288 lines
32 KiB
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288 lines
32 KiB
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<document id="81FFD903E3D8B259E1688FDE000DFD4A" ID-CLB-Dataset="46097" ID-DOI="10.11646/zootaxa.3784.4.4" ID-GBIF-Dataset="16f488d9-eb77-4ff5-a7ce-99eeb7875371" ID-ISSN="1175-5326" ID-Zenodo-Dep="252218" ID-ZooBank="D5E03502-7BD3-41F4-A4CF-5537B1462A23" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460404217522" checkinUser="plazi" docAuthor="Makarkin, Vladimir N. & Archibald, S. Bruce" docDate="2014" docId="039287A3FE0B371D23C76F21FAD0FD9A" docLanguage="en" docName="zt03784p444.pdf" docOrigin="Zootaxa 3784 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaraphidia elegans Cockerell 1907" docType="treatment" docVersion="9" lastPageNumber="416" masterDocId="FFABFFDBFE06371223506A3EFFE0FF82" masterDocTitle="A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha" masterLastPageNumber="444" masterPageNumber="401" pageNumber="414" updateTime="1698299590966" updateUser="ExternalLinkService">
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<mods:title id="087A13C900740B03DC4E7187E0E2D401">A revision of the late Eocene snakeflies (Raphidioptera) of the Florissant Formation, Colorado, with special reference to the wing venation of the Raphidiomorpha</mods:title>
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<mods:namePart id="3955EFCA482761BC38CEEE0DA4C38A68">Archibald, S. Bruce</mods:namePart>
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<mods:date id="C74829441B158D90B01D632AC5BA7A2D">2014</mods:date>
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<treatment id="039287A3FE0B371D23C76F21FAD0FD9A" ID-DOI="http://doi.org/10.5281/zenodo.6131151" ID-GBIF-Taxon="119420885" ID-Zenodo-Dep="6131151" LSID="urn:lsid:plazi:treatment:039287A3FE0B371D23C76F21FAD0FD9A" httpUri="http://treatment.plazi.org/id/039287A3FE0B371D23C76F21FAD0FD9A" lastPageId="15" lastPageNumber="416" pageId="13" pageNumber="414">
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<subSubSection id="C321653EFE0B371F23C76F21FF0BFAD9" pageId="13" pageNumber="414" type="nomenclature">
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<paragraph id="8B8436B5FE0B371F23C76F21FD8DFAB8" blockId="13.[151,621,1311,1371]" box="[151,621,1311,1338]" pageId="13" pageNumber="414">
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<heading id="D0CC81D9FE0B371F23C76F21FD8DFAB8" bold="true" box="[151,621,1311,1338]" fontSize="11" level="1" pageId="13" pageNumber="414" reason="1">
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<taxonomicName id="4C3B4D36FE0B371F23C76F21FD8DFAB8" ID-CoL="3YWZH" authority="Cockerell, 1907" authorityName="Cockerell" authorityYear="1907" box="[151,621,1311,1338]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="B94FEAA7FE0B371F23C76F21FD8DFAB8" bold="true" box="[151,621,1311,1338]" pageId="13" pageNumber="414">
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<emphasis id="B94FEAA7FE0B371F23C76F21FE44FABB" bold="true" box="[151,420,1311,1337]" italics="true" pageId="13" pageNumber="414">Megaraphidia elegans</emphasis>
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<bibRefCitation id="EFAA4B44FE0B371F22FC6F1EFD8DFAB8" author="Cockerell" box="[428,621,1312,1338]" pageId="13" pageNumber="414" refString="Cockerell, T. D. A. (1907) Some fossil arthropods from Florissant, Colorado. Bulletin of the American Museum of Natural History, 23, 605 - 616." type="journal article" year="1907">Cockerell, 1907</bibRefCitation>
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<paragraph id="8B8436B5FE0B371F23C76F7CFF0BFAD9" blockId="13.[151,621,1311,1371]" box="[151,235,1346,1371]" pageId="13" pageNumber="414">
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(
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<figureCitation id="13002A30FE0B371F23CF6F7CFF02FAD9" box="[159,226,1346,1371]" captionStart="FIGURE 5" captionStartId="14.[151,249,1944,1966]" captionTargetBox="[151,1436,553,1920]" captionTargetId="figure@14.[151,1436,553,1923]" captionTargetPageId="14" captionText="FIGURE 5. Megaraphidia elegans Cockerell, 1907, holotype. A, part UCM 8602. B, counterpart AMNH 35861 (converted to right). C, forewing venation. Scale bar = 4 mm." httpUri="https://zenodo.org/record/252223/files/figure.png" pageId="13" pageNumber="414">Fig. 5</figureCitation>
