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<document id="A2B5BDED53D719AE2099FE9089B2C464" ID-DOI="10.37520/aemnp.2023.027" ID-ISSN="1804-6487" ID-Zenodo-Dep="10621277" ID-ZooBank="4F6482C1-A2F2-4406-A115-B8E0F205806C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707164847204" checkinUser="felipe" docAuthor="Batelka, Jan, Rosová, Kateřina &amp; Prokop, Jakub" docDate="2023" docId="039A7E58F2628C68FEBDFC84C65FFB1C" docLanguage="en" docName="ActaEntMusNatPra.63.2.451-478.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 63 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2023.027" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Baltistena aurata Batelka &amp; Rosová &amp; Prokop 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="462" masterDocId="FFA30620F2698C63FF87FFB0C330FF89" masterDocTitle="Diversity and morphology of Eocene and Oligocene Mordellidae (Coleoptera)" masterLastPageNumber="478" masterPageNumber="451" pageNumber="462" updateTime="1707165608224" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="D709F619DC569491EB43C895F296B80D">Diversity and morphology of Eocene and Oligocene Mordellidae (Coleoptera)</mods:title>
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<mods:namePart id="45B8A6C49D215F4E612B8A2985C5A999">Rosová, Kateřina</mods:namePart>
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<subSubSection id="C3299CC5F2628C68FEBDFC84C165FCC5" box="[314,597,820,844]" pageId="11" pageNumber="462" type="nomenclature">
<paragraph id="8B8CCF4EF2628C68FEBDFC84C165FCC5" blockId="11.[314,597,820,844]" box="[314,597,820,844]" pageId="11" pageNumber="462">
<heading id="D0C47822F2628C68FEBDFC84C165FCC5" box="[314,597,820,844]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="462" reason="3">
<emphasis id="B947135CF2628C68FEBDFC84C165FCC5" bold="true" box="[314,597,820,844]" pageId="11" pageNumber="462">
<taxonomicName id="4C33B4CDF2628C68FEBDFC84C2C9FCC2" authority="Batelka &amp; Rosová &amp; Prokop, 2023" authorityName="Batelka &amp; Rosová &amp; Prokop" authorityYear="2023" box="[314,505,820,844]" class="Insecta" family="Mordellidae" genus="Baltistena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="aurata" status="sp. nov.">
<emphasis id="B947135CF2628C68FEBDFC84C2C9FCC2" bold="true" box="[314,505,820,844]" italics="true" pageId="11" pageNumber="462">Baltistena aurata</emphasis>
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<taxonomicNameLabel id="A274AE27F2628C68FE78FC85C165FCC5" box="[511,597,821,844]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3299CC5F2628C68FE16FCE3C2CEFCEF" box="[401,510,851,870]" pageId="11" pageNumber="462" type="description">
<paragraph id="8B8CCF4EF2628C68FE16FCE3C2CEFCEF" blockId="11.[401,510,851,870]" box="[401,510,851,870]" pageId="11" pageNumber="462">
(
<figureCitation id="1308D3CBF2628C68FE10FCE3C2C8FCEF" box="[407,504,851,870]" captionStart="Figs 6" captionStartId="14.[151,188,1743,1762]" captionTargetBox="[151,1436,189,1720]" captionTargetId="figure-13@14.[151,1436,189,1720]" captionTargetPageId="14" captionText="Figs 6. AG Mordellistenini, Baltic amber.AC Baltistena ultima sp. nov., holotype, PřFUK65:A habitus; B antenna; C hind leg and hypopygium. DG Baltistena aurata sp. nov., holotype, PřFUK66: D habitus; E hind leg; F head frontally (maxillary palpomere IV marked by black arrow); G hypopygium. Scale bars: A, D 0.5 mm; C, E, F 0.3 mm; B, G 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10621291" httpUri="https://zenodo.org/record/10621291/files/figure.png" pageId="11" pageNumber="462">Figs 6DG</figureCitation>
)
</paragraph>
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<subSubSection id="C3299CC5F2628C68FF10FCCDC1C7FC23" pageId="11" pageNumber="462" type="materials_examined">
<paragraph id="8B8CCF4EF2628C68FF10FCCDC1C7FC23" blockId="11.[151,761,893,938]" pageId="11" pageNumber="462">
<emphasis id="B947135CF2628C68FF10FCCDC22FFC19" bold="true" box="[151,287,893,912]" pageId="11" pageNumber="462">Type material.</emphasis>
<typeStatus id="548871ECF2628C68FEA0FCCDC2B2FC19" box="[295,386,893,912]" pageId="11" pageNumber="462" type="holotype">HOLOTYPE</typeStatus>
: PřfUK66, obtained from rmvveta (www. ambertreasure4u.com) (original number 224239532480): Baltic amber.
</paragraph>
</subSubSection>
<subSubSection id="C3299CC5F2628C68FF10FC73C753FD5C" pageId="11" pageNumber="462" type="description">
<paragraph id="8B8CCF4EF2628C68FF10FC73C753FD5C" blockId="11.[151,761,963,2074]" lastBlockId="11.[828,1437,190,1173]" pageId="11" pageNumber="462">
<emphasis id="B947135CF2628C68FF10FC73C212FC53" bold="true" box="[151,290,963,986]" pageId="11" pageNumber="462">Description.</emphasis>
Head subglobular, frons continuously convex, posterior edge of eye near posterior margin of head, elytra convex, metacoxae broad, comb formula 3//3/1/0/0, pygidium long. Main characters: Right antenna clearly visible, length ratios of antennomeres as follows: 1.25- 1.4-1-&gt;1-&gt;1-1.4-1.25-1.4-1.4-1.25-&lt;2 (antennomeres IV and V only slightly longer than antennomere III), antennomeres IX cylindrical, antennomere XI malleiform, widest in middle, all antennomeres distinctly longer than wide with ratio: 1.75-2-1.5-1.45-1.5-2-2.3-2.6-2.6-1.9-3.9. Maxillary palpomere II robust, almost as long as wide, palpomere III short, triangular, palpomere IV malleiform of
<taxonomicName id="4C33B4CDF2628C68FF32FAF3C264FAD3" authorityName="Emery" authorityYear="1876" box="[181,340,1347,1370]" class="Insecta" family="Mordellidae" genus="Mordellochroa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="B947135CF2628C68FF32FAF3C264FAD3" box="[181,340,1347,1370]" italics="true" pageId="11" pageNumber="462">Mordellochroa</emphasis>
</taxonomicName>
-
<typeStatus id="548871ECF2628C68FEDCFAF3C2BBFAD3" box="[347,395,1347,1370]" pageId="11" pageNumber="462">type</typeStatus>
(
<bibRefCitation id="EFA2B2BFF2628C68FE1DFAF3C181FAD3" author="FRANCISCOLO M. E." box="[410,689,1347,1370]" pageId="11" pageNumber="462" pagination="207 - 291" refId="ref20488" refString="FRANCISCOLO M. E. 1957 a: Chapter V. Coleoptera: Mordellidae. A monograph of the South African genera and species. 1. Morphology, subfamily Ctenidiinae and tribe Stenaliini. Pp. 207 - 291. In: HAN- STROM B., BRINCK P. &amp; RUDEBECK G. (eds): South African Animal Life, Results of the Lund University Expedition in 1950 - 1951. Vol. 4. Almqvist &amp; Wiksell, Stockholm, 508 pp." type="book chapter" year="1957">FRANCISCOLO 1957a: 216</bibRefCitation>
). Eyes roughly faceted. Eyes glabrous. Form of scutellum indiscernible. Form of basal side of pronotum indiscernible. Elytra 3.1× as long as pronotal disc. Form of protarsi: protarsomeres cylindrical, tarsomere I 6.25× as long as wide; tarsomere II 4× as long as wide; tarsomere III 3× as long as wide; tarsomere IV oblique at apex but without visible emargination; tarsomere V about 3× as long as wide. Structure of protibia indiscernible. Form of mesotarsomere IV: mesotarsomeres cylindrical, tarsomere I 5.6× as long as wide; tarsomere II 3.6× as long as wide; tarsomere III 3.5× as long as wide; tarsomere IV 2× as long as wide excised almost down to middle of its length; tarsomere V about 2.5× as long as wide. Mesotibia 1.5× as long as mesotarsal segments combined.
<typeStatus id="548871ECF2628C68FDC4F8B3C148F893" box="[579,632,1795,1818]" pageId="11" pageNumber="462">Type</typeStatus>
of comb on metatibia: three distinct lateral combs of scale-like setae including preapical comb, combs do not reach middle of metatibia; preapical comb runs parallel with apical fringe of setae, remaining two combs oriented at different angles; few isolated patches of scale-like setae present behind proximal dorso-lateral comb. Metatibia with two spurs, outer spur shorter and sharply pointed. Metatarsomere I with row of spiniform setae consisting of about 20 short strong setae ventrally, metatarsomeres IIIV with row of spiniform setae consisting ventrally of about 17 short strong setae; metatarsomere I with three short lateral combs of scale-like setae; metatarsomere II with one short lateral comb of scale-like setae, metatarsomeres IIIIV without combs; length ratio of metatarsal segments 1.75-1-1-1. Pygidium long and straight, 3.1× as long as hypopygium. Shape of metanepisternum: long and narrow, lower corner of posterior edge rounded, anterior edge 3× as long as posterior edge, ventral edge 5.5× as long as posterior edge. Metakatepisternum not discernible. Pretarsal claws long and straight, with 23 indistinct teeth on ventral edge. Tip of pygidium with distinct cluster of black scale-like setae. Lengths (in mm): pronotal disc = 0.54, elytra = 2.16, mesotibia = 0.8, metatarsomere I = 0.34, pygidium = 0.9.
<emphasis id="B947135CF2628C68FCBBFD2EC08CFD3C" bold="true" box="[828,956,670,693]" pageId="11" pageNumber="462">Etymology.</emphasis>
The species name refers to the golden appearance of the body; adjective.
</paragraph>
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<subSubSection id="C3299CC5F2628C68FCBBFD6EC65FFB1C" pageId="11" pageNumber="462" type="diagnosis">
<paragraph id="8B8CCF4EF2628C68FCBBFD6EC65FFB1C" blockId="11.[828,1437,190,1173]" pageId="11" pageNumber="462">
<emphasis id="B947135CF2628C68FCBBFD6EC704FD7C" bold="true" box="[828,1076,734,757]" pageId="11" pageNumber="462">Differential diagnosis.</emphasis>
Although the structure of mouthparts, shape of antennae, legs and pronotal disc, and overall habitus of
<taxonomicName id="4C33B4CDF2628C68FC2AFCAEC757FCBC" authority="Batelka &amp; Rosová &amp; Prokop, 2023" authorityName="Batelka &amp; Rosová &amp; Prokop" authorityYear="2023" box="[941,1127,798,821]" class="Insecta" family="Mordellidae" genus="Baltistena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="aurata" status="sp. nov.">
<emphasis id="B947135CF2628C68FC2AFCAEC757FCBC" box="[941,1127,798,821]" italics="true" pageId="11" pageNumber="462">Baltistena aurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A274AE27F2628C68FBE9FCAEC7F1FCBC" box="[1134,1217,798,821]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
appear familiar, this species cannot be placed in any extant genus of
<taxonomicName id="4C33B4CDF2628C68FAABFC8EC049FCFC" class="Insecta" family="Mordellidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="tribe" tribe="Mordellistenini">Mordellistenini</taxonomicName>
because of the following characters: length over
<quantity id="4CCB62ABF2628C68FCBBFCCEC0B0FC1C" box="[828,896,894,917]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="462" unit="mm" value="4.0">4 mm</quantity>
; eyes glabrous; frons convex; terminal maxillary palpomere malleiform; antenna filiform, antennomeres cylindrical; antennomeres VX 1.52.6× as long as wide; antennomere XI 1.6× as long as preceding one; anterior edge of pronotal disc rounded; pro- and mesotarsomere IV emarginated, shallowly bilobed; metatibial combs short; both metatibial spurs long, only slightly dissimilar. For its comparison with 37 extant genera and subgenera having pro- and mesotarsomere IV emarginated see
<tableCitation id="C6B1FAF5F2628C68FA9EFBCEC658FB1C" box="[1305,1384,1150,1173]" captionStart="Table 1" captionStartId="12.[151,199,190,209]" captionText="Table 1. Comparison of Baltistena aurata sp. nov. with extant genera of Mordellistenini." pageId="11" pageNumber="462">Table 1</tableCitation>
.
</paragraph>
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