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<document id="EE2E9E7328586B4E8F6F908FF62539D6" ID-CLB-Dataset="7365" ID-DOI="10.11646/zootaxa.4947.1.1" ID-GBIF-Dataset="74dc998a-dc7a-4b02-b43c-a2064c9c6dc2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4614567" ID-ZooBank="7F3B4794-65CF-4691-A330-78A423F2606F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1616002389958" checkinUser="plazi" docAuthor="Ebert, David A., Wintner, Sabine P. &amp; Kyne, Peter M." docDate="2021" docId="039287A19538FFDFCFF75F1BFDE927B3" docLanguage="en" docName="zootaxa.4947.1.1.pdf" docOrigin="Zootaxa 4947 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Malacoraja spinacidermis" docType="treatment" docVersion="10" lastPageNumber="82" masterDocId="FFABFFD99569FF8ECF605A0DFFF9206F" masterDocTitle="An annotated checklist of the chondrichthyans of South Africa" masterLastPageNumber="127" masterPageNumber="1" pageNumber="82" updateTime="1698934036276" updateUser="ExternalLinkService">
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<mods:title id="0036351509FF21BBD160E8D11AFEAA88">An annotated checklist of the chondrichthyans of South Africa</mods:title>
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<mods:namePart id="91A62FF6ED66ACBD623FADBE37C9F70C">Ebert, David A.</mods:namePart>
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<mods:namePart id="1CDF1C0AF63F13CFDF031C08D4F2023D">Kyne, Peter M.</mods:namePart>
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<treatment id="039287A19538FFDFCFF75F1BFDE927B3" ID-DOI="http://doi.org/10.5281/zenodo.4697724" ID-GBIF-Taxon="181006947" ID-Zenodo-Dep="4697724" LSID="urn:lsid:plazi:treatment:039287A19538FFDFCFF75F1BFDE927B3" httpUri="http://treatment.plazi.org/id/039287A19538FFDFCFF75F1BFDE927B3" lastPageNumber="82" pageId="81" pageNumber="82">
<subSubSection id="C321653C9538FFDFCFF75F1BFD72255F" box="[151,651,1302,1329]" pageId="81" pageNumber="82" type="nomenclature">
<paragraph id="8B8436B79538FFDFCFF75F1BFD72255F" blockId="81.[151,1437,1302,2013]" box="[151,651,1302,1329]" pageId="81" pageNumber="82">
<heading id="D0CC81DB9538FFDFCFF75F1BFD72255F" bold="true" box="[151,651,1302,1329]" fontSize="11" level="2" pageId="81" pageNumber="82" reason="4">
<taxonomicName id="4C3B4D349538FFDFCFF75F1BFD72255F" ID-CoL="6R4RP" ID-ENA="1090676" authority="(Barnard, 1923)" baseAuthorityName="Barnard" baseAuthorityYear="1923" box="[151,651,1302,1329]" class="Elasmobranchii" family="Rajidae" genus="Malacoraja" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="spinacidermis">
<emphasis id="B94FEAA59538FFDFCFF75F1BFD72255F" bold="true" box="[151,651,1302,1329]" pageId="81" pageNumber="82">
<emphasis id="B94FEAA59538FFDFCFF75F1BFE3D255E" bold="true" box="[151,452,1302,1329]" italics="true" pageId="81" pageNumber="82">Malacoraja spinacidermis</emphasis>
(
<bibRefCitation id="EFAA4B469538FFDFCEB55F1BFD7B255F" author="Barnard" box="[469,642,1302,1328]" firstAuthor="Barnard" pageId="81" pageNumber="82" pagination="439 - 445" refId="ref92395" refString="Barnard, K. H. (1923) Diagnoses of new species of marine fishes from South African waters. Annals of the South African Museum, 13 (Pt. 8), No. 14, 439 - 445." type="journal article" year="1923">Barnard, 1923</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321653C9538FFDFCFF75F37FEAF253B" box="[151,342,1338,1364]" pageId="81" pageNumber="82" type="vernacular_names">
<paragraph id="8B8436B79538FFDFCFF75F37FEAF253B" blockId="81.[151,1437,1302,2013]" box="[151,342,1338,1364]" pageId="81" pageNumber="82">Roughskin Skate</paragraph>
</subSubSection>
<subSubSection id="C321653C9538FFDFCFF75F52FC97276A" pageId="81" pageNumber="82" type="reference_group">
<paragraph id="8B8436B79538FFDFCFF75F52FE6325F2" blockId="81.[151,1437,1302,2013]" pageId="81" pageNumber="82">
<treatmentCitation id="0A9A10A69538FFDFCFF75F52FDAB2517" author="Barnard, K. H." box="[151,594,1374,1400]" page="440" pageId="81" pageNumber="82" year="1923">