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)
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</paragraph>
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<subSubSection id="C321653EFE0B371F23C76FB4FDF0F9B8" pageId="13" pageNumber="414" type="reference_group">
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<paragraph id="8B8436B5FE0B371F23C76FB4FE82FA5C" blockId="13.[151,1436,1418,1594]" pageId="13" pageNumber="414">
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<taxonomicName id="4C3B4D36FE0B371F23C76FB4FD86FA22" ID-CoL="3YWZH" authority="Cockerell, 1907: 607" authorityName="Cockerell" authorityPageNumber="607" authorityYear="1907" box="[151,614,1418,1440]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="B94FEAA7FE0B371F23C76FB4FE9EFA22" box="[151,382,1418,1440]" italics="true" pageId="13" pageNumber="414">Megaraphidia elegans</emphasis>
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<treatmentCitation id="0A9A10A4FE0B371F22D86FB4FD86FA22" author="Cockerell" box="[392,614,1418,1440]" page="607" pageId="13" pageNumber="414" year="1907">
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<bibRefCitation id="EFAA4B44FE0B371F22D86FB4FDD0FA22" author="Cockerell" box="[392,560,1418,1440]" pageId="13" pageNumber="414" refString="Cockerell, T. D. A. (1907) Some fossil arthropods from Florissant, Colorado. Bulletin of the American Museum of Natural History, 23, 605 - 616." type="journal article" year="1907">Cockerell, 1907</bibRefCitation>
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: 607
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</taxonomicName>
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,
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<figureCitation id="13002A30FE0B371F21236FB4FD53FA22" box="[627,691,1418,1440]" captionStart="FIGURE 3" captionStartId="10.[151,249,1760,1782]" captionTargetBox="[151,1436,783,1726]" captionTargetId="figure@10.[151,1436,783,1738]" captionTargetPageId="10" captionText="FIGURE 3. Dictyoraphidia veterana (Scudder, 1890). Holotype PU 6385, EM 1.385. A, the specimen as preserved today. B, forewing. C, hind wing. FW, forewing; HW, hind wing. Scale bars = 4 mm (B, C to scale)." httpUri="https://zenodo.org/record/252221/files/figure.png" pageId="13" pageNumber="414">Fig. 3</figureCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F21936FB4FC7AFA22" author="Cockerell" box="[707,922,1418,1440]" page="78" pageId="13" pageNumber="414" year="1909">
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<bibRefCitation id="EFAA4B44FE0B371F21936FB4FC92FA22" author="Cockerell" box="[707,882,1418,1440]" pageId="13" pageNumber="414" refString="Cockerell, T. D. A. (1909 b) A catalogue of the generic names based on American insects and arachnids from the tertiary rocks, with indications of the type species. Bulletin of the American Museum of Natural History, 26, 77 - 86." type="journal article" year="1909" yearSuffix="b">Cockerell 1909b</bibRefCitation>
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: 78
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F20F86FB4FB8EFA22" author="Rohwer" box="[936,1134,1418,1440]" page="534" pageId="13" pageNumber="414" year="1909">
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<bibRefCitation id="EFAA4B44FE0B371F20F86FB4FBD8FA22" author="Rohwer" box="[936,1080,1418,1440]" pageId="13" pageNumber="414" refString="Rohwer, S. A. (1909) Three new fossil insects from Florissant, Colorado. American Journal of Science, 28, 533 - 536. http: // dx. doi. org / 10.2475 / ajs. s 4 - 28.168.533" type="journal article" year="1909">Rohwer 1909</bibRefCitation>
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: 534
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F272C6FB4FA80FA22" author="Handlirsch" box="[1148,1376,1418,1440]" page="103" pageId="13" pageNumber="414" year="1910">
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<bibRefCitation id="EFAA4B44FE0B371F272C6FB4FACCFA22" author="Handlirsch" box="[1148,1324,1418,1440]" pageId="13" pageNumber="414" refString="Handlirsch, A. (1910) Canadian fossil insects. 5. Insects from the Tertiary lake deposits of the Southern Interior of British Columbia collected by Mr. Lawrence Lambe, in 1906. In: Contributions to Canadian Palaeontology. Vol. II. Part III. Memoirs of the Geological Survey of Canada, 12 - P, pp. 93 - 129." type="book chapter" year="1910">Handlirsch 1910</bibRefCitation>
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: 103
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</treatmentCitation>
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, 104;
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<treatmentCitation id="0A9A10A4FE0B371F23976F97FE7BFA3D" author="Cockerell" box="[199,411,1449,1471]" page="216" pageId="13" pageNumber="414" year="1912">
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<bibRefCitation id="EFAA4B44FE0B371F23976F97FE87FA3D" author="Cockerell" box="[199,359,1449,1471]" pageId="13" pageNumber="414" refString="Cockerell, T. D. A. (1912) A fossil Raphidia (Neur. Planip.). Entomological News, 23, 215 - 216." type="journal article" year="1912">Cockerell 1912</bibRefCitation>
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: 216
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F22F66F97FD66FA3D" author="Handlirsch" box="[422,646,1449,1471]" page="812" pageId="13" pageNumber="414" year="1913">
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<bibRefCitation id="EFAA4B44FE0B371F22F66F97FDB2FA3D" author="Handlirsch" box="[422,594,1449,1471]" pageId="13" pageNumber="414" refString="Handlirsch, A. (1913) Subphylum C. Insecta (Hexapoda). Insects. In: Zittel, K. A. von (Ed.), Text-book of paleontology, edited by Charles R. Eastman. Vol. 1. 2 nd Edition. Macmillan and Co. Limited, London, pp. 794 - 822." type="journal article" year="1913">Handlirsch 1913</bibRefCitation>
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: 812
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F21C16F97FC29FA3D" author="Cockerell" box="[657,969,1449,1471]" page="295" pageId="13" pageNumber="414" year="1925">
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<bibRefCitation id="EFAA4B44FE0B371F21C16F97FC75FA3D" author="Cockerell" box="[657,917,1449,1471]" pageId="13" pageNumber="414" refString="Cockerell, T. D. A. & Custer, C. (1925) A new fossil Inocellia (Neuroptera) from Florissant. The Entomologist, 58, 295 - 297." type="journal article" year="1925">Cockerell & Custer 1925</bibRefCitation>
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: 295
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F20856F97FB73FA3D" author="Rodeck" box="[981,1171,1449,1471]" page="290" pageId="13" pageNumber="414" year="1938">
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<bibRefCitation id="EFAA4B44FE0B371F20856F97FBBFFA3D" author="Rodeck" box="[981,1119,1449,1471]" pageId="13" pageNumber="414" refString="Rodeck, H. G. (1938) Type specimens of fossils in the University of Colorado Museum. The University of Colorado Studies, A, 25, 281 - 304." type="journal article" year="1938">Rodeck 1938</bibRefCitation>
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: 290
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F27CF6F97FA78FA3D" author="Aspock" box="[1183,1432,1449,1471]" page="535" pageId="13" pageNumber="414" year="1991">
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<bibRefCitation id="EFAA4B44FE0B371F27CF6F97FA85FA3D" author="Aspock" box="[1183,1381,1449,1471]" pageId="13" pageNumber="414" refString="Aspock, H., Aspock, U. & Rausch, H. (1991) Die Raphidiopteren der Erde. Eine monographische Darstellung der Systematik, Taxonomie, Biologie, Okologie und Chorologie der rezenten Raphidiopteren der Erde, mit einer zusammenfassenden Ubersicht der fossilen Raphidiopteren (Insecta: Neuropteroidea). 2 Vols. Goecke & Evers, Krefeld, 730 & 550 pp." type="book" year="1991">
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Aspöck
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<emphasis id="B94FEAA7FE0B371F27A46F94FAC8FA3D" box="[1268,1320,1449,1471]" italics="true" pageId="13" pageNumber="414">et al.</emphasis>
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1991
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</bibRefCitation>
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: 535
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</treatmentCitation>
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, 665, Fig. 3046.
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</paragraph>
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<paragraph id="8B8436B5FE0B371F23C76FD8FEC4F999" blockId="13.[151,1436,1418,1594]" pageId="13" pageNumber="414">
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<taxonomicName id="4C3B4D36FE0B371F23C76FD8FDCAFA7E" ID-CoL="794V2" authority="Carpenter 1936: 145" authorityName="Carpenter" authorityPageNumber="145" authorityYear="1936" box="[151,554,1510,1532]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="elegans">
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<emphasis id="B94FEAA7FE0B371F23C76FD8FEA8FA7E" box="[151,328,1510,1532]" italics="true" pageId="13" pageNumber="414">Raphidia elegans</emphasis>
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:
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<treatmentCitation id="0A9A10A4FE0B371F22066FD8FDCAFA7E" author="Carpenter" box="[342,554,1510,1532]" page="145" pageId="13" pageNumber="414" year="1936">
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<bibRefCitation id="EFAA4B44FE0B371F22066FD8FE17FA7E" author="Carpenter" box="[342,503,1510,1532]" pageId="13" pageNumber="414" refString="Carpenter, F. M. (1936) Revision of the Nearctic Raphidiodea (recent and fossil). Proceedings of the American Academy of Arts and Science, 71, 89 - 157, 2 pls." type="journal article" year="1936">Carpenter 1936</bibRefCitation>
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: 145
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, 146;
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<treatmentCitation id="0A9A10A4FE0B371F21396FD8FCC8FA7E" author="Oswald" box="[617,808,1510,1532]" page="160" pageId="13" pageNumber="414" year="1990">
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<bibRefCitation id="EFAA4B44FE0B371F21396FD8FD15FA7E" author="Oswald" box="[617,757,1510,1532]" pageId="13" pageNumber="414" refString="Oswald, J. D. (1990) Raphidioptera. In: Grimaldi, D. A. (Ed.), Insects from the Santana Formation, Lower Cretaceous, of Brazil. Bulletin of the American Museum of Natural History, 195, pp. 154 - 163." type="journal article" year="1990">Oswald 1990</bibRefCitation>
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: 160
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F20636FD8FC3FFA7E" author="Engel" box="[819,991,1510,1532]" page="125" pageId="13" pageNumber="414" year="2003">
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<bibRefCitation id="EFAA4B44FE0B371F20636FD8FC4BFA7E" author="Engel" box="[819,939,1510,1532]" pageId="13" pageNumber="414" refString="Engel, M. S. (2003) A new Eocene-Oligocene snakefly from Florissant, Colorado (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Sciences, 106, 124 - 128. http: // dx. doi. org / 10.1660 / 0022 - 8443 (2003) 106 [0124: anesff] 2.0. co; 2" type="journal article" year="2003">Engel 2003</bibRefCitation>
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: 125
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</treatmentCitation>
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;
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<treatmentCitation id="0A9A10A4FE0B371F20BA6FD8FB73FA7E" author="Engel" box="[1002,1171,1510,1532]" page="213" pageId="13" pageNumber="414" year="2009">
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<bibRefCitation id="EFAA4B44FE0B371F20BA6FD8FB82FA7E" author="Engel" box="[1002,1122,1510,1532]" pageId="13" pageNumber="414" refString="Engel, M. S. (2009) A Miocene snakefly from Stewart Valley, Nevada (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 112, 211 - 214. http: // dx. doi. org / 10.1660 / 062.112.0409" type="journal article" year="2009">Engel 2009</bibRefCitation>
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: 213
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(Tabl. 1);
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<treatmentCitation id="0A9A10A4FE0B371F26506FD8FA7BFA7E" author="Engel" box="[1280,1435,1510,1532]" page="79" pageId="13" pageNumber="414" year="2011">
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<bibRefCitation id="EFAA4B44FE0B371F26506FD8FA98FA7E" author="Engel" box="[1280,1400,1510,1532]" pageId="13" pageNumber="414" refString="Engel, M. S. (2011) A new snakefly from the Eocene Green River Formation (Raphidioptera: Raphidiidae). Transactions of the Kansas Academy of Science, 114, 77 - 87. http: // dx. doi. org / 10.1660 / 062.114.0107" type="journal article" year="2011">Engel 2011</bibRefCitation>
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: 79
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(Tabl. 2).
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</paragraph>
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<paragraph id="8B8436B5FE0B371F23C76C1AFDF0F9B8" blockId="13.[151,1436,1418,1594]" box="[151,528,1572,1594]" pageId="13" pageNumber="414">
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<treatmentCitationGroup id="AB2B119BFE0B371F23C76C1AFDF0F9B8" box="[151,528,1572,1594]" pageId="13" pageNumber="414">
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“
|
|
<emphasis id="B94FEAA7FE0B371F23F26C1AFE80F9B8" box="[162,352,1572,1594]" italics="true" pageId="13" pageNumber="414">
|
|
<taxonomicName id="4C3B4D36FE0B371F23F26C1AFEE6F9B8" ID-CoL="775X" box="[162,262,1572,1594]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="genus">Raphidia</taxonomicName>
|
|
”
|
|
<taxonomicName id="4C3B4D36FE0B371F22436C1AFE80F9B8" ID-CoL="794V2" authority="Engel 2002: 22" authorityName="Engel" authorityPageNumber="22" authorityYear="2002" box="[275,352,1572,1594]" class="Insecta" family="Raphidiidae" genus="Raphidia" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
|
</emphasis>
|
|
:
|
|
<treatmentCitation id="0A9A10A4FE0B371F223E6C1AFDEBF9B8" author="Engel" box="[366,523,1572,1594]" page="22" pageId="13" pageNumber="414" year="2002">
|
|
<bibRefCitation id="EFAA4B44FE0B371F223E6C1AFE05F9B8" author="Engel" box="[366,485,1572,1594]" pageId="13" pageNumber="414" refString="Engel, M. S. (2002) The smallest snakefly (Raphidioptera: Mesoraphidiidae): a new species in Cretaceous amber from Myanmar, with a catalog of fossil snakeflies. American Museum Novitates, 3363, 1 - 22. http: // dx. doi. org / 10.1206 / 0003 - 0082 (2002) 363 <0001: tssrma> 2.0. co; 2" type="journal article" year="2002">Engel 2002</bibRefCitation>
|
|
: 22
|
|
</treatmentCitation>
|
|
.
|
|
</treatmentCitationGroup>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C321653EFE0B371C23C76C58FC7FFE76" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414" type="diagnosis">
|
|
<paragraph id="8B8436B5FE0B371F23C76C58FBD1F920" blockId="13.[151,1436,1638,2023]" pageId="13" pageNumber="414">
|
|
<emphasis id="B94FEAA7FE0B371F23C76C58FEF1F9FD" bold="true" box="[151,273,1638,1663]" pageId="13" pageNumber="414">Diagnosis.</emphasis>
|
|
May be distinguished from
|
|
<taxonomicName id="4C3B4D36FE0B371F21046C59FD07F9FD" box="[596,743,1638,1663]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="exhumata">
|
|
<emphasis id="B94FEAA7FE0B371F21046C59FD07F9FD" box="[596,743,1638,1663]" italics="true" pageId="13" pageNumber="414">M. exhumata</emphasis>
|
|
</taxonomicName>
|
|
by AA1 touching AA2 [these connecting by crossvein in
|
|
<taxonomicName id="4C3B4D36FE0B371F262D6C59FEE4F921" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="exhumata">
|
|
<emphasis id="B94FEAA7FE0B371F262D6C59FEE4F921" italics="true" pageId="13" pageNumber="414">M. exhumata</emphasis>
|
|
</taxonomicName>
|
|
]; by two branches of CuA before fusion with MP [one in
|
|
<taxonomicName id="4C3B4D36FE0B371F20DE6CB2FBC0F921" box="[910,1056,1674,1699]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="exhumata">
|
|
<emphasis id="B94FEAA7FE0B371F20DE6CB2FBC0F921" box="[910,1056,1674,1699]" italics="true" pageId="13" pageNumber="414">M. exhumata</emphasis>
|
|
</taxonomicName>
|
|
].
|
|
</paragraph>
|
|
<paragraph id="8B8436B5FE0B371C23976C90FC7FFE76" blockId="13.[151,1436,1638,2023]" lastBlockId="14.[151,1437,151,501]" lastPageId="14" lastPageNumber="415" pageId="13" pageNumber="414">
|
|
<emphasis id="B94FEAA7FE0B371F23976C90FE5CF945" bold="true" box="[199,444,1710,1735]" pageId="13" pageNumber="414">Revised description.</emphasis>
|
|
Forewing 14.0 mm long,
|
|
<quantity id="4CC39B50FE0B371F21A46C90FCAEF944" box="[756,846,1710,1735]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="414" unit="mm" value="4.5">4.5 mm</quantity>
|
|
wide (length/width ratio 3.1). Costal space broad, strongly dilated at proximal 1/4 of length; eight preserved subcostal veinlets simple, widely spaced. ScP relatively long, terminating on costal margin at ca. 1/3 of wing length, slightly distad crossvein 2ra-rp; length from termination of ScP to pterostigma shorter than length of pterostigma. Subcostal space broad, with one crossvein located in middle between origin of
|
|
<collectingCountry id="F32C7625FE0B371F21606D01FDB2F8D4" box="[560,594,1855,1878]" name="Poland" pageId="13" pageNumber="414">RP</collectingCountry>
|
|
, divergence of M, CuA; crossvein closing pterostigma proximally straight, slightly inclined to wing base. Pterostigma rather long, weakly pigmented. Four RA branches: one incorporated in pterostigma proximally clearly visible; other at distal end of pterostigma; two distad pterostigma, one long, other short, very shallowly forked. Posterior trace of RA after pterostigma long, smoothly incurved; entering margin before wing apex. RA space with two crossveins forming two radial cells: 2ra-rp located well proximad pterostigma, slightly proximad termination of ScP; 3ra-rp located in distal part of pterostigma.
|
|
<collectingCountry id="F32C7625FE08371C27B36AA6FAE5FF2D" box="[1251,1285,152,175]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
originates at nearly half wing length, with six pectinate branches:
|
|
<collectingCountry id="F32C7625FE08371C21A66A82FCFFFF56" box="[758,799,188,212]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
1,
|
|
<collectingCountry id="F32C7625FE08371C20666A82FCB9FF56" box="[822,857,188,212]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
2 deeply forked;
|
|
<collectingCountry id="F32C7625FE08371C274D6A82FBA3FF56" box="[1053,1091,188,212]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
3,
|
|
<collectingCountry id="F32C7625FE08371C270D6A82FB62FF56" box="[1117,1154,188,212]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
4,
|
|
<collectingCountry id="F32C7625FE08371C27CD6A82FB20FF57" box="[1181,1216,188,213]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
6 shallowly forked;
|
|
<collectingCountry id="F32C7625FE08371C23C76AE1FF5BFF7A" box="[151,187,223,248]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
5 simple. One crossvein in radial space, between
|
|
<collectingCountry id="F32C7625FE08371C21B16ADEFCEAFF7A" box="[737,778,224,248]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
1,
|
|
<collectingCountry id="F32C7625FE08371C204F6ADEFCA5FF7A" box="[799,837,224,248]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
2. Three crossvein between
|
|
<collectingCountry id="F32C7625FE08371C272B6ADEFB7DFF75" box="[1147,1181,224,247]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
, MA. M appears fused with R for considerable distance, and then fused with CuA for short distance; forked well proximad origin of
|
|
<collectingCountry id="F32C7625FE08371C26266B3BFA79FE9E" box="[1398,1433,261,284]" name="Poland" pageId="14" pageNumber="415">RP</collectingCountry>
|
|
. MA deeply dichotomously forked. MP zigzagged, anterior trace deeply forked distally; with three long pectinate branches: two deeply forked, one simple. Two intramedian crossveins form two
|
|
<emphasis id="B94FEAA7FE08371C27086B72FB9CFEE7" box="[1112,1148,332,357]" italics="true" pageId="14" pageNumber="415">doi</emphasis>
|
|
. Cu basally not visible; dividing into CuA, CuP far from wing base. CuA crossvein-like before fusion with M and perpendicular to Cu; after fusion running parallel to CuP, then its anterior trace strongly zigzagged, shallowly forked, with two pectinate long simple branches. CuP simple, continuing Cu. Crossvein between CuA, CuP long. Crossvein cu-aa short. AA1 to AA3 simple. AA1, AA2 touching slightly proximad origin of CuP.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="DF44663DFE08371C23C76DA6FD98F84F" httpUri="https://zenodo.org/record/252223/files/figure.png" pageId="14" pageNumber="415" targetBox="[151,1436,553,1920]" targetPageId="14">
|
|
<paragraph id="8B8436B5FE08371C23C76DA6FD98F84F" blockId="14.[151,1435,1944,1997]" pageId="14" pageNumber="415">
|
|
<emphasis id="B94FEAA7FE08371C23C76DA6FEF2F82F" bold="true" box="[151,274,1944,1966]" pageId="14" pageNumber="415">FIGURE 5.</emphasis>
|
|
<taxonomicName id="4C3B4D36FE08371C22496DA6FD48F82C" authority="Cockerell, 1907" authorityName="Cockerell" authorityYear="1907" box="[281,680,1944,1966]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="14" pageNumber="415" phylum="Arthropoda" rank="species" species="elegans">
|
|
<emphasis id="B94FEAA7FE08371C22496DA6FE1DF82C" box="[281,509,1944,1966]" italics="true" pageId="14" pageNumber="415">Megaraphidia elegans</emphasis>
|
|
Cockerell, 1907
|
|
</taxonomicName>
|
|
, holotype. A, part UCM 8602. B, counterpart AMNH 35861 (converted to right). C, forewing venation. Scale bar = 4 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="C321653EFE09371D23976AA9FC74FE2E" pageId="15" pageNumber="416" type="materials_examined">
|
|
<paragraph id="8B8436B5FE09371D23976AA9FE90FE0A" blockId="15.[151,1437,151,536]" pageId="15" pageNumber="416">
|
|
<emphasis id="B94FEAA7FE09371D23976AA9FE98FF32" bold="true" box="[199,376,151,176]" pageId="15" pageNumber="416">
|
|
<typeStatus id="54808817FE09371D23976AA9FEE2FF32" box="[199,258,151,176]" pageId="15" pageNumber="416">Type</typeStatus>
|
|
material.
|
|
</emphasis>
|
|
<typeStatus id="54808817FE09371D22D36AA9FE0FFF32" box="[387,495,151,176]" pageId="15" pageNumber="416" type="holotype">Holotype</typeStatus>
|
|
: Specimen No. 35861, deposited in
|
|
<collectionCode id="ED2AAE70FE09371D20C86AA6FC0DFF32" LSID="urn:lsid:biocol.org:col:34925" box="[920,1005,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="15" pageNumber="416">AMNH</collectionCode>
|
|
(part), No. 8602, deposited in
|
|
<collectionCode id="ED2AAE70FE09371D260C6AA6FA7CFF32" LSID="urn:lsid:biocol.org:col:34854" box="[1372,1436,152,176]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="15" pageNumber="416">UCM</collectionCode>
|
|
(counterpart), collected by W.P. Cockerell in 1906. A well-preserved nearly complete forewing. The counterpart 8602: Verbatim label data [typed and hand written]: “R / B /
|
|
<collectionCode id="ED2AAE70FE09371D20046ADEFC74FF7A" LSID="urn:lsid:biocol.org:col:34854" box="[852,916,224,248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="15" pageNumber="416">UCM</collectionCode>
|
|
Paleontology
|
|
<collectionCode id="ED2AAE70FE09371D27696ADEFB9FFF75" box="[1081,1151,224,247]" pageId="15" pageNumber="416">
|
|
<typeStatus id="54808817FE09371D27696ADEFB9FFF75" box="[1081,1151,224,247]" pageId="15" pageNumber="416">TYPE</typeStatus>
|
|
</collectionCode>
|
|
Collection /
|
|
<taxonomicName id="4C3B4D36FE09371D26466ADEFA79FF75" box="[1302,1433,224,247]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
|
|
:
|
|
<taxonomicName id="4C3B4D36FE09371D23C76B3AFEFEFE9F" box="[151,286,260,285]" class="Insecta" family="Raphidiidae" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="family">Raphidiidae</taxonomicName>
|
|
/
|
|
<collectionCode id="ED2AAE70FE09371D22636B3AFE96FE9E" LSID="urn:lsid:biocol.org:col:34854" box="[307,374,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="15" pageNumber="416">UCM</collectionCode>
|
|
#: 8602 / Locality #: 83013 Description: wing / ID:
|
|
<taxonomicName id="4C3B4D36FE09371D20E06B3AFAF9FE9F" authority="Cockerell" authorityName="Cockerell" box="[944,1305,260,285]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="elegans">Megaraphidia elegans Cockerell</taxonomicName>
|
|
/ Age: Late Eocene Collector: WP Cockerell 1906 / Fm:
|
|
<collectionCode id="ED2AAE70FE09371D21DD6B16FD31FEC2" box="[653,721,296,320]" pageId="15" pageNumber="416">FLFO</collectionCode>
|
|
[Florissant Formation] Citation: Toohey 55210”. The specimen has black ink writing on the rock that says “
|
|
<taxonomicName id="4C3B4D36FE09371D21D96B72FCC7FEE7" box="[649,807,332,357]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="genus">Megaraphidia</taxonomicName>
|
|
/
|
|
<taxonomicName id="4C3B4D36FE09371D206F6B72FC73FEE7" box="[831,915,332,357]" class="Insecta" family="Raphidiidae" genus="Megaraphidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Raphidioptera" pageId="15" pageNumber="416" phylum="Arthropoda" rank="species" species="elegans">elegans</taxonomicName>
|
|
/ Ckll / Reverse [counterpart] /
|
|
<collectionCode id="ED2AAE70FE09371D27AE6B73FAA4FEE6" box="[1278,1348,333,356]" pageId="15" pageNumber="416">
|
|
<typeStatus id="54808817FE09371D27AE6B73FAA4FEE6" box="[1278,1348,333,356]" pageId="15" pageNumber="416">TYPE</typeStatus>
|
|
</collectionCode>
|
|
/
|
|
<collectionCode id="ED2AAE70FE09371D260C6B72FA7CFEE6" LSID="urn:lsid:biocol.org:col:34854" box="[1372,1436,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="15" pageNumber="416">UCM</collectionCode>
|
|
8602”, and “N 13”.
|
|
</paragraph>
|
|
<paragraph id="8B8436B5FE09371D23976BAAFC74FE2E" blockId="15.[151,1437,151,536]" box="[199,916,404,429]" pageId="15" pageNumber="416">
|
|
<emphasis id="B94FEAA7FE09371D23976BAAFE85FE2F" bold="true" box="[199,357,404,429]" pageId="15" pageNumber="416">
|
|
<typeStatus id="54808817FE09371D23976BAAFEE2FE2F" box="[199,258,404,429]" pageId="15" pageNumber="416">Type</typeStatus>
|
|
locality.
|
|
</emphasis>
|
|
Florissant: Station 13 (
|
|
<collectionCode id="ED2AAE70FE09371D21396BAAFD4BFE2E" LSID="urn:lsid:biocol.org:col:34854" box="[617,683,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34854" name="University of Colorado Museum of Natural History" pageId="15" pageNumber="416">UCM</collectionCode>
|
|
locality No. 83013).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C321653EFE09371D23976B89FAD0FD9A" pageId="15" pageNumber="416" type="discussion">
|
|
<paragraph id="8B8436B5FE09371D23976B89FAD0FD9A" blockId="15.[151,1437,151,536]" pageId="15" pageNumber="416">
|
|
<emphasis id="B94FEAA7FE09371D23976B89FEDBFE52" bold="true" box="[199,315,439,464]" pageId="15" pageNumber="416">Remarks.</emphasis>
|
|
The species was described only from the part, in which the central portion of the forewing is missing. Our description and drawing are based on both the part and counterpart, allowing reconstruction of the complete forewing venation, with which we provide our revised genus diagnosis and species description.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |