C33B4CDF26D8C67FEB3FD0EC137FD5C" box="[308,519,702,725]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="scripta">
<emphasis id="B947135CF26D8C67FEB3FD0EC137FD5C" box="[308,519,702,725]" italics="true" pageId="4" pageNumber="455">Mordellaria scripta</emphasis>
</taxonomicName>
stored in The Natural History Museum, London has revealed that the dorsolateral ridge is present on the metatibia and metatarsomere I only, and that the differential diagnosis provided by
<bibRefCitation id="EFA2B2BFF26D8C67FDF9FCAEC240FCDC" author="PERKOVSKY E. E. &amp; ODNOSUM V. K." pageId="4" pageNumber="455" pagination="177 - 179" refId="ref21948" refString="PERKOVSKY E. E. &amp; ODNOSUM V. K. 2013: A new species of tumbling flower beetles of the genus Mordellaria (Insecta: Coleoptera: Mordellidae) from the Baltic amber. Paleontological Journal 47: 177 - 179." type="journal article" year="2013">PERKOVSKY &amp; ODNOSUM (2013)</bibRefCitation>
is incorrect also with respect to the
<typeStatus id="548871ECF26D8C67FF10FCEEC3F5FCFC" box="[151,197,862,885]" pageId="4" pageNumber="455">type</typeStatus>
species of the genus.
</paragraph>
<paragraph id="8B8CCF4EF26D8C67FF32FCCEC1C5FC5C" blockId="4.[151,760,190,981]" pageId="4" pageNumber="455">
Because characters of
<taxonomicName id="4C33B4CDF26D8C67FE21FCCEC11FFC1C" box="[422,559,894,917]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="friedrichi">
<emphasis id="B947135CF26D8C67FE21FCCEC11FFC1C" box="[422,559,894,917]" italics="true" pageId="4" pageNumber="455">M. friedrichi</emphasis>
</taxonomicName>
do not correspond with the generic characteristic of
<taxonomicName id="4C33B4CDF26D8C67FE7CFC2EC14CFC3C" authorityName="Ermisch" authorityYear="1950" box="[507,636,926,949]" class="Insecta" family="Mordellidae" genus="Mordellaria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF26D8C67FE7CFC2EC14CFC3C" box="[507,636,926,949]" italics="true" pageId="4" pageNumber="455">Mordellaria</emphasis>
</taxonomicName>
the species is transferred here into
<taxonomicName id="4C33B4CDF26D8C67FE09FC0EC137FC5C" box="[398,519,958,981]" class="Insecta" family="Mordellidae" genus="Baltimorda" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF26D8C67FE09FC0EC137FC5C" box="[398,519,958,981]" italics="true" pageId="4" pageNumber="455">Baltimorda</emphasis>
</taxonomicName>
new collective group.
</paragraph>
</subSubSection>
</treatment>
</document>
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