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<document id="12F7A53195206C50CD7EF2BB4D4B4F71" ID-DOI="10.5281/zenodo.182511" ID-GBIF-Dataset="6b9264fd-88d3-4865-9d3b-9654ecd088d5" ID-ISSN="1175-5326" ID-Zenodo-Dep="182511" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459868260011" checkinUser="plazi" docAuthor="Tuf, Ivan Hadrián, Wytwer, Jolanta &amp; Tajovský, Karel" docDate="2008" docId="0392AA42FFCEFFE934C4F9BD51AC81F1" docLanguage="en" docName="zt01788p046.pdf" docOrigin="Zootaxa 1788" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Lithobius erythrocephalus subsp. mohelensis Dobroruka 1959" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="FFABD23AFFCCFFEA3453FFD052038363" masterDocTitle="On the identity of the species described in the genus Lithobius Leach, 1814 by L. J. Dobroruka from the former Czechoslovakia (Czech and Slovak Republics) (Chilopoda: Lithobiomorpha)" masterLastPageNumber="46" masterPageNumber="37" pageNumber="39" updateTime="1698226387028" updateUser="plazi">
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<mods:title id="A3B13D89F3A8617B9F81952E4D376734">On the identity of the species described in the genus Lithobius Leach, 1814 by L. J. Dobroruka from the former Czechoslovakia (Czech and Slovak Republics) (Chilopoda: Lithobiomorpha)</mods:title>
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<mods:namePart id="170E25E8FA9A4BC82EC057445130F8B1">Tuf, Ivan Hadrián</mods:namePart>
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<mods:namePart id="117BF2464B32BE910BEB9BFA0D791E26">Wytwer, Jolanta</mods:namePart>
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<mods:namePart id="B760D8C00CFAE07151259B152D07F754">Tajovský, Karel</mods:namePart>
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<mods:date id="34580B0AF56674B9019CC9761AD56F46">2008</mods:date>
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<treatment id="0392AA42FFCEFFE934C4F9BD51AC81F1" ID-DOI="http://doi.org/10.5281/zenodo.6229875" ID-GBIF-Taxon="119369019" ID-Zenodo-Dep="6229875" LSID="urn:lsid:plazi:treatment:0392AA42FFCEFFE934C4F9BD51AC81F1" httpUri="http://treatment.plazi.org/id/0392AA42FFCEFFE934C4F9BD51AC81F1" lastPageId="3" lastPageNumber="40" pageId="2" pageNumber="39">
<subSubSection id="C32148DFFFCEFFE834C4F9BD514385E4" box="[151,832,1645,1671]" pageId="2" pageNumber="39" type="nomenclature">
<paragraph id="8B841B54FFCEFFE834C4F9BD514385E4" blockId="2.[151,832,1645,1671]" box="[151,832,1645,1671]" pageId="2" pageNumber="39">
<heading id="D0CCAC38FFCEFFE834C4F9BD514385E4" bold="true" box="[151,832,1645,1671]" fontSize="11" level="1" pageId="2" pageNumber="39" reason="1">
<taxonomicName id="4C3B60D7FFCEFFE834C4F9BD514385E4" ID-CoL="7JY58" authority="Dobroruka, 1959" authorityName="Dobroruka" authorityYear="1959" box="[151,832,1645,1671]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="erythrocephalus" subSpecies="mohelensis">
<emphasis id="B94FC746FFCEFFE834C4F9BD514385E4" bold="true" box="[151,832,1645,1671]" pageId="2" pageNumber="39">
<emphasis id="B94FC746FFCEFFE834C4F9BD506085E4" bold="true" box="[151,611,1645,1671]" italics="true" pageId="2" pageNumber="39">Lithobius erythrocephalus mohelensis</emphasis>
<bibRefCitation id="EFAA66A5FFCEFFE83638F9BD514385E4" author="Dobroruka" box="[619,832,1645,1671]" pageId="2" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka, 1959</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C32148DFFFCEFFE834C4F96851B385AC" box="[151,944,1720,1743]" pageId="2" pageNumber="39" type="reference_group">
<paragraph id="8B841B54FFCEFFE834C4F96851B385AC" blockId="2.[151,944,1720,1743]" box="[151,944,1720,1743]" pageId="2" pageNumber="39">
<treatmentCitationGroup id="AB2B3C7AFFCEFFE834C4F96851B385AC" box="[151,944,1720,1743]" pageId="2" pageNumber="39">
<taxonomicName id="4C3B60D7FFCEFFE834C4F968511985AC" ID-CoL="7JY58" authority="Dobroruka 1959: 105" authorityName="Dobroruka" authorityPageNumber="105" authorityYear="1959" box="[151,794,1720,1743]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="erythrocephalus" subSpecies="mohelensis">
<emphasis id="B94FC746FFCEFFE834C4F968502F85AC" box="[151,556,1720,1743]" italics="true" pageId="2" pageNumber="39">Lithobius erythrocephalus mohelensis</emphasis>
<treatmentCitation id="0A9A3D45FFCEFFE83661F968511985AC" author="Dobroruka" box="[562,794,1720,1743]" page="105" pageId="2" pageNumber="39" year="1959">
<bibRefCitation id="EFAA66A5FFCEFFE83661F96850E685AC" author="Dobroruka" box="[562,741,1720,1743]" pageId="2" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka 1959</bibRefCitation>
: 105
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</taxonomicName>
, tab., figs 3, 4
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</paragraph>
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<subSubSection id="C32148DFFFCEFFE834C4F92F56138481" pageId="2" pageNumber="39" type="materials_examined">
<paragraph id="8B841B54FFCEFFE834C4F92F53C08421" blockId="2.[151,1436,1791,2018]" pageId="2" pageNumber="39">
<emphasis id="B94FC746FFCEFFE834C4F92F533E847A" bold="true" box="[151,317,1791,1817]" pageId="2" pageNumber="39">
<typeStatus id="5480A5F6FFCEFFE834C4F92F52D7847A" box="[151,212,1791,1817]" pageId="2" pageNumber="39">Type</typeStatus>
locality.
</emphasis>
Mohelenská hadcová step National Nature Reserve (NNR) (= Serpentine Steppe of Mohelno), Moravia,
<collectingCountry id="F32C5BC4FFCEFFE8355AF8F853C08421" box="[265,451,1832,1858]" name="Czech Republic" pageId="2" pageNumber="39">Czech Republic</collectingCountry>
</paragraph>
<paragraph id="8B841B54FFCEFFE83495F89F535B84F1" blockId="2.[151,1436,1791,2018]" pageId="2" pageNumber="39">
<emphasis id="B94FC746FFCEFFE83495F89F5390840A" bold="true" box="[198,403,1871,1897]" pageId="2" pageNumber="39">
<typeStatus id="5480A5F6FFCEFFE83495F89F5300840A" box="[198,259,1871,1897]" pageId="2" pageNumber="39">Type</typeStatus>
specimens.
</emphasis>
The original description of
<taxonomicName id="4C3B60D7FFCEFFE836B1F88051A1840A" box="[738,930,1872,1897]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="2" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="erythrocephalus" subSpecies="mohelensis">
<emphasis id="B94FC746FFCEFFE836B1F88051A1840A" box="[738,930,1872,1897]" italics="true" pageId="2" pageNumber="39">L. e. mohelensis</emphasis>
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was based on 48 individuals (29 ɗ, 19 Ψ), coll. Dobroruka.
</paragraph>
<paragraph id="8B841B54FFCEFFE83495F84F56138481" blockId="2.[151,1436,1791,2018]" pageId="2" pageNumber="39">
<emphasis id="B94FC746FFCEFFE83495F84F53B884DA" bold="true" box="[198,443,1951,1977]" pageId="2" pageNumber="39">Material examined.</emphasis>
2 ɗ and 1 Ψ with the rest of a label reading “
<emphasis id="B94FC746FFCEFFE8378AF870565F84DA" box="[985,1116,1952,1977]" italics="true" pageId="2" pageNumber="39">mohelensis</emphasis>
n. spp.” from Dobrorukas “
<typeStatus id="5480A5F6FFCEFFE834F2F81852D58481" box="[161,214,1992,2018]" pageId="2" pageNumber="39">type</typeStatus>
collection”. The material was dried up and mildewed, now in alcohol.
</paragraph>
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<subSubSection id="C32148DFFFCFFFE93495FF4751AC81F1" pageId="3" pageNumber="40" type="discussion">
<paragraph id="8B841B54FFCFFFE93495FF4753CF8249" blockId="3.[151,1436,151,658]" pageId="3" pageNumber="40">
<emphasis id="B94FC746FFCFFFE93495FF47534383D2" bold="true" box="[198,320,151,177]" pageId="3" pageNumber="40">Remarks.</emphasis>
This taxon was described in Dobrorukas paper on
<taxonomicName id="4C3B60D7FFCFFFE937CEFF48536D83B9" authority="C.L. Koch, 1847 (Dobroruka 1962)" authorityName="C.L. Koch, 1847 (Dobroruka" authorityYear="1962" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="erythrocephalus">
<emphasis id="B94FC746FFCFFFE937CEFF4856D383D2" box="[925,1232,152,177]" italics="true" pageId="3" pageNumber="40">Lithobius erythrocephalus</emphasis>
<bibRefCitation id="EFAA66A5FFCFFFE93084FF48579883D1" author="Koch" box="[1239,1435,152,178]" pageId="3" pageNumber="46" refString="Koch, C. L. (1847) System der Myriapoden. In: Panzer, G. &amp; Herrich-Schaffer, L. (Eds.), Kritische Revision der Insectenfauna Deutschlands 3. Friedrich Pustet, Regensburg, pp. 1 - 270." type="journal article" year="1847">C.L. Koch, 1847</bibRefCitation>
(
<bibRefCitation id="EFAA66A5FFCFFFE934CDFF10536583B9" author="Dobroruka" box="[158,358,192,218]" pageId="3" pageNumber="45" refString="Dobroruka, L. J. (1962) Uber Lithobius erythrocephalus C. Koch 1847 (Chilopoda). Zoologischer Anzeiger, 168, 43 - 45." type="journal article" year="1962">Dobroruka 1962</bibRefCitation>
)
</taxonomicName>
and it has never been recorded since then. The differences from the nominal subspecies were stated as lying in the larger size (
<quantity id="4CC3B6B1FFCFFFE93635FF3850EF8261" box="[614,748,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="3" pageNumber="40" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">1922 mm</quantity>
), the weak sulci on the 14th and 15th male tibiae and the very smooth female spurs.
</paragraph>
<paragraph id="8B841B54FFCFFFE93495FEE8563B8121" blockId="3.[151,1436,151,658]" pageId="3" pageNumber="40">
The re-examined material was badly damaged, as almost all the legs were broken off. However, part of the 15th male leg was recognized and a very shallow sulcus was evident on the tibia. The length of these specimens is only
<quantity id="4CC3B6B1FFCFFFE93562FE5853B182C1" box="[305,434,392,418]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="3" pageNumber="40" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">1215 mm</quantity>
, yet the body may have become contracted as a result of drying up. In
<taxonomicName id="4C3B60D7FFCFFFE930B4FE5853A582A9" authority="C.L. Koch, 1847" authorityName="C.L. Koch" authorityYear="1847" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="erythrocephalus">
<emphasis id="B94FC746FFCFFFE930B4FE5852CB82AA" italics="true" pageId="3" pageNumber="40">L. erythrocephalus</emphasis>
<bibRefCitation id="EFAA66A5FFCFFFE93487FE6053A582A9" author="Koch" box="[212,422,432,458]" pageId="3" pageNumber="46" refString="Koch, C. L. (1847) System der Myriapoden. In: Panzer, G. &amp; Herrich-Schaffer, L. (Eds.), Kritische Revision der Insectenfauna Deutschlands 3. Friedrich Pustet, Regensburg, pp. 1 - 270." type="journal article" year="1847">C.L. Koch, 1847</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFAA66A5FFCFFFE935E4FE60505A82A9" author="Eason" box="[439,601,432,458]" pageId="3" pageNumber="45" refString="Eason, E. H. (1972) The type specimens and identity of the species described in the genus Lithobius by C. L. Koch and L. Koch from 1841 to 1878 (Chilopoda: Lithobiomorpha). Bulletin of the British Museum (Natural History), Zoology, 22, 105 - 150." type="journal article" year="1972">Eason (1972)</bibRefCitation>
mentioned intrapopulation variability of the male secondary sexual characters on both last pairs of legs. In addition, there are no differences between the shape of female spurs on the gonopods as depicted by
<bibRefCitation id="EFAA66A5FFCFFFE935B4FDD050F68179" author="Dobroruka" box="[487,757,512,538]" pageId="3" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka (1959: 105)</bibRefCitation>
and
<bibRefCitation id="EFAA66A5FFCFFFE9377EFDD056048179" author="Eason" box="[813,1031,512,538]" pageId="3" pageNumber="45" refString="Eason, E. H. (1972) The type specimens and identity of the species described in the genus Lithobius by C. L. Koch and L. Koch from 1841 to 1878 (Chilopoda: Lithobiomorpha). Bulletin of the British Museum (Natural History), Zoology, 22, 105 - 150." type="journal article" year="1972">Eason (1972: 143)</bibRefCitation>
.
