181 lines
28 KiB
XML
181 lines
28 KiB
XML
<document id="BD6BF6E20ED0EA46FB9246D87CFF01B7" ID-ISSN="0024-4082" ID-ZooBank="72117203-4FF3-4E2B-837E-646B330B30D4" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630103366865" checkinUser="felipe" docAuthor="Bisconti, Michelangelo, Damarco, Piero, Pavia, Marco, Sorce, Barbara & Carnevale, Giorgio" docDate="2021" docId="0392E34BFFDDA44AFF0FB31D00F0FE41" docLanguage="en" docName="zlaa131.pdf" docOrigin="Zoological Journal of the Linnean Society 192" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Marzanoptera tersillae Bisconti & Damarco & Pavia & Sorce & Carnevale 2021, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="1278" masterDocId="FFAB9B33FFC5A453FFE4B55C014CFFF3" masterDocTitle="Marzanoptera tersillae, a new balaenopterid genus and species from the Pliocene of Piedmont, north-west Italy" masterLastPageNumber="1292" masterPageNumber="1253" pageNumber="1277" updateTime="1699153126933" updateUser="plazi">
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<mods:title id="3E69C5149AC25C47A899E1D7757DB3DD">Marzanoptera tersillae, a new balaenopterid genus and species from the Pliocene of Piedmont, north-west Italy</mods:title>
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<mods:namePart id="6B10B51EBE59CF0C48455D36E35D1120">Bisconti, Michelangelo</mods:namePart>
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<mods:namePart id="D95A74016897D437C682A97B6A6B75B1">Damarco, Piero</mods:namePart>
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<mods:roleTerm id="14E930121EB727CCBE60BD4B15D0661D">Author</mods:roleTerm>
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<mods:namePart id="DCB04F447B927CECCAD304405088713C">Pavia, Marco</mods:namePart>
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<mods:namePart id="58F97FFC6956AC0F2572A36B2C5AE412">Sorce, Barbara</mods:namePart>
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<mods:namePart id="7F1F6D94EC7E0BD38A7822124D847BD9">Carnevale, Giorgio</mods:namePart>
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<mods:title id="969C798E15D0344C2A0B065AC9E2713C">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="213F6EE3D9DB0BDA5FE522915CC592D5">
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<mods:date id="31B444A6B2AC01BC871C76329A556D6B">2021</mods:date>
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<mods:detail id="B6590CD2F62C232C4D71347CC5D70DAE" type="volume">
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<mods:number id="5403D4422ED58306A582E059AFBE06A1">192</mods:number>
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<mods:start id="A79CEDBF449C1CC055B21D2A4127D4C4">1253</mods:start>
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<mods:end id="FEA449417FC1E69D75285BCC99541A87">1292</mods:end>
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<subSubSection id="C32101D6FFDDA44BFF0FB31D03D2F9AA" box="[235,670,1601,1625]" pageId="24" pageNumber="1277" type="nomenclature">
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<paragraph id="8B84525DFFDDA44BFF0FB31D03D2F9AA" blockId="24.[235,670,1601,1625]" box="[235,670,1601,1625]" pageId="24" pageNumber="1277">
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<heading id="D0CCE531FFDDA44BFF0FB31D03D2F9AA" box="[235,670,1601,1625]" centered="true" fontSize="9" level="2" pageId="24" pageNumber="1277" reason="2">
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<taxonomicName id="4C3B29DEFFDDA44BFF0FB31D0378F9A4" authority="Bisconti & Damarco & Pavia & Sorce & Carnevale, 2021" authorityName="Bisconti & Damarco & Pavia & Sorce & Carnevale" authorityYear="2021" box="[235,564,1601,1624]" class="Mammalia" family="Balaenopteridae" genus="Marzanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="24" pageNumber="1292" phylum="Chordata" rank="species" species="tersillae" status="sp. nov.">MARZANOPTERA TERSILLAE</taxonomicName>
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<emphasis id="B94F8E4FFFDDA44BFDDFB31803D2F9AA" bold="true" box="[571,670,1601,1625]" pageId="24" pageNumber="1277">
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<taxonomicNameLabel id="A27C3334FFDDA44BFDDFB31803D2F9AA" box="[571,670,1601,1625]" pageId="24" pageNumber="1277" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32101D6FFDDA44AFE66B33100F0FE41" lastPageId="25" lastPageNumber="1278" pageId="24" pageNumber="1277" type="description">