<taxonomicName id="4C3B4D349538FFDFCFF75F52FDAB2517" ID-CoL="793L8" authority="Barnard, 1923: 440" authorityName=": Barnard" authorityPageNumber="440" authorityYear="1923" baseAuthorityName="Barnard" baseAuthorityYear="1923" box="[151,594,1374,1400]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="spinacidermis">
<emphasis id="B94FEAA59538FFDFCFF75F52FE892517" box="[151,368,1374,1400]" italics="true" pageId="81" pageNumber="82">Raia spinacidermis</emphasis>
<bibRefCitation id="EFAA4B469538FFDFCE165F53FDAB2517" author="Barnard" box="[374,594,1374,1400]" firstAuthor="Barnard" page="440" pageId="81" pageNumber="82" pagination="439 - 445" refId="ref92395" refString="Barnard, K. H. (1923) Diagnoses of new species of marine fishes from South African waters. Annals of the South African Museum, 13 (Pt. 8), No. 14, 439 - 445." type="journal article" year="1923">Barnard, 1923: 440</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="548088159538FFDFCD3B5F53FD3A2517" box="[603,707,1374,1400]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
(unique):
<specimenCode id="DB9D9ECC9538FFDFCC575F52FC1A2516" box="[823,995,1374,1401]" collectionCode="BMNH" pageId="81" pageNumber="82">BMNH 1935.7</specimenCode>
.19.7. Type locality:
<collectingCountry id="F32C76279538FFDFCBA45F53FAAB2517" box="[1220,1362,1374,1400]" name="South Africa" pageId="81" pageNumber="82">South Africa</collectingCountry>
(exact locality unknown).
</paragraph>
<paragraph id="8B8436B79538FFDFCFF75FABFC97276A" blockId="81.[151,1437,1302,2013]" pageId="81" pageNumber="82">
<emphasis id="B94FEAA59538FFDFCFF75FABFEA425AF" bold="true" box="[151,349,1446,1472]" pageId="81" pageNumber="82">Local synonymy</emphasis>
:
<taxonomicName id="4C3B4D349538FFDFCE0F5FAAFCC425AF" ID-CoL="793L8" authority=": Barnard, 1923: 440" authorityName=": Barnard" authorityPageNumber="440" authorityYear="1923" baseAuthorityName="Barnard" baseAuthorityYear="1923" box="[367,829,1446,1473]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="spinacidermis">
<emphasis id="B94FEAA59538FFDFCE0F5FAAFDB325AF" box="[367,586,1446,1472]" italics="true" pageId="81" pageNumber="82">Raia spinacidermis</emphasis>
:
<bibRefCitation id="EFAA4B469538FFDFCD3C5FABFCC425AF" author="Barnard" box="[604,829,1446,1473]" firstAuthor="Barnard" page="440" pageId="81" pageNumber="82" pagination="439 - 445" refId="ref92395" refString="Barnard, K. H. (1923) Diagnoses of new species of marine fishes from South African waters. Annals of the South African Museum, 13 (Pt. 8), No. 14, 439 - 445." type="journal article" year="1923">Barnard, 1923: 440</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B469538FFDFCC2C5FABFBE725AF" author="Barnard" box="[844,1054,1446,1473]" firstAuthor="Barnard" page="73" pageId="81" pageNumber="82" pagination="1 - 418" refId="ref92438" refString="Barnard, K. H. (1925) A monograph of the marine fishes of South Africa. Part I (Amphioxus, Cyclostomata, Elasmobranchii, and Teleostei-Isospondyli to Heterosomata). Annals of the South African Museum, 21, 1 - 418." type="journal article" year="1925">Barnard, 1925: 73</bibRefCitation>
, fig. 6, pl. 4:
<bibRefCitation id="EFAA4B469538FFDFCBA25FABFA6E25AF" author="Fowler" box="[1218,1431,1446,1472]" firstAuthor="Fowler" page="392" pageId="81" pageNumber="82" pagination="1 - 879" refId="ref101290" refString="Fowler, H. W. (1941) The fishes of the groups Elasmobranchii, Holocephali, Isospondyli, and Ostarophysi obtained by the U. S. Bureau of Fisheries streamer &quot; Albatross &quot; in 1907 to 1910, chiefly in the Philippine Islands and adjacent seas. Bulletin United States National Museum, 100, 1 - 879." type="journal article" year="1941">Fowler, 1941: 392</bibRefCitation>
;
<treatmentCitation id="0A9A10A69538FFDFCFA65FC7FE71258B" box="[198,392,1482,1508]" pageId="81" pageNumber="82">
<bibRefCitation id="EFAA4B469538FFDFCFA65FC7FE71258B" author="Smith, J. L. B." box="[198,392,1482,1508]" pageId="81" pageNumber="82" refId="ref114538" refString="Smith, J. L. B. (1949 a) The Sea Fishes of Southern Africa. South Africa Central News Agency Ltd., 550 pp." type="book" year="1949">Smith, 1949a: 66</bibRefCitation>
</treatmentCitation>
(in part,
<collectingCountry id="F32C76279538FFDFCE915FC7FD7A258B" box="[497,643,1482,1508]" name="South Africa" pageId="81" pageNumber="82">South Africa</collectingCountry>
, not
<taxonomicName id="4C3B4D349538FFDFCDDB5FC6FC78258B" ID-CoL="6X3KQ" authority="Barnard, 1923: 440" authorityName=": von Bonde &amp; Swart" authorityPageNumber="440" authorityYear="1923" box="[699,897,1482,1508]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="durbanensis">
<emphasis id="B94FEAA59538FFDFCDDB5FC6FC78258B" box="[699,897,1482,1508]" italics="true" pageId="81" pageNumber="82">Raia durbanensis</emphasis>
</taxonomicName>
, in synonymy);
<bibRefCitation id="EFAA4B469538FFDFCB5C5FC7FB08258B" author="Smith, J. L. B." box="[1084,1265,1482,1509]" pageId="81" pageNumber="82" refId="ref115700" refString="Smith, J. L. B. (1965) The Sea Fishes of Southern Africa. 5 th Edition. Central News Agency Ltd., 580 pp." type="book" year="1965">Smith, 1965: 66</bibRefCitation>
(in part,
<collectingCountry id="F32C76279538FFDFCA3B5FC7FEEA2666" name="South Africa" pageId="81" pageNumber="82">South Africa</collectingCountry>
).?
<taxonomicName id="4C3B4D349538FFDFCE5B5FE2FE2F2667" authority=": Barnard, 1925: 68" authorityName=": Barnard" authorityPageNumber="68" authorityYear="1925" box="[315,470,1518,1544]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="plutonia">
<emphasis id="B94FEAA59538FFDFCE5B5FE2FE2F2667" box="[315,470,1518,1544]" italics="true" pageId="81" pageNumber="82">Raia plutonia</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9A10A69538FFDFCE865FE3FD4C2667" box="[486,693,1518,1545]" pageId="81" pageNumber="82">
<bibRefCitation id="EFAA4B469538FFDFCE865FE3FD4C2667" author="Barnard" box="[486,693,1518,1545]" firstAuthor="Barnard" page="68" pageId="81" pageNumber="82" pagination="1 - 418" refId="ref92438" refString="Barnard, K. H. (1925) A monograph of the marine fishes of South Africa. Part I (Amphioxus, Cyclostomata, Elasmobranchii, and Teleostei-Isospondyli to Heterosomata). Annals of the South African Museum, 21, 1 - 418." type="journal article" year="1925">Barnard, 1925: 68</bibRefCitation>
</treatmentCitation>
(in part, also including
<typeStatus id="548088159538FFDFCCA85FE3FBD32667" box="[968,1066,1518,1544]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3B4D349538FFDFCB325FE2FEB22643" ID-CoL="78RKK" authority="von Bonde &amp; Swart, 1923" authorityName=": von Bonde &amp; Swart" authorityPageNumber="440" authorityYear="1923" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="albalinea">
<emphasis id="B94FEAA59538FFDFCB325FE2FB032667" box="[1106,1274,1518,1544]" italics="true" pageId="81" pageNumber="82">Raia albalinea</emphasis>
<bibRefCitation id="EFAA4B469538FFDFCA625FE2FEB22643" author="von Bonde, C. &amp; Swart, D. B." pageId="81" pageNumber="82" pagination="1 - 22" refId="ref118185" refString="von Bonde, C. &amp; Swart, D. B. (1923) The Platosomia (skates and rays) collected by the S. S. &quot; Pickle &quot;. University of South Africa, Fisheries and Marine Biological Survey Report, 3 (5), 1 - 22." type="journal article" year="1923">von Bonde &amp; Swart, 1923</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C3B4D349538FFDFCE055C1EFDE52643" ID-CoL="6X3KL" authority="(Barnard, 1923)" authorityName=": Smith" authorityYear="1949" box="[357,540,1554,1580]" class="Elasmobranchii" family="Rajidae" genus="Raia" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="caudaspinosa">
<emphasis id="B94FEAA59538FFDFCE055C1EFDE52643" box="[357,540,1554,1580]" italics="true" pageId="81" pageNumber="82">R. caudaspinosa</emphasis>
</taxonomicName>
).
<taxonomicName id="4C3B4D349538FFDFCD525C1EFC182643" ID-CoL="6X3CD" authority=": Norman, 1935: 46" authorityName="Norman" authorityPageNumber="46" authorityYear="1935" box="[562,993,1554,1581]" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="spinacidermis">
<emphasis id="B94FEAA59538FFDFCD525C1EFCF02643" box="[562,777,1554,1580]" italics="true" pageId="81" pageNumber="82">Raja spinacidermis</emphasis>
:
<bibRefCitation id="EFAA4B469538FFDFCC765C1EFC182643" author="Norman" box="[790,993,1554,1581]" firstAuthor="Norman" page="46" pageId="81" pageNumber="82" pagination="1 - 58" refId="ref110714" refString="Norman, J. R. (1935) Coast fishes, Part. I. The South Atlantic. ' Discovery' Reports, 12, 1 - 58." type="journal article" year="1935">Norman, 1935: 46</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B469538FFDFCC8B5C1FFB4A2643" author="Hulley" box="[1003,1203,1554,1580]" firstAuthor="Hulley" page="173" pageId="81" pageNumber="82" pagination="151 - 220" refId="ref104963" refString="Hulley, P. A. (1970) An investigation of the Rajidae of the west and south coasts of southern Africa. Annals of the South African Museum, 55, 151 - 220." type="journal article" year="1970">Hulley, 1970: 173</bibRefCitation>
, pl. 4;
<bibRefCitation id="EFAA4B469538FFDFCB995C1FFF1D263F" author="Hulley" firstAuthor="Hulley" page="86" pageId="81" pageNumber="82" pagination="1 - 103" refId="ref105041" refString="Hulley, P. A. (1972 a) The origin, interrelationships and distribution of southern African Rajidae (Chondrichthyes, Batoidei). Annals of the South African Museum, 60, 1 - 103. https: // doi. org / 10.2307 / 1442499" type="journal article" year="1972" yearSuffix="a">Hulley, 1972a: 86</bibRefCitation>
, figs. 5859;
<bibRefCitation id="EFAA4B469538FFDFCEE25C3BFD15263F" author="Hulley &amp; Stehmann" box="[386,748,1590,1616]" firstAuthor="Hulley" page="227" pageId="81" pageNumber="82" pagination="227 - 237" refId="ref105277" refString="Hulley, P. A. &amp; Stehmann, M. (1977) The validity of Malacoraja Stehmann, 1970 (Chondrichthyes; Batoidei, Rajidae) and its phylogenetic significance. Annals of the South African Museum, 72 (12), 227 - 237." type="journal article" year="1977">Hulley &amp; Stehmann, 1977: 227</bibRefCitation>
, figs. 15;
<bibRefCitation id="EFAA4B469538FFDFCC0D5C3BFBC5263E" author="Hulley" box="[877,1084,1590,1617]" firstAuthor="Hulley" page="125" pageId="81" pageNumber="82" pagination="115 - 128" refId="ref105183" refString="Hulley, P. A. (1986) Family Rajidae, Anacanthobatidae. In: Smith, M. M. &amp; Heemstra, P. C. (Eds.), Smith's Sea Fishes. Macmillan, Johannesburg, pp. 115 - 128." type="book chapter" year="1986">Hulley, 1986: 125</bibRefCitation>
, fig. 25.19.
<taxonomicName id="4C3B4D349538FFDFCBA85C3AFD7A261B" ID-CoL="6X3CD" authority=": Compagno et al., 1989" authorityName="Compagno" authorityYear="1989" class="Elasmobranchii" family="Rajidae" genus="Raja" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="spinacidermis" subGenus="Malacoraja">
<emphasis id="B94FEAA59538FFDFCBA85C3AFB04263F" box="[1224,1277,1591,1616]" italics="true" pageId="81" pageNumber="82">Raja</emphasis>
(
<emphasis id="B94FEAA59538FFDFCA6F5C3BFA6A263F" box="[1295,1427,1590,1616]" italics="true" pageId="81" pageNumber="82">Malacoraja</emphasis>
)
<emphasis id="B94FEAA59538FFDFCFA65C57FE9A261B" box="[198,355,1626,1652]" italics="true" pageId="81" pageNumber="82">spinacidermis</emphasis>
:
<bibRefCitation id="EFAA4B469538FFDFCE155C57FD7A261B" author="Compagno" box="[373,643,1626,1653]" etAl="et al." firstAuthor="Compagno" pageId="81" pageNumber="82" refId="ref96138" refString="Compagno, L. J. V., Ebert, D. A. &amp; Smale, M. J. (1989) Guide to the Sharks and Rays of Southern Africa. Struik Publishers, Cape Town, 158 pp." type="book" year="1989">
Compagno
<emphasis id="B94FEAA59538FFDFCE9B5C56FDC9261B" box="[507,560,1626,1652]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
., 1989
</bibRefCitation>
</taxonomicName>
, pl.;
<bibRefCitation id="EFAA4B469538FFDFCDA15C57FC05261B" author="Compagno" box="[705,1020,1626,1653]" etAl="et al." firstAuthor="Compagno" page="97" pageId="81" pageNumber="82" pagination="43 - 139" refId="ref96066" refString="Compagno, L. J. V., Ebert, D. A. &amp; Cowley, P. D. (1991) Distribution of offshore demersal cartilaginous fishes (class Chondrichthyes) of the west coast of southern Africa, with notes on their systematics. South African Journal of Marine Science, 11, 43 - 139. https: // doi. org / 10.2989 / 025776191784287664" type="journal article" year="1991">
Compagno
<emphasis id="B94FEAA59538FFDFCC265C56FC82261B" box="[838,891,1626,1652]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
., 1991: 97
</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCB6C5C57FAF0261B" author="Ebert" box="[1036,1289,1626,1653]" etAl="et al." firstAuthor="Ebert" page="76" pageId="81" pageNumber="82" pagination="71 - 81" refId="ref99172" refString="Ebert, D. A., Cowley, P. D. &amp; Compagno, L. J. V. (1991) A preliminary investigation of the feeding ecology of skates (Batoidea: Rajidae) off the west coast of southern Africa. South African Journal of Marine Science, 10, 71 - 81. https: // doi. org / 10.2989 / 02577619109504621" type="journal article" year="1991">
Ebert
<emphasis id="B94FEAA59538FFDFCB335C56FB71261B" box="[1107,1160,1626,1652]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
., 1991: 76
</bibRefCitation>
.
<taxonomicName id="4C3B4D349538FFDFCA785C57FD8A26F7" ID-CoL="6R4RP" authority=": Compagno, 1999: 116" authorityName="Compagno" authorityPageNumber="116" authorityYear="1999" class="Elasmobranchii" family="Rajidae" genus="Malacoraja" kingdom="Animalia" order="Rajiformes" pageId="81" pageNumber="82" phylum="Chordata" rank="species" species="spinacidermis">
<emphasis id="B94FEAA59538FFDFCA785C57FE9A26F7" italics="true" pageId="81" pageNumber="82">Malacoraja spinacidermis</emphasis>
:
<treatmentCitation id="0A9A10A69538FFDFCE145C73FD8A26F7" box="[372,627,1662,1688]" pageId="81" pageNumber="82">
<bibRefCitation id="EFAA4B469538FFDFCE145C73FD8A26F7" author="Compagno" box="[372,627,1662,1688]" firstAuthor="Compagno" page="116" pageId="81" pageNumber="82" pagination="75 - 120" refId="ref95720" refString="Compagno, L. J. V. (1999) An overview of chondrichthyan systematics and biodiversity in southern Africa. Transactions of the Royal Society of South Africa, 54, 75 - 120. https: // doi. org / 10.1080 / 00359199909520406" type="journal article" year="1999">Compagno, 1999: 116</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<bibRefCitation id="EFAA4B469538FFDFCDE15C73FC9126F7" author="Stehmann, M." box="[641,872,1662,1689]" pageId="81" pageNumber="82" pagination="1 - 119" refId="ref116339" refString="Stehmann, M. (1995) First and new records of skates (Chondrichthyes, Rajiformes, Rajidae) from the West African continental slope (Morocco to South Africa), with descriptions of two new species. Archiv f ̡ r Fischereiwissenschaft, 43 (1), 1 - 119." type="journal article" year="1995">Stehmann, 1995: 28</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCC165C73FB2026F7" author="Compagno &amp; Ebert" box="[886,1241,1662,1688]" firstAuthor="Compagno" page="137" pageId="81" pageNumber="82" pagination="125 - 145" refId="ref96013" refString="Compagno, L. J. V. &amp; Ebert, D. A. (2007) Southern African skate biodiversity and distribution. Environmental Biology of Fishes, 80, 125 - 145. https: // doi. org / 10.1007 / s 10641 - 007 - 9243 - 4" type="journal article" year="2007">Compagno &amp; Ebert, 2007: 137</bibRefCitation>
, fig. 5e;
<bibRefCitation id="EFAA4B469538FFDFCA215C73FE3F26D3" author="Ebert &amp; Compagno" firstAuthor="Ebert" page="120" pageId="81" pageNumber="82" pagination="125 - 145" refId="ref96013" refString="Compagno, L. J. V. &amp; Ebert, D. A. (2007) Southern African skate biodiversity and distribution. Environmental Biology of Fishes, 80, 125 - 145. https: // doi. org / 10.1007 / s 10641 - 007 - 9243 - 4" type="journal article" year="2007">Ebert &amp; Compagno, 2007: 120</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCEB35CAFFD3126D3" author="Ebert" box="[467,712,1698,1725]" etAl="et al." firstAuthor="Ebert" page="91" pageId="81" pageNumber="82" pagination="81 - 102" refId="ref98910" refString="Ebert, D. A., Compagno, L. J. V. &amp; Cowley, P. D. (2008) Aspects on the reproductive biology of skates (Chondrichthyes: Rajiformes: Rajoidei) from southern African waters. ICES Journal of Marine Science, 65, 81 - 102. https: // doi. org / 10.1093 / icesjms / fsm 169" type="journal article" year="2008">
Ebert
<emphasis id="B94FEAA59538FFDFCD785CAEFDB226D3" box="[536,587,1698,1724]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
., 2008: 91
</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCDB65CAFFC6026D2" author="NPOA" box="[726,921,1698,1725]" firstAuthor="NPOA" page="55" pageId="81" pageNumber="82" refId="ref110949" refString="NPOA. (2013) National Plan of Action for the Conservation and Management of Sharks (NPOA-Sharks). Department of Agriculture, Forestry and Fisheries (DAFF), Rogge Bay, Cape Town, 63 pp." type="book" year="2013">NPOA, 2013: 55</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCCC65CAFFB3026D3" author="da Silva" box="[934,1225,1698,1725]" etAl="et al." firstAuthor="da Silva" page="247" pageId="81" pageNumber="82" pagination="233 - 248" refId="ref96965" refString="da Silva C., Booth, A. J., Dudley, S. F. J., Kerwath, S. E., Lamberth, S. J., Leslie, R. W., McCord, M. E., Sauer, W. H. H. &amp; Zweig, T. (2015) A description and updated overview of the status and management of South Africa's chondrichthyan fisheries. South African Journal of Marine Science, 37, 233 - 248. https: // doi. org / 10.2989 / 1814232 X. 2015.1044471" type="journal article" year="2015">
da Silva
<emphasis id="B94FEAA59538FFDFCB6B5CAEFBC726D3" box="[1035,1086,1698,1724]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
., 2015: 247
</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCBB75CAFFA6E26D3" author="Ebert, D. A." box="[1239,1431,1698,1725]" pageId="81" pageNumber="82" refId="ref98428" refString="Ebert, D. A. (2015) Deep-sea cartilaginous fishes of the Southeastern Atlantic Ocean. FAO Species Catalogue for Fishery Purposes No. 9. FAO, Rome, 251 pp." type="book" year="2015">Ebert, 2015: 181</bibRefCitation>
, fig. 205;
<bibRefCitation id="EFAA4B469538FFDFCE4C5CCBFD8C268F" author="Ebert &amp; van Hees" box="[300,629,1734,1761]" firstAuthor="Ebert" page="147" pageId="81" pageNumber="82" pagination="141 - 156" refId="ref99877" refString="Ebert, D. A. &amp; van Hees, K. E. (2015) Beyond jaws: rediscovering the &quot; Lost Sharks &quot; of southern Africa. African Journal of Marine Science, 37, 141 - 156. https: // doi. org / 10.2989 / 1814232 X. 2015.1048730" type="journal article" year="2015">Ebert &amp; van Hees, 2015: 147</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCDE15CCAFC97268F" author="Last" box="[641,878,1734,1761]" etAl="et al." firstAuthor="Last" page="23" pageId="81" pageNumber="82" pagination="11 - 34" refId="ref107873" refString="Last, P. R., Weigmann, S. &amp; Yang, L. (2016 f) Changes to the nomenclature of the skates (Chondrichthyes: Rajiformes). In: Last, P. R. &amp; Yearsley, G. K. (Eds.), Rays of the World: Supplementary Information. CSIRO Australian National Fish Collection, Hobart, pp. 11 - 34." type="book chapter" year="2016" yearSuffix="f">
Last
<emphasis id="B94FEAA59538FFDFCDD85CCAFD13268F" box="[696,746,1734,1760]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
., 2016f: 23
</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCC195CCAFB81268F" author="Last" box="[889,1144,1734,1761]" etAl="et al." firstAuthor="Last" page="296" pageId="81" pageNumber="82" pagination="204 - 363" refId="ref107642" refString="Last, P. R., Seret, B., Stehmann, M. F. W. &amp; Weigmann, S. (2016 g) Skates. Family Rajidae. In: Last, P. R., White, W. T., de Carvalho, M. E., Seret, B., Stehmann, M. F. W. &amp; Naylor, G. J. P. (Eds.), Rays of the World. Cornell University Press, Ithaca, New York, pp. 204 - 363. https: // doi. org / 10.1071 / 9780643109148" type="book chapter" year="2016" yearSuffix="g">
Last
<emphasis id="B94FEAA59538FFDFCCD15CCAFC1B268F" box="[945,994,1734,1760]" italics="true" pageId="81" pageNumber="82">et al</emphasis>
.,
<quantity id="4CC39B529538FFDFCC975CCBFBB8268F" box="[1015,1089,1734,1760]" metricMagnitude="0" metricUnit="kg" metricValue="2.016" pageId="81" pageNumber="82" unit="g" value="2016.0">2016g</quantity>
: 296
</bibRefCitation>
, fig. 19.87; McEachran &amp;
<bibRefCitation id="EFAA4B469538FFDFCFA65CE7FE76276B" author="Seret, B." box="[198,399,1770,1796]" pageId="81" pageNumber="82" pagination="1404 - 1433" refId="ref113064" refString="Seret, B. (2016) Batoid fishes. Multiple families. In: Carpenter, K. E. &amp; De Angelis, N. (Eds.), The Living Marine Resources of the Eastern Central Atlantic. Vol. 2. Bivalves, Gastropods, Hagfishes, Sharks, Batoid Fishes, and Chimaeras. FAO Species Identification Guide for Fishery Purposes. FAO, Rome, pp. 1337 - 1369, 1404 - 1433." type="journal article" year="2016">Séret, 2016: 1386</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCEFB5CE7FD9D276A" author="Seret, B." box="[411,612,1770,1797]" pageId="81" pageNumber="82" pagination="1404 - 1433" refId="ref113064" refString="Seret, B. (2016) Batoid fishes. Multiple families. In: Carpenter, K. E. &amp; De Angelis, N. (Eds.), The Living Marine Resources of the Eastern Central Atlantic. Vol. 2. Bivalves, Gastropods, Hagfishes, Sharks, Batoid Fishes, and Chimaeras. FAO Species Identification Guide for Fishery Purposes. FAO, Rome, pp. 1337 - 1369, 1404 - 1433." type="journal article" year="2016">Séret, 2016: 1345</bibRefCitation>
;
<bibRefCitation id="EFAA4B469538FFDFCD105CE7FC92276A" author="Weigmann, S." box="[624,875,1770,1797]" pageId="81" pageNumber="82" pagination="837 - 1037" refId="ref118762" refString="Weigmann, S. (2016) Annotated checklist of the living sharks, batoids and chimaeras (Chondrichthyes) of the world, with a focus on biogeographical diversity. Journal of Fish Biology, 88, 837 - 1037. https: // doi. org / 10.1111 / jfb. 12874" type="journal article" year="2016">Weigmann, 2016: 950</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321653C9538FFDFCFF75D03FB6E2747" box="[151,1175,1806,1833]" pageId="81" pageNumber="82" type="materials_examined">
<paragraph id="8B8436B79538FFDFCFF75D03FB6E2747" blockId="81.[151,1437,1302,2013]" box="[151,1175,1806,1833]" pageId="81" pageNumber="82">
<materialsCitation id="3B533CEA9538FFDFCFF75D03FB6E2747" ID-GBIF-Occurrence="3072574314" box="[151,1175,1806,1833]" collectionCode="BMNH, SAIAB" country="South Africa" location="South Africa" pageId="81" pageNumber="82" specimenCode="BMNH 1935.7, SAIAB 27330" specimenCount="1" typeStatus="holotype">
<emphasis id="B94FEAA59538FFDFCFF75D03FDFB2747" bold="true" box="[151,514,1806,1832]" pageId="81" pageNumber="82">
<collectingCountry id="F32C76279538FFDFCFF75D03FED62747" box="[151,303,1806,1832]" name="South Africa" pageId="81" pageNumber="82">South Africa</collectingCountry>
voucher material
</emphasis>
:
<typeStatus id="548088159538FFDFCD715D03FD872747" box="[529,638,1806,1832]" pageId="81" pageNumber="82" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DB9D9ECC9538FFDFCDE85D02FCCC2746" box="[648,821,1806,1833]" collectionCode="BMNH" pageId="81" pageNumber="82">BMNH 1935.7</specimenCode>
.19.7. Non-type:
<specimenCode id="DB9D9ECC9538FFDFCC8E5D03FB6A2747" box="[1006,1171,1806,1832]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="81" pageNumber="82" type="Other type of research institution/biorepository">SAIAB 27330</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C321653C9538FFDFCFF75D3FFC2E2723" box="[151,983,1842,1869]" pageId="81" pageNumber="82" type="distribution">
<paragraph id="8B8436B79538FFDFCFF75D3FFC2E2723" blockId="81.[151,1437,1302,2013]" box="[151,983,1842,1869]" pageId="81" pageNumber="82">
<emphasis id="B94FEAA59538FFDFCFF75D3FFE2A2723" bold="true" box="[151,467,1842,1868]" pageId="81" pageNumber="82">South African distribution</emphasis>
: The Orange River (NP) to
<collectingRegion id="49FFF8559538FFDFCC6D5D3FFCBE2723" box="[781,839,1842,1868]" country="South Africa" name="Western Cape" pageId="81" pageNumber="82">Cape</collectingRegion>
Point (WC).
</paragraph>
</subSubSection>
<subSubSection id="C321653C9538FFDFCFF75D5BFDE927B3" pageId="81" pageNumber="82" type="discussion">
<subSubSection id="C321653C9538FFDFCFF75D5BFD3727D7" pageId="81" pageNumber="82" type="discussion">
<paragraph id="8B8436B79538FFDFCFF75D5BFD3727D7" blockId="81.[151,1437,1302,2013]" pageId="81" pageNumber="82">
<emphasis id="B94FEAA59538FFDFCFF75D5BFEFD271F" bold="true" box="[151,260,1878,1904]" pageId="81" pageNumber="82">Remarks</emphasis>
: This species also occurs in the North Atlantic, but its distribution north of
<collectingCountry id="F32C76279538FFDFCBF05D5BFADA271F" box="[1168,1315,1878,1904]" name="South Africa" pageId="81" pageNumber="82">South Africa</collectingCountry>
is patchy. Morphological and molecular comparison of specimens from
<collectingCountry id="F32C76279538FFDFCC195D77FBFE27FB" box="[889,1031,1914,1940]" name="South Africa" pageId="81" pageNumber="82">South Africa</collectingCountry>
and the North Atlantic may confirm if it is the same species or if they are different.
</paragraph>
</subSubSection>
<subSubSection id="C321653C9538FFDFCFF75DCFFDE927B3" box="[151,528,1986,2013]" pageId="81" pageNumber="82" type="multiple">
<paragraph id="8B8436B79538FFDFCFF75DCFFDE927B3" blockId="81.[151,1437,1302,2013]" box="[151,528,1986,2013]" pageId="81" pageNumber="82">
<emphasis id="B94FEAA59538FFDFCFF75DCFFE7D27B3" bold="true" box="[151,388,1986,2012]" pageId="81" pageNumber="82">Conservation status</emphasis>
: LC (2020).
</paragraph>
</subSubSection>
</subSubSection>
</treatment>
</document>