<bibRefCitation id="EFAA66A5FFCFFFE93041FDD056AD8179" author="Eason" box="[1042,1198,512,538]" pageId="3" pageNumber="45" refString="Eason, E. H. (1972) The type specimens and identity of the species described in the genus Lithobius by C. L. Koch and L. Koch from 1841 to 1878 (Chilopoda: Lithobiomorpha). Bulletin of the British Museum (Natural History), Zoology, 22, 105 - 150." type="journal article" year="1972">Eason (1972)</bibRefCitation>
noted that the shape of the tibia of male legs is not a valid character for establishing any subspecies.
</paragraph>
<paragraph id="8B841B54FFCFFFE93495FD9F51AC81F1" blockId="3.[151,1436,151,658]" pageId="3" pageNumber="40">
<emphasis id="B94FC746FFCFFFE93495FD9F5397810A" bold="true" box="[198,404,591,617]" pageId="3" pageNumber="40">Taxonomic note.</emphasis>
We consider
<taxonomicName id="4C3B60D7FFCFFFE93660FD8056C38109" authority="Dobroruka, 1959" authorityName="Dobroruka" authorityYear="1959" box="[563,1216,592,618]" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" order="Lithobiomorpha" pageId="3" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="erythrocephalus" subSpecies="mohelensis">
<emphasis id="B94FC746FFCFFFE93660FD8051EC810A" box="[563,1007,592,617]" italics="true" pageId="3" pageNumber="40">Lithobius erythrocephalus mohelensis</emphasis>
<bibRefCitation id="EFAA66A5FFCFFFE937A4FD8056C38109" author="Dobroruka" box="[1015,1216,592,618]" pageId="3" pageNumber="45" refString="Dobroruka, L. J. (1959) Chilopoda statni prirodni reservace Mohelno. [Chilopoden des staatlichen Naturschutzgebeites Mohelno.] Ochrana p r irody, 14, 104 - 106. (in Czech, with German summary)" type="journal article" year="1959">Dobroruka, 1959</bibRefCitation>
</taxonomicName>
as a junior subjective synonym of
<taxonomicName id="4C3B60D7FFCFFFE9350FFDA8515581F1" authority="C.L. Koch, 1847" authorityName="C.L. Koch" authorityYear="1847" box="[348,854,632,658]" genus="Lithobus" pageId="3" pageNumber="40" rank="species" species="erythrocephalus">
<emphasis id="B94FC746FFCFFFE9350FFDA8508B81F2" box="[348,648,632,657]" italics="true" pageId="3" pageNumber="40">Lithobus erythrocephalus</emphasis>
<bibRefCitation id="EFAA66A5FFCFFFE936DCFDA8515581F1" author="Koch" box="[655,854,632,658]" pageId="3" pageNumber="46" refString="Koch, C. L. (1847) System der Myriapoden. In: Panzer, G. &amp; Herrich-Schaffer, L. (Eds.), Kritische Revision der Insectenfauna Deutschlands 3. Friedrich Pustet, Regensburg, pp. 1 - 270." type="journal article" year="1847">C.L. Koch, 1847</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A27C7A3DFFCFFFE93731FDA851AC81F1" box="[866,943,632,658]" pageId="3" pageNumber="40">syn. n.</taxonomicNameLabel>
</paragraph>
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</treatment>
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