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<paragraph id="8B84525DFFDDA44BFE66B331034BF975" blockId="24.[386,519,1645,1670]" box="[386,519,1645,1670]" pageId="24" pageNumber="1277">
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(
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<figureCitation id="13004ED8FFDDA44BFE6EB331034CF975" box="[394,512,1645,1670]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-10="Figure 14" captionStart-11="Figure 15" captionStart-12="Figure 16" captionStart-2="Figure 6" captionStart-3="Figure 7" captionStart-4="Figure 8" captionStart-5="Figure 9" captionStart-6="Figure 10" captionStart-7="Figure 11" captionStart-8="Figure 12" captionStart-9="Figure 13" captionStartId-0="6.[145,223,1377,1399]" captionStartId-1="7.[163,243,1137,1159]" captionStartId-10="16.[145,223,1508,1530]" captionStartId-11="18.[144,224,1632,1654]" captionStartId-12="21.[163,249,1700,1722]" captionStartId-2="9.[163,242,1417,1439]" captionStartId-3="10.[145,224,760,782]" captionStartId-4="10.[145,224,1714,1736]" captionStartId-5="11.[163,243,1678,1700]" captionStartId-6="12.[145,224,1846,1868]" captionStartId-7="13.[162,243,943,965]" captionStartId-8="14.[145,223,1671,1693]" captionStartId-9="15.[163,242,1616,1638]" captionTargetBox-0="[386,1185,195,1337]" captionTargetBox-1="[387,1218,195,1097]" captionTargetBox-10="[227,1342,199,1464]" captionTargetBox-11="[337,1233,195,1592]" captionTargetBox-12="[232,710,907,1660]" captionTargetBox-2="[402,1202,197,1376]" captionTargetBox-3="[208,1361,197,719]" captionTargetBox-4="[226,1343,999,1674]" captionTargetBox-5="[416,1190,197,1637]" captionTargetBox-6="[369,1200,1099,1807]" captionTargetBox-7="[389,1215,196,902]" captionTargetBox-8="[338,1233,195,1631]" captionTargetBox-9="[325,1281,197,1575]" captionTargetId-0="figure-255@6.[385,1185,195,1337]" captionTargetId-1="figure-342@7.[387,1218,195,1097]" captionTargetId-10="figure-144@16.[223,1346,195,1468]" captionTargetId-11="figure-70@18.[337,1233,195,1592]" captionTargetId-12="figure-667@21.[231,711,906,1661]" captionTargetId-2="figure-196@9.[401,1204,195,1377]" captionTargetId-3="figure-26@10.[207,1362,195,720]" captionTargetId-4="figure-130@10.[225,1345,998,1675]" captionTargetId-5="figure-24@11.[414,1192,195,1638]" captionTargetId-6="figure-474@12.[369,1200,1099,1807]" captionTargetId-7="figure-479@13.[388,1216,195,903]" captionTargetId-8="figure-80@14.[337,1233,195,1631]" captionTargetId-9="figure-94@15.[322,1283,195,1576]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-10="16" captionTargetPageId-11="18" captionTargetPageId-12="21" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="Figure 3. Holotype skull of Marzanoptera tersillae in dorsal view.A, left premaxilla. B, right premaxilla. C, photographic representation of the skull. D, anatomical interpretation of the skull. Scale bar: 20 cm." captionText-1="Figure 4. Computed tomography (CT) scan reconstruction of the holotype skull of Marzanoptera tersillae in ventral view. Note that the squamosals are not included because of size limitations for the CT scanning machine. Scale bar: 20 cm." captionText-10="Figure 14. Postcranial skeleton of Marzanoptera tersillae (holotype). A, atlas in anterior view. B, atlas in posterior view. C, axis in anterior view. D, axis in posterior view. E, C3 in anterior view. F, C4 in anterior view. G, C5 in anterior view. H, C6 in anterior view. I, C7 in anterior view. J, T1 in anterior view. K–M, anterior thoracic vertebrae in anterior view. N, vertebral epiphyses. Scale bars: 20 cm." captionText-11="Figure 15. Postcranial skeleton of Marzanoptera tersillae (holotype). A, B, thoracic vertebrae in ventrolateral view, showing a ventral keel (not to scale; A, same vertebra as shown in K; B, same vertebra as shown in L). C, D, undetermined thoracic vertebra. E, F, additional undetermined thoracic vertebra. G, H, undetermined lumbar vertebra. I, J, undetermined thoracic vertebra. K, ribs in anterior view. Scale bars: 15 cm in C–J; 10 cm in K." captionText-12="Figure 16. Evidence of shark interactions. Ribs of Marzanoptera tersillae (holotype) showing tips of shark teeth embedded into the bone. A, double-headed rib. B, proximal diaphysis of a double-headed rib. C, proximal diaphysis of a possibly double-headed rib. D, diaphysis of a rib. White triangles indicate the shark teeth. Scale bars: 10 cm." captionText-2="Figure 6. Holotype skull of Marzanoptera tersillae in lateral view. A, photographic representation of the skull in right lateral view. B, anatomical interpretation. C, photographic representation of the skull in left lateral view. D, anatomical interpretation. Scale bar: 10 cm." captionText-3="Figure 7. Holotype skull of Marzanoptera tersillae in anterior and posterior views. Note that the squamosals are not included because of size limitations for the computed tomography (CT) scanning machine. A, CT scan-based tridimensional rendering of the skull in anterior view.B, cross-section of the skull at the level of the interorbital region of the frontal, showing the squared outline of the depression medial to the supraorbital processes of the frontal. C, CT scan-based tridimensional rendering of the skull in posterior view. B, photographic representation of the skull in posterior view from a slightly ventral point of view. Scale bar: 30 cm." captionText-4="Figure 8. Vertex of the holotype skull of Marzanoptera tersillae. A, computed tomography scan view of the vertex. B, anatomical interpretation. Scale bar: 30 cm." captionText-5="Figure 9. Temporal fossa of the holotype skull of Marzanoptera tersillae. Note that the squamosals are not included because of size limitations for the computed tomography (CT) scanning machine. A, CT scan-based tridimensional reconstruction of the right temporal fossa and the skull in anterolateral view. B, anatomical interpretation. C, CT scan-based tridimensional reconstruction of the left temporal fossa and the skull in anterolateral view. D, anatomical interpretation. Scale bar: 30 cm." captionText-6="Figure 10. Right posterolateral portion of the holotype skull of Marzanoptera tersillae showing the posterior lacerate foramen and associated anatomical structures based on computed tomography scan reconstruction. Scale bar: 10 cm." captionText-7="Figure 11. Left posterolateral portion of the holotype skull of Marzanoptera tersillae showing the left periotic in articulation with the skull and other associated anatomical structures based on computed tomography scan reconstruction. Scale bar: 10 cm." captionText-8="Figure 12. Left periotic of Marzanoptera tersillae (holotype skull) based on computed tomography scan reconstructions. A, ventral view. B, posterolateral view. C, posterior view. D–F, details of the pars cochlearis of the left periotic in medial (D) posterior (E) and posteromedial (F) views. Scale bars: 5 cm." captionText-9="Figure 13. Virtual cross-sections of the pars cochlearis of the left periotic of Marzanoptera tersillae showing the path of the facial nerve within the promontorium based on computed tomography scan reconstructions. In A–D, the arrowheads indicate the positions of the cross-sections of the periotic in ventral view. E–H, close-up views of the pars cochlearis in medial view. I–L, anatomical interpretations of images in E–H. Grey areas represent section areas. Not to scale." pageId="24" pageNumber="1277">FIGS 3–16</figureCitation>
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)
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</paragraph>
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<paragraph id="8B84525DFFDDA44BFF75B3EA03B4F919" blockId="24.[145,762,1717,1770]" pageId="24" pageNumber="1277">
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<emphasis id="B94F8E4FFFDDA44BFF75B3EA009AF938" box="[145,470,1717,1739]" italics="true" pageId="24" pageNumber="1277">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n: l s i d: z o o b a n k. org:act:
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<uuid id="FF9D6888FFDDA44BFF3BB38803B4F919" box="[223,760,1748,1770]" pageId="24" pageNumber="1277">F3B4AA64-BEBC-47FA-8DE5-19B777B75BB1</uuid>
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</paragraph>
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<paragraph id="8B84525DFFDDA44BFF75B24E0324F8B4" blockId="24.[145,760,1810,1863]" pageId="24" pageNumber="1277">
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<emphasis id="B94F8E4FFFDDA44BFF75B24E005AF8D4" box="[145,278,1810,1831]" italics="true" pageId="24" pageNumber="1277">Etymology:</emphasis>
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The fossil skeleton was discovered by Tersilla Argenta, for whom it was named.
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</paragraph>
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<paragraph id="8B84525DFFDDA44BFCCDB5990459FEC4" blockId="24.[809,1424,197,311]" pageId="24" pageNumber="1277">
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<materialsCitation id="3B535800FFDDA44BFCCDB5990459FEC4" collectionCode="EGPPA-MPTA" pageId="24" pageNumber="1277" specimenCount="1" typeStatus="holotype">
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<emphasis id="B94F8E4FFFDDA44BFCCDB59902D2FF29" box="[809,926,197,218]" italics="true" pageId="24" pageNumber="1277">
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<typeStatus id="5480ECFFFFDDA44BFCCDB59902DBFF29" box="[809,919,197,218]" pageId="24" pageNumber="1277" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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Specimen
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<emphasis id="B94F8E4FFFDDA44BFBDEB59905F5FF29" box="[1082,1209,197,218]" italics="true" pageId="24" pageNumber="1277">207.13307</emphasis>
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of the inventory of
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<collectionCode id="ED2ACA98FFDDA44BFCACB5B802BBFF0A" box="[840,1015,228,249]" pageId="24" pageNumber="1277">EGPPA-MPTA</collectionCode>
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institution in Asti. The specimen includes most of the skull, with one periotic in articulation, seven vertebrae and nine ribs.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B84525DFFDDA44BFCCDB43C028AF96F" blockId="24.[809,1425,352,1692]" pageId="24" pageNumber="1277">
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<emphasis id="B94F8E4FFFDDA44BFCCDB43C0578FE86" box="[809,1076,352,373]" italics="true" pageId="24" pageNumber="1277">Horizon and locality:</emphasis>
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The
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<typeStatus id="5480ECFFFFDDA44BFB9FB43C05A8FE85" box="[1147,1252,352,374]" pageId="24" pageNumber="1277" type="holotype">holotype</typeStatus>
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skeleton was discovered near San Marzanotto d’Asti (~
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<quantity id="4CC3FFB8FFDDA44BFAABB42204DCFE67" box="[1359,1424,382,404]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="24" pageNumber="1277" unit="km" value="6.0">6 km</quantity>
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South from Asti,
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<collectingRegion id="49FF9CBFFFDDA44BFBE3B4C105CDFE40" box="[1031,1153,413,435]" country="Italy" name="Piemonte" pageId="24" pageNumber="1277">Piedmont</collectingRegion>
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, northern
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<collectingCountry id="F32C12CDFFDDA44BFAEFB4C10406FE40" box="[1291,1354,413,435]" name="Italy" pageId="24" pageNumber="1277">Italy</collectingCountry>
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). The geographical coordinates of the site of the discovery are
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<geoCoordinate id="EE0F349AFFDDA44BFCBFB4860546FE03" box="[859,1034,473,496]" degrees="44" direction="north" minutes="51" orientation="latitude" pageId="24" pageNumber="1277" precision="1" seconds="54.36" value="44.865097">44°51′54.36″N</geoCoordinate>
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and
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<geoCoordinate id="EE0F349AFFDDA44BFBA9B48705A0FE03" box="[1101,1260,473,496]" degrees="8" direction="east" minutes="13" orientation="longitude" pageId="24" pageNumber="1277" precision="1" seconds="13.33" value="8.220369">8°13′13.33″E</geoCoordinate>
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(
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<figureCitation id="13004ED8FFDDA44BFAE4B486040BFE03" box="[1280,1351,474,496]" captionStart="Figure 1" captionStartId="1.[827,906,1251,1273]" captionTargetBox="[895,1375,195,1211]" captionTargetId="figure-684@1.[895,1375,195,1211]" captionTargetPageId="1" captionText="Figure 1. Map showing the discovery site of the holotype skeleton of Marzanoptera tersillae. A, the Italian peninsula. B, the locality of the discovery. The arrow indicates the site of San Marzanotto, which is located ~5 km South from Asti." pageId="24" pageNumber="1277">Fig. 1</figureCitation>
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;
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<bibRefCitation id="EFAA2FACFFDDA44BFABCB48602B0FDFD" author="Boni A & Casnedi R" pageId="24" pageNumber="1277" refId="ref23268" refString="Boni A, Casnedi R. 1970. Note illustrative della Carta d'Italia. F. 69. Asti, Roma: ISPRA." type="book" year="1970">Boni & Casnedi, 1970</bibRefCitation>
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). The specimen was found in a hilly area in Asti Province, a site previously known because of the discovery of several marine mammals, including an almost complete, 9-m-long skeleton of a large balaenopterid whale assigned to ‘
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<taxonomicName id="4C3B29DEFFDDA44BFB11B728056FFD5B" authorityName="' (Caretto" authorityYear="1970" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="24" pageNumber="1277" phylum="Chordata" rank="subSpecies" species="acutorostrata" subSpecies="cuvieri">
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<emphasis id="B94F8E4FFFDDA44BFB11B728056FFD5B" italics="true" pageId="24" pageNumber="1277">Balaenoptera acutorostrata cuvieri</emphasis>
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</taxonomicName>
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’ by
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<bibRefCitation id="EFAA2FACFFDDA44BFBB2B7CF0448FD5A" author="Caretto PG" box="[1110,1284,659,681]" pageId="24" pageNumber="1277" pagination="3 - 75" refId="ref23606" refString="Caretto PG. 1970. La balenottera delle sabbie plioceniche di Valmontasca (Vigliano d'Asti). Bollettino della Societa Paleontologica Italiana 9: 3 - 75." type="journal article" year="1970">Caretto (1970)</bibRefCitation>
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, which was discovered in Valmontasca (Vigliano d’Asti). That skeleton was subsequently regarded as a new taxon (still to be described formally) by
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<bibRefCitation id="EFAA2FACFFDDA44BFB53B7B20425FCF6" author="Bisconti M" box="[1207,1385,750,773]" pageId="24" pageNumber="1277" pagination="147 - 156" refId="ref22590" refString="Bisconti M. 2009. Taxonomy and evolution of the Italian Pliocene Mysticeti (Mammalia, Cetacea): a state of the art. Bollettino della Societa Paleontologica Italiana 48: 147 - 156." type="journal article" year="2009">Bisconti (2009)</bibRefCitation>
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. In this area,
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<bibRefCitation id="EFAA2FACFFDDA44BFC42B651051AFCD0" author="Caretto PG" box="[934,1110,781,803]" pageId="24" pageNumber="1277" pagination="3 - 75" refId="ref23606" refString="Caretto PG. 1970. La balenottera delle sabbie plioceniche di Valmontasca (Vigliano d'Asti). Bollettino della Societa Paleontologica Italiana 9: 3 - 75." type="journal article" year="1970">Caretto (1970)</bibRefCitation>
|
||
described a stratigraphic sequence ~
|
||
<quantity id="4CC3FFB8FFDDA44BFC42B67002ACFCB2" box="[934,992,812,834]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="24" pageNumber="1277" unit="m" value="53.0">53 m</quantity>
|
||
thick. At the bottom of this sequence, there are
|
||
<quantity id="4CC3FFB8FFDDA44BFC73B616028DFC93" box="[919,961,842,864]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="1277" unit="m" value="5.0">5 m</quantity>
|
||
of clayey sediments in Piacenzian facies, overlapped by ~
|
||
<quantity id="4CC3FFB8FFDDA44BFC01B635056CFC8C" box="[997,1056,873,895]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="24" pageNumber="1277" unit="m" value="48.0">48 m</quantity>
|
||
of sands alternating with silty sand. In this second part, there are two layers with abundant molluscs, dominated by bivalves (Supporting Information,
|
||
<tableCitation id="C6B967E6FFDDA44BFC2AB6990577FC29" box="[974,1083,965,987]" captionStart="Table S" captionStartId="39.[163,233,658,679]" captionText="Table S1. List of molluscan taxa found in close association with the holotype skeleton of Marzanoptera tersillae. Table S2. Measurements of the holotype skull of Marzanoptera tersillae (EGPPA-MPTA No. 207.13307, Asti). Measurements are in millimetres. Numbers on the right are from the right side; numbers on the left are from the left side." pageId="24" pageNumber="1277">Table S1</tableCitation>
|
||
;
|
||
<figureCitation id="13004ED8FFDDA44BFBA8B69905E8FC29" box="[1100,1188,965,987]" captionStart="Figure 1" captionStartId="1.[827,906,1251,1273]" captionTargetBox="[895,1375,195,1211]" captionTargetId="figure-684@1.[895,1375,195,1211]" captionTargetPageId="1" captionText="Figure 1. Map showing the discovery site of the holotype skeleton of Marzanoptera tersillae. A, the Italian peninsula. B, the locality of the discovery. The arrow indicates the site of San Marzanotto, which is located ~5 km South from Asti." pageId="24" pageNumber="1277">Fig. S1</figureCitation>
|
||
). Polychaete tubes (
|
||
<taxonomicName id="4C3B29DEFFDDA44BFCD5B6B80544FC0A" baseAuthorityName="Linnaeus" baseAuthorityYear="1767" box="[817,1032,996,1018]" class="Polychaeta" family="Serpulidae" genus="Ditrupa" kingdom="Animalia" order="Sabellida" pageId="24" pageNumber="1277" phylum="Annelida" rank="species" species="cornea">
|
||
<emphasis id="B94F8E4FFFDDA44BFCD5B6B802C3FC0A" box="[817,911,996,1017]" italics="true" pageId="24" pageNumber="1277">Ditrupa</emphasis>
|
||
cf.
|
||
<emphasis id="B94F8E4FFFDDA44BFC5FB6B80544FC0A" box="[955,1032,996,1017]" italics="true" pageId="24" pageNumber="1277">cornea</emphasis>
|
||
</taxonomicName>
|
||
) are also abundant in this fossil community lying beneath and near the bones. The particular diagenetic conditions associated with the water circulation in the sediments did not allow the preservation of the calcareous tests of foraminiferans except in a few cases. The few foraminiferans found showed worn tests and unrecognizable morphological features. Therefore, the study of the foraminiferans found at the site did not allow an accurate stratigraphic reconstruction. To obtain a more accurate chronostratigraphic assessment of the site, the mollusc assemblage was compared with assemblages from other sites of the Asti area where foraminiferans were more abundant (
|
||
<bibRefCitation id="EFAA2FACFFDDA44BFC1CB02E05E0FA74" author="Scarselli S" box="[1016,1196,1394,1416]" pageId="24" pageNumber="1277" refId="ref26574" refString="Scarselli S. 1990. Variazioni nelle malacofaune piloceniche della sommita delle Argille di Lugagnano ad Ovest di Asti. Unpublished MSc Thesis, Universita di Torino." type="book" year="1990">Scarselli, 1990</bibRefCitation>
|
||
). This comparison resulted in the conclusion that the whale-bearing horizon corresponds approximately to the higher part of the Argille Azzurre Formation, close to the overlying Sabbie d’Asti Formation (
|
||
<bibRefCitation id="EFAA2FACFFDDA44BFBB4B0B105A3F9F0" author="Carraro F" box="[1104,1263,1517,1539]" pageId="24" pageNumber="1277" pagination="36 - 38" refId="ref23634" refString="Carraro F. 1996. Revisione del Villafranchiano nell'area-tipo di Villafranca d'Asti. Il Quaternario 9: 36 - 38." type="journal article" year="1996">Carraro, 1996</bibRefCitation>
|
||
). The mollusc assemblage and the overall stratigraphic data support the hypothesis that the estimated age of the whale-bearing horizon is earliest Piacenzian, MPL4a foraminiferal zone of
|
||
<bibRefCitation id="EFAA2FACFFDDA44BFBCCB33405A8F98D" author="Rio D & Channell JET & Bertoldi R & Poli MS & Vergerio PP & Raffi I & Sprovieri R & Thunell RC" box="[1064,1252,1640,1662]" pageId="24" pageNumber="1277" pagination="1 - 25" refId="ref26475" refString="Rio D, Channell JET, Bertoldi R, Poli MS, Vergerio PP, Raffi I, Sprovieri R, Thunell RC. 1997. Pliocene sapropels in the northern Adriatic area: chronology and paleoenvironmental significance. Palaeogeography, Palaeoclimatology, Palaeoecology 135: 1 - 25." type="journal article" year="1997">
|
||
Rio
|
||
<emphasis id="B94F8E4FFFDDA44BFBB3B33405DEF98E" box="[1111,1170,1640,1661]" italics="true" pageId="24" pageNumber="1277">et al.</emphasis>
|
||
(1997)
|
||
</bibRefCitation>
|
||
(between ~3.6 and 3.2 Mya).
|
||
</paragraph>
|
||
<paragraph id="8B84525DFFDDA44BFCCDB38F04C3F890" blockId="24.[809,1425,1746,1891]" pageId="24" pageNumber="1277">
|
||
<emphasis id="B94F8E4FFFDDA44BFCCDB38F02E6F91B" box="[809,938,1747,1768]" italics="true" pageId="24" pageNumber="1277">Diagnosis:</emphasis>
|
||
Given that in the text it is suggested that ‘
|
||
<taxonomicName id="4C3B29DEFFDDA44BFC89B3AD0558F8F5" authorityName="Anderson" authorityYear="1878" box="[877,1044,1777,1798]" class="Mammalia" family="Balaenopteridae" genus="Balaenoptera" kingdom="Animalia" order="Cetacea" pageId="24" pageNumber="1277" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94F8E4FFFDDA44BFC89B3AD0558F8F5" box="[877,1044,1777,1798]" italics="true" pageId="24" pageNumber="1277">Balaenoptera</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="4C3B29DEFFDDA44BFBC2B3AD043DF8F5" authority="Boessenecker, 2013" authorityName="Boessenecker" authorityYear="2013" box="[1062,1393,1777,1799]" class="Mammalia" family="Balaenopteridae" genus="Marzanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="24" pageNumber="1277" phylum="Chordata" rank="species" species="bertae">
|
||
<emphasis id="B94F8E4FFFDDA44BFBC2B3AD053EF8F5" box="[1062,1138,1777,1798]" italics="true" pageId="24" pageNumber="1277">bertae</emphasis>
|
||
<bibRefCitation id="EFAA2FACFFDDA44BFB99B3AD043DF8F5" author="Boessenecker RW" box="[1149,1393,1777,1799]" pageId="24" pageNumber="1277" pagination="815 - 940" refId="ref23138" refString="Boessenecker RW. 2013. A new marine vertebrate assemblage from the Late Neogene Purisima Formation in Central California, part II: pinnipeds and cetaceans. Geodiversitas 35: 815 - 940." type="journal article" year="2013">Boessenecker, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
is to be assigned to
|
||
<taxonomicName id="4C3B29DEFFDDA44BFC1BB24C05E5F8D6" authorityName="Bisconti & Damarco & Pavia & Sorce & Carnevale" authorityYear="2021" box="[1023,1193,1808,1829]" class="Mammalia" family="Balaenopteridae" genus="Marzanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="24" pageNumber="1277" phylum="Chordata" rank="genus">
|
||
<emphasis id="B94F8E4FFFDDA44BFC1BB24C05E5F8D6" box="[1023,1193,1808,1829]" italics="true" pageId="24" pageNumber="1277">Marzanoptera</emphasis>
|
||
</taxonomicName>
|
||
, a diagnosis of the new species is provided here. In proportion to the length of the supraoccipital (from the superior
|
||
</paragraph>
|
||
<paragraph id="8B84525DFFDCA44AFF47B59900F0FE41" blockId="25.[163,778,197,434]" pageId="25" pageNumber="1278">
|
||
border of the foramen magnum to the anterior border),
|
||
<taxonomicName id="4C3B29DEFFDCA44AFEF8B5B9031BFF09" authorityName="Bisconti & Damarco & Pavia & Sorce & Carnevale" authorityYear="2021" box="[284,599,228,250]" class="Mammalia" family="Balaenopteridae" genus="Marzanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="25" pageNumber="1278" phylum="Chordata" rank="species" species="tersillae">
|
||
<emphasis id="B94F8E4FFFDCA44AFEF8B5B9031BFF09" box="[284,599,228,250]" italics="true" pageId="25" pageNumber="1278">Marzanoptera tersillae</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C3B29DEFFDCA44AFF47B45F00ECFEE4" authorityName="Boessenecker" authorityYear="2013" box="[163,416,258,280]" class="Mammalia" family="Balaenopteridae" genus="Marzanoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cetacea" pageId="25" pageNumber="1278" phylum="Chordata" rank="species" species="bertae">
|
||
<emphasis id="B94F8E4FFFDCA44AFF47B45F00ECFEE4" box="[163,416,258,280]" italics="true" pageId="25" pageNumber="1278">Marzanoptera bertae</emphasis>
|
||
</taxonomicName>
|
||
in having transversely wider intertemporal region of the frontal, transversely s h o r t e r s u p r a o r b i t a l p r o c e s s e s o f t h e f r o n t a l, slightly shorter and stockier zygomatic process of the squamosal, and dorsal surface of the skull less rounded in lateral view.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |