565 lines
76 KiB
XML
565 lines
76 KiB
XML
<document id="E3F5E6CB6468638E9CCBA713A815EB29" ID-CLB-Dataset="31038" ID-DOI="10.11646/zootaxa.4370.2.5" ID-GBIF-Dataset="b1eaefaf-c0cf-4527-bd3f-dc374e43cf35" ID-ISSN="1175-5326" ID-Zenodo-Dep="1143921" ID-ZooBank="5921F955-95FE-47EC-B94E-7B72D39B8D2C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1515573681623" checkinUser="plazi" docAuthor="Graf, Wolfram, Pauls, Steffen U. & Vitecek, Simon" docDate="2018" docId="03928791EA5E2F7243BCFF3BFE57FD96" docLanguage="en" docName="zootaxa.4370.2.5.pdf" docOrigin="Zootaxa 4370 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isoperla vjosae Graf & Pauls & Vitecek 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="178" masterDocId="FFABFFE9EA5D2F75432BFFE7FFB7FFAB" masterDocTitle="Isoperla vjosae sp. n., a new species of the Isoperla tripartita group from Albania (Plecoptera: Perlodidae)" masterLastPageNumber="179" masterPageNumber="171" pageNumber="174" updateTime="1698513581474" updateUser="ExternalLinkService">
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<mods:mods id="F0C1D4FD6C75136A0A46AD05BB1AD779" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AD7C9FA94A4FD4AACB018389DACA7ADD">
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<mods:title id="FBE3D376B25AEE57FED3A6752106A8A5">Isoperla vjosae sp. n., a new species of the Isoperla tripartita group from Albania (Plecoptera: Perlodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C37591AD31D9A89DED8F943A8A1078BB" type="personal">
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<mods:role id="E759113AFDD1767788C45777C6767F44">
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<mods:roleTerm id="2D11C98E8134657ADF85CC0136DBC068">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5E8BBA2103790391D1B435CA788B81DA">Graf, Wolfram</mods:namePart>
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</mods:name>
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<mods:roleTerm id="847DD1F4C3AC1D12164D3F9C3C600C18">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E93B4BBFF8F27AA1F70D987A5DA81966">Pauls, Steffen U.</mods:namePart>
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</mods:name>
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<mods:name id="FC4758D4421F261C0B78C27904C71D01" type="personal">
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<mods:roleTerm id="DAB9CF406E4FFC890E1658FCE9A52BAC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F2941289EB2886190AA7B6F0E41A6C9C">Vitecek, Simon</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2973A8F9104B228B337F9164B5FCA6EF">text</mods:typeOfResource>
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<mods:titleInfo id="5783552E29605A5286239B7659D9C71B">
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<mods:title id="FEBDE3E172EB3D69EF3B8C9F272C0657">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CFC7DA18458287406269E8B5DA499F3C">
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<mods:date id="0B5D52B7BF5019C6771FCD762E321295">2018</mods:date>
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<mods:number id="D086EB3217B7E792883EBF01619402CA">2018-01-10</mods:number>
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<mods:number id="EF90D9B1732234A51C201EA38CCDB7BD">4370</mods:number>
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</mods:detail>
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<mods:detail id="97E0C967A6FBCD10D2FB61ED8CC4DAF2" type="issue">
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<mods:number id="5798CDD4A2EA86D820AADD9BFA506453">2</mods:number>
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<mods:start id="BC0BC3EE395FE2E94DC6AB46D254ADA3">171</mods:start>
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<mods:classification id="FB9C67ED97FE2573FEC8CAFF0677394E">journal article</mods:classification>
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<mods:identifier id="E3BBCAF055A7D161DA8EB5496F131ABD" type="CLB-Dataset">31038</mods:identifier>
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<mods:identifier id="7CB6329E3236B6D9647F66A7A731C31F" type="DOI">10.11646/zootaxa.4370.2.5</mods:identifier>
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<mods:identifier id="C462A25451C76343A4FDCECCC01D12E6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FCE89A1CA99353083C8109B66A4C4552" type="Zenodo-Dep">1143921</mods:identifier>
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</mods:mods>
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<treatment id="03928791EA5E2F7243BCFF3BFE57FD96" ID-DOI="http://doi.org/10.5281/zenodo.5961733" ID-GBIF-Taxon="140236757" ID-Zenodo-Dep="5961733" LSID="urn:lsid:plazi:treatment:03928791EA5E2F7243BCFF3BFE57FD96" httpUri="http://treatment.plazi.org/id/03928791EA5E2F7243BCFF3BFE57FD96" lastPageId="7" lastPageNumber="178" pageId="3" pageNumber="174">
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<subSubSection id="C321650CEA5E2F7643BCFF3BFE9AFEB3" pageId="3" pageNumber="174" type="nomenclature">
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<paragraph id="8B843687EA5E2F7643BCFF3BFD33FF5C" blockId="3.[151,644,220,280]" box="[151,644,220,247]" pageId="3" pageNumber="174">
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<heading id="D0CC81EBEA5E2F7643BCFF3BFD33FF5C" bold="true" box="[151,644,220,247]" fontSize="11" level="1" pageId="3" pageNumber="174" reason="1">
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<emphasis id="B94FEA95EA5E2F7643BCFF3BFD33FF5C" bold="true" box="[151,644,220,247]" pageId="3" pageNumber="174">
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<taxonomicName id="4C3B4D04EA5E2F7643BCFF3BFEE6FF5D" ID-CoL="3QB2N" authority="Graf & Pauls & Vitecek, 2018" authorityName="Graf & Pauls & Vitecek" authorityYear="2018" box="[151,337,220,246]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="vjosae" status="sp. nov.">
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<emphasis id="B94FEA95EA5E2F7643BCFF3BFEE6FF5D" bold="true" box="[151,337,220,246]" italics="true" pageId="3" pageNumber="174">Isoperla vjosae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27C57EEEA5E2F764273FF3AFE0FFF5C" box="[344,440,221,247]" pageId="3" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
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Graf & Vitecek
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B843687EA5E2F7643BCFF18FE9AFEB3" blockId="3.[151,644,220,280]" box="[151,301,255,280]" pageId="3" pageNumber="174">
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||
<figureCitation id="13002A02EA5E2F7643BCFF18FE9AFEB3" box="[151,301,255,280]" captionStart-0="FIGURES 2–13" captionStart-1="FIGURE 14" captionStartId-0="4.[151,264,1895,1918]" captionStartId-1="5.[151,250,876,898]" captionTargetBox-0="[184,1394,194,1867]" captionTargetBox-1="[151,1436,193,854]" captionTargetId-0="figure@4.[183,1395,193,1868]" captionTargetId-1="figure@5.[151,1436,193,855]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." captionText-1="FIGURE 14. Isoperla vjosae sp. nov., lacinia; FIGURES 15–17. Isoperla tripartita tripartita; 15: medial penial armatUres; 16: male meso- and metasternUm; 17: vesicle of sternite VIII; FIGURE 18. Isoperla pesici, medial penial armatUres.Scale bar represents 500 µm." httpUri-0="https://zenodo.org/record/1143925/files/figure.png" httpUri-1="https://zenodo.org/record/1143929/files/figure.png" pageId="3" pageNumber="174">Figures 2–14</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C321650CEA5E2F7643BCFEA5FBADFD41" pageId="3" pageNumber="174" type="materials_examined">
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<paragraph id="8B843687EA5E2F7643BCFEA5FBD7FE14" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
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<materialsCitation id="3B533CDAEA5E2F7643BCFEA5FCC3FED7" ID-GBIF-Occurrence="1802552355" collectingDate="2017-04-25" collectionCode="SMF" collectorName="Graf & Itri" country="Albania" latitude="40.4726" location="Vjosa River" longLatPrecision="42" longitude="19.749" municipality="Kute" pageId="3" pageNumber="174" specimenCount="2" typeStatus="holotype">
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<emphasis id="B94FEA95EA5E2F7643BCFEA5FEE7FEF0" bold="true" box="[151,336,322,347]" pageId="3" pageNumber="174">
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||
<typeStatus id="54808825EA5E2F7643BCFEA5FF64FEF0" box="[151,211,322,347]" pageId="3" pageNumber="174">Type</typeStatus>
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material.
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</emphasis>
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||
<typeStatus id="54808825EA5E2F764275FEA5FE7EFEF0" box="[350,457,322,347]" pageId="3" pageNumber="174" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="F32C7617EA5E2F76410EFEA4FD1FFEF0" box="[549,680,323,347]" name="Albania" pageId="3" pageNumber="174">ALBANIA</collectingCountry>
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,
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<location id="8EE4605CEA5E2F764190FEA5FCFBFEF0" LSID="urn:lsid:plazi:treatment:03928791EA5E2F7243BCFF3BFE57FD96:8EE4605CEA5E2F764190FEA5FCFBFEF0" box="[699,844,322,347]" country="Albania" latitude="40.4726" longLatPrecision="42" longitude="19.749" municipality="Kute" name="Vjosa River" pageId="3" pageNumber="174">Vjosa River</location>
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||
near the village of
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||
<collectingMunicipality id="6BE0ACFDEA5E2F764764FEA4FB3DFEF1" box="[1103,1162,323,346]" pageId="3" pageNumber="174">Kutë</collectingMunicipality>
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||
;
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||
<geoCoordinate id="EE0F5040EA5E2F7647B6FEA5FAADFEF0" box="[1181,1306,322,347]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="5" value="40.4726">40.4726°N</geoCoordinate>
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||
<geoCoordinate id="EE0F5040EA5E2F764603FEA4FA2FFEF0" box="[1320,1432,323,347]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="55" value="19.749">19.749°E</geoCoordinate>
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||
;
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||
<collectingDate id="EFC1E9AFEA5E2F7643BCFE83FEA5FED6" box="[151,274,356,381]" pageId="3" pageNumber="174" value="2017-04-25">
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||
<date id="FF851047EA5E2F7643BCFE83FEA5FED6" box="[151,274,356,381]" pageId="3" pageNumber="174" value="2017-04-25">25.iv.2017</date>
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||
</collectingDate>
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||
, leg.
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||
<collectorName id="26CE5351EA5E2F764272FE83FE3AFED6" box="[345,397,356,381]" pageId="3" pageNumber="174">Graf</collectorName>
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||
(specimen identifier:
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<collectorName id="26CE5351EA5E2F7641B2FE83FD0AFED6" box="[665,701,356,381]" pageId="3" pageNumber="174">Itri</collectorName>
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||
0301M [
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||
<collectionCode id="ED2AAE42EA5E2F764009FE83FCD3FED7" box="[802,868,356,380]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="174">SMF</collectionCode>
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||
])
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B533CDAEA5E2F7640A8FE82FBEBFE14" ID-GBIF-Occurrence="1802552356" collectionCode="SMF" collectorName="Graf" country="Austria" location="Biologiezentrum Linz" municipality="Linz" pageId="3" pageNumber="174" specimenCount="46" specimenCount-female="10" specimenCount-larva="13" specimenCount-male="23" typeStatus="paratype">
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||
<typeStatus id="54808825EA5E2F7640A8FE82FC40FED6" box="[899,1015,357,381]" pageId="3" pageNumber="174" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="9D3DFD0EEA5E2F76472CFE83FBC1FED6" box="[1031,1142,356,381]" pageId="3" pageNumber="174" type="male">21 males</specimenCount>
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||
,
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||
<specimenCount id="9D3DFD0EEA5E2F7647ADFE82FB4BFED6" box="[1158,1276,356,381]" pageId="3" pageNumber="174" type="female">7 females</specimenCount>
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||
,
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||
<specimenCount id="9D3DFD0EEA5E2F764627FE83FACAFED6" box="[1292,1405,356,381]" pageId="3" pageNumber="174" type="larva">11 larvae</specimenCount>
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, 2 exuviae; same collection data (
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<specimenCount id="9D3DFD0EEA5E2F7642D4FE61FDD4FE36" box="[511,611,390,413]" pageId="3" pageNumber="174" type="female">1 female</specimenCount>
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||
, 1 larva,
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||
<collectionCode id="ED2AAE42EA5E2F7641F9FE61FCBBFE36" box="[722,780,390,413]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="174">SMF</collectionCode>
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||
; all other specimens currently in coll.
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<collectorName id="26CE5351EA5E2F7647E3FE61FB4DFE36" box="[1224,1274,390,413]" pageId="3" pageNumber="174">Graf</collectorName>
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||
–
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||
<specimenCount id="9D3DFD0EEA5E2F764634FE61FAC9FE35" box="[1311,1406,389,414]" pageId="3" pageNumber="174" type="male">2 males</specimenCount>
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||
,
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||
<specimenCount id="9D3DFD0EEA5E2F7646A6FE61FF47FE14" pageId="3" pageNumber="174" type="female">2 females</specimenCount>
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||
and
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||
<specimenCount id="9D3DFD0EEA5E2F764203FE40FE34FE14" box="[296,387,422,447]" pageId="3" pageNumber="174" type="larva">2 larvae</specimenCount>
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||
will be transferred to the
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<location id="8EE4605CEA5E2F764186FE41FC1CFE14" LSID="urn:lsid:plazi:treatment:03928791EA5E2F7243BCFF3BFE57FD96:8EE4605CEA5E2F764186FE41FC1CFE14" box="[685,939,422,447]" country="Austria" municipality="Linz" name="Biologiezentrum Linz" pageId="3" pageNumber="174">Biologiezentrum Linz</location>
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||
,
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<collectingMunicipality id="6BE0ACFDEA5E2F76409CFE41FC59FE14" box="[951,1006,422,447]" pageId="3" pageNumber="174">Linz</collectingMunicipality>
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||
,
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<collectingCountry id="F32C7617EA5E2F7640D1FE41FBE3FE14" box="[1018,1108,422,447]" name="Austria" pageId="3" pageNumber="174">Austria</collectingCountry>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8B843687EA5E2F7643ECFE20FD9FFDA9" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
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||
Additional material:
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||
<specimenCount id="9D3DFD0EEA5E2F7642E8FE2FFDADFE4B" box="[451,538,455,480]" pageId="3" pageNumber="174" type="male">1 male</specimenCount>
|
||
;
|
||
<collectingCountry id="F32C7617EA5E2F764107FE2FFD18FE4B" box="[556,687,456,480]" name="Albania" pageId="3" pageNumber="174">ALBANIA</collectingCountry>
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||
, Vjosa River near the village of Kutë;
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||
<geoCoordinate id="EE0F5040EA5E2F7647B5FE20FAACFE4B" box="[1182,1307,455,480]" direction="north" orientation="latitude" pageId="3" pageNumber="174" precision="5" value="40.4726">40.4726°N</geoCoordinate>
|
||
<geoCoordinate id="EE0F5040EA5E2F764602FE2FFA2FFE4B" box="[1321,1432,456,480]" direction="east" orientation="longitude" pageId="3" pageNumber="174" precision="55" value="19.749">19.749°E</geoCoordinate>
|
||
;
|
||
<date id="FF851047EA5E2F7643BCFE0EFEA3FDA9" box="[151,276,489,514]" pageId="3" pageNumber="174" value="2017-06-14">14.vi.2017</date>
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||
, leg. Graf, in coll. Graf.
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5E2F7643ECFDEDFDCCFDCD" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
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||
<emphasis id="B94FEA95EA5E2F7643ECFDEDFEF3FD88" bold="true" box="[199,324,522,547]" pageId="3" pageNumber="174">DiagnoSiS.</emphasis>
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Ventral lobe of the penis with partitioned medial penial armature comprising a distal and a proximal part, both subdivided into diverging, elongated left and right portions. Scales of the penial armature spike-like. Lateral penial armatures small.
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</paragraph>
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<paragraph id="8B843687EA5E2F7643ECFD88FD68FD03" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
|
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In all other species of
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<taxonomicName id="4C3B4D04EA5E2F7642E4FD89FDD1FD2D" authorityName="tripartita Illies" authorityYear="1954" box="[463,614,622,647]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
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||
the
|
||
<emphasis id="B94FEA95EA5E2F7642D7FD88FDD1FD2D" box="[508,614,623,646]" italics="true" pageId="3" pageNumber="174">tripartita</emphasis>
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</taxonomicName>
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group either the distal or the proximal part of the medial ventral lobe penial armature forms a continuous band of scales.
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</paragraph>
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<paragraph id="8B843687EA5E2F7643ECFD56FBADFD41" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94FEA95EA5E2F7643ECFD56FEEBFD61" bold="true" box="[199,348,689,714]" pageId="3" pageNumber="174">DeSCription.</emphasis>
|
||
Medium to large species, macropterous. Body length:
|
||
<typeStatus id="54808825EA5E2F7640F8FD56FB80FD61" box="[979,1079,689,714]" pageId="3" pageNumber="174" type="holotype">holotype</typeStatus>
|
||
male
|
||
<quantity id="4CC39B62EA5E2F764755FD56FB6FFD62" box="[1150,1240,689,714]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="174" unit="mm" value="9.5">9.5 mm</quantity>
|
||
,
|
||
<typeStatus id="54808825EA5E2F7647CFFD55FAF0FD61" box="[1252,1351,690,714]" pageId="3" pageNumber="174" type="paratype">paratype</typeStatus>
|
||
female 11.0 mm; forewing length:
|
||
<typeStatus id="54808825EA5E2F7642FBFD35FD83FD40" box="[464,564,722,747]" pageId="3" pageNumber="174" type="holotype">holotype</typeStatus>
|
||
male 11.0 mm,
|
||
<typeStatus id="54808825EA5E2F7641C5FD34FCE6FD40" box="[750,849,723,747]" pageId="3" pageNumber="174" type="paratype">paratype</typeStatus>
|
||
female 14.0 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321650CEA5E2F7043BCFD13FEE4F923" lastPageId="5" lastPageNumber="176" pageId="3" pageNumber="174" type="description">
|
||
<paragraph id="8B843687EA5E2F7643BCFD13FD58FB16" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94FEA95EA5E2F7643BCFD13FEF4FCA6" bold="true" box="[151,323,756,781]" pageId="3" pageNumber="174">Adult habituS</emphasis>
|
||
: General colour yellowish but head and pronotum mostly yellow with dark brown markings; body and legs pilose. Ground colour of the head yellow, with rectangular brown marking connecting the three ocelli, leaving a pale subrhombic area between them; with brownish band extending rostrally from the M-line; M-line rather distinct, yellowish (
|
||
<figureCitation id="13002A02EA5E2F764106FCB8FD12FCD3" box="[557,677,863,888]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Figs. 2, 3</figureCitation>
|
||
); tentorial callosities inconspicuous, yellow; occiput with even surface, in some specimens with two dark brown semicircular areas meeting at the coronal suture. Eyes as large as the area delimited by the three ocelli. Scape dorsally dark brown; pedicel and the following ten antennomeres dorsally creamy; scape, pedicel and basal antennomeres ventrally yellowish; distal part of the antenna brown. Palpi cream-coloured. Pronotum yellow, rectangular, with angled edges; areas with textured surface dark brown, the surrounding area smooth and cream-coloured, anterior and posterior lines located distally and anteriorly of areas with textured surface dark brown. Mesonotum dark brown but yellow anteriorly, metanotum dark brown. Wings translucent yellowish; venation dark brown. Ventral surface of thorax pale, meso- and metabasisternum off-whitish, furcasternites pale, furcal pits inconspicuous (
|
||
<figureCitation id="13002A02EA5E2F76472AFB98FBFEFB33" box="[1025,1097,1151,1176]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Fig. 4</figureCitation>
|
||
). Femora brown dorsally and ventrally. Tibiae uniformly brown; tarsi dark brown.
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5E2F7643ECFB20FAEDFA13" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94FEA95EA5E2F7643ECFB20FEC8FB4B" bold="true" box="[199,383,1223,1248]" pageId="3" pageNumber="174">Male abdomen</emphasis>
|
||
: Tergites I-IX dorsally with prominent broad dark posterior margin; tergite X uniformly yellow. Tergite I-IV dorsally entirely brown; tergites IV-IX subsequently getting more yellowish mediolaterally; tergites VIII to IX dorsally with brown longitudinal band (
|
||
<figureCitation id="13002A02EA5E2F764013FAE8FCC9FA83" box="[824,894,1295,1320]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Fig. 5</figureCitation>
|
||
). Tergites I–IX dorsally with transverse row of four pigmented spots; laterally with two to three spots; all spots with pale centre. Ventral surface of abdomen pale. Sternite VIII vesicle brownish, slightly longer than wide, its posterior margin evenly rounded; nearly as long as half the segment’s length; posterior margin of sternite VIII with additional spines (
|
||
<figureCitation id="13002A02EA5E2F764797FA9BFAB2FA3E" box="[1212,1285,1404,1429]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Fig. 6</figureCitation>
|
||
). Sternite IX yellowish, paraprocts yellowish; cerci yellowish with brown ring at terminal end of each segment (
|
||
<figureCitation id="13002A02EA5E2F76462EFA78FAFCFA13" box="[1285,1355,1439,1464]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5E2F7643ECFA23FB26F973" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94FEA95EA5E2F7643ECFA23FE37FA76" bold="true" box="[199,384,1476,1501]" pageId="3" pageNumber="174">PeniS (everted)</emphasis>
|
||
: Medial penial armature on ventral surface of penis divided into a distal and a proximal part, both coloured; distal armature located on medial lobe adjacent to ventral lobe, divided into suboval left and right portions, length of each portion 290-300 µm, width 100-110 µm; proximal armature located on central part of ventral lobe, divided by 85-93 µm gap lacking large coloured scales into left and right portions, length of each portion 250-260 µm, width 100-125 µm; both distal and proximal armatures proximally diverging, usually forming an upturned ‘V’ (
|
||
<figureCitation id="13002A02EA5E2F7642EAF990FD93F93B" box="[449,548,1655,1680]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Fig. 7, 8</figureCitation>
|
||
). Scales of medial penial armature spike-like and straight, scales of proximal part usually longer: length of distal part scales 44-58 µm, width 4-7µm; length of proximal part scales 97-138 µm, width comparable to distal part scales. Lateral penial armatures small and indistinct.
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5E2F7643ECF903FB17F88B" blockId="3.[151,1437,322,1861]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94FEA95EA5E2F7643ECF903FE2BF956" bold="true" box="[199,412,1764,1789]" pageId="3" pageNumber="174">Female abdomen</emphasis>
|
||
: All tergites uniformly dark brown. Sternites and cerci pale yellow. Subgenital plate evenly rounded, covering half of sternite VIII and the anterior third of sternite IX (
|
||
<figureCitation id="13002A02EA5E2F764767F8E0FB24F88B" box="[1100,1171,1799,1824]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="3" pageNumber="174">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5E2F7643ECF8CBFCE1F8EE" blockId="3.[151,1437,322,1861]" box="[199,854,1836,1861]" pageId="3" pageNumber="174">
|
||
<emphasis id="B94FEA95EA5E2F7643ECF8CBFF40F8EE" bold="true" box="[199,247,1836,1861]" pageId="3" pageNumber="174">Egg</emphasis>
|
||
: unknown. No mature eggs were available for study.
|
||
</paragraph>
|
||
<caption id="DF44660FEA592F7143BCF880FAE5F871" httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="4" pageNumber="175" targetBox="[184,1394,194,1867]" targetPageId="4">
|
||
<paragraph id="8B843687EA592F7143BCF880FAE5F871" blockId="4.[151,1436,1895,2010]" pageId="4" pageNumber="175">
|
||
<emphasis id="B94FEA95EA592F7143BCF880FEE7F8D6" bold="true" box="[151,336,1895,1918]" pageId="4" pageNumber="175">FIGURES 2–13:</emphasis>
|
||
<taxonomicName id="4C3B4D04EA592F714276F88FFE48F8D5" authority="Graf & Pauls & Vitecek, 2018" authorityName="Graf & Pauls & Vitecek" authorityYear="2018" box="[349,511,1896,1918]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="vjosae" status="sp. nov.">
|
||
<emphasis id="B94FEA95EA592F714276F88FFE48F8D5" box="[349,511,1896,1918]" italics="true" pageId="4" pageNumber="175">Isoperla vjosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57EEEA592F714127F88FFDD3F8D6" box="[524,612,1896,1917]" pageId="4" pageNumber="175" rank="species">
|
||
<emphasis id="B94FEA95EA592F714127F88FFDD3F8D6" bold="true" box="[524,612,1896,1917]" pageId="4" pageNumber="175">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval head and pronotUm; 12: larval meso-and metathorax; 13: larval abdominal tergites. Scale bar represents 500 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF44660FEA582F7043BCFC8BFEEEFC14" httpUri="https://zenodo.org/record/1143929/files/figure.png" pageId="5" pageNumber="176" targetBox="[151,1436,193,854]" targetPageId="5">
|
||
<paragraph id="8B843687EA582F7043BCFC8BFEEEFC14" blockId="5.[151,1436,876,959]" pageId="5" pageNumber="176">
|
||
<emphasis id="B94FEA95EA582F7043BCFC8BFE96FC2A" bold="true" box="[151,289,876,898]" pageId="5" pageNumber="176">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C3B4D04EA582F704202FC8BFE71FC29" authority="Graf & Pauls & Vitecek, 2018" authorityName="Graf & Pauls & Vitecek" authorityYear="2018" box="[297,454,876,898]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="vjosae" status="sp. nov.">
|
||
<emphasis id="B94FEA95EA582F704202FC8BFE71FC29" box="[297,454,876,898]" italics="true" pageId="5" pageNumber="176">Isoperla vjosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27C57EEEA582F7042E5FC8AFD95FC29" box="[462,546,877,898]" pageId="5" pageNumber="176" rank="species">
|
||
<emphasis id="B94FEA95EA582F7042E5FC8AFD95FC29" bold="true" box="[462,546,877,898]" pageId="5" pageNumber="176">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, lacinia;
|
||
<emphasis id="B94FEA95EA582F7041A8FC8BFCF6FC2A" bold="true" box="[643,833,876,899]" pageId="5" pageNumber="176">FIGURES 15–17.</emphasis>
|
||
<taxonomicName id="4C3B4D04EA582F704062FC8BFBD0FC29" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[841,1127,876,898]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94FEA95EA582F704062FC8BFBD0FC29" box="[841,1127,876,898]" italics="true" pageId="5" pageNumber="176">Isoperla tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
; 15: medial penial armatUres; 16: male meso- and metasternUm; 17: vesicle of sternite VIII;
|
||
<emphasis id="B94FEA95EA582F704026FC6CFC22FC0B" bold="true" box="[781,917,907,929]" pageId="5" pageNumber="176">FIGURE 18.</emphasis>
|
||
<taxonomicName id="4C3B4D04EA582F7040B7FC6CFB85FC0B" authorityName="Muranyi" authorityYear="2011" box="[924,1074,907,928]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="pesici">
|
||
<emphasis id="B94FEA95EA582F7040B7FC6CFB85FC0B" box="[924,1074,907,928]" italics="true" pageId="5" pageNumber="176">Isoperla pesici</emphasis>
|
||
</taxonomicName>
|
||
, medial penial armatUres. Scale bar represents 500 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B843687EA582F7043ECFBECFEE4F923" blockId="5.[151,1437,1035,1996]" pageId="5" pageNumber="176">
|
||
<emphasis id="B94FEA95EA582F7043ECFBECFEDDFB8F" bold="true" box="[199,362,1035,1060]" pageId="5" pageNumber="176">Mature larva</emphasis>
|
||
: Body length
|
||
<quantity id="4CC39B62EA582F704131FBECFD08FB88" box="[538,703,1035,1060]" metricMagnitude="-2" metricUnit="m" metricValue="1.085" metricValueMax="1.1199999999999999" metricValueMin="1.05" pageId="5" pageNumber="176" unit="mm" value="10.85" valueMax="11.2" valueMin="10.5">10.5-11.2 mm</quantity>
|
||
. General colour pale yellow with dark markings on head and abdomen (
|
||
<figureCitation id="13002A02EA582F70423BFBC8FED0FBEC" box="[272,359,1071,1096]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="5" pageNumber="176">Fig. 10</figureCitation>
|
||
); body and legs pilose, pilosity as typical for the genus; pronotal, posterior tergal and cercal fringes relatively long and acute; swimming hairs present on femora, tibiae and tarsi, cerci without dorsal hair fringe. Ground colour of the head pale yellow with dark brown transversal mask connecting eyes and ocelli and extending rostrally (
|
||
<figureCitation id="13002A02EA582F704255FB7CFE65FB18" box="[382,466,1179,1204]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="5" pageNumber="176">Fig. 11</figureCitation>
|
||
); M-line indistinct; occiput with row of lateral occipital setae, lateral occipital setae medially thinning. Eyes well-developed, not reduced. Scape, pedicel and antennomeres light brown, mouthparts light brown. Lacinia bidentate, triangular, inner lacinial margin with seven to eight stout setae and a row of short thin setae below subapical tooth (
|
||
<figureCitation id="13002A02EA582F704124FAE0FDD4FAB4" box="[527,611,1287,1312]" captionStart="FIGURE 14" captionStartId="5.[151,250,876,898]" captionTargetBox="[151,1436,193,854]" captionTargetId="figure@5.[151,1436,193,855]" captionTargetPageId="5" captionText="FIGURE 14. Isoperla vjosae sp. nov., lacinia; FIGURES 15–17. Isoperla tripartita tripartita; 15: medial penial armatUres; 16: male meso- and metasternUm; 17: vesicle of sternite VIII; FIGURE 18. Isoperla pesici, medial penial armatUres.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143929/files/figure.png" pageId="5" pageNumber="176">Fig. 14</figureCitation>
|
||
). Pronotum rounded, pale with obscure darker patterns laterally; pronotal anterior and posterior margin slightly darker, interrupted medially; pronotal setal fringe with predominantly short bristles at anterior and lateral margin, at posterior margin with additional large setae. Prosternum inconspicuous. Mesonotum and metanotum both pale with a brownish line anteriorly and posteriorly; wingpads whitish, with brown spots proximally. Meso- and metasternum pale, furcasternites and furcal pits slightly darker. Legs pale. Abdominal tergites I and II dorsally pale yellow with three longitudinal brown bands, abdominal tergites III to VI dorsally entirely brown, abdominal tergites VII to X dorsally with expanding pale areas leaving some brown markings medially and laterally. Ventral surface of abdomen pale yellow proximally, continually darkens distally. Paraprocts and cerci uniform pale; setation on distal section of cercal segments consisting of rather uniform setae with larger single dorsomedial and ventromedial setae, from cercal segment XIV with dense lateral fringe of swimming hairs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321650CEA582F7343ECF974FA2BF847" lastPageId="6" lastPageNumber="177" pageId="5" pageNumber="176" type="etymology">
|
||
<paragraph id="8B843687EA582F7043ECF974FE42F97B" blockId="5.[151,1437,1035,1996]" pageId="5" pageNumber="176">
|
||
<emphasis id="B94FEA95EA582F7043ECF974FEF9F907" bold="true" box="[199,334,1683,1708]" pageId="5" pageNumber="176">Etymology.</emphasis>
|
||
Named after the Vjosa River; Latinised Shqip (Albanian) “Vjosë” to genitive “
|
||
<taxonomicName id="4C3B4D04EA582F704637F974FADFF907" authorityName="Graf & Pauls & Vitecek" authorityYear="2018" box="[1308,1384,1683,1708]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="vjosae">vjosae</taxonomicName>
|
||
” — «belonging to the river Vjosa».
|
||
</paragraph>
|
||
<paragraph id="8B843687EA582F7043ECF93CFEF1F860" blockId="5.[151,1437,1035,1996]" pageId="5" pageNumber="176">
|
||
<emphasis id="B94FEA95EA582F7043ECF93CFE8BF95F" bold="true" box="[199,316,1755,1780]" pageId="5" pageNumber="176">AffinitieS.</emphasis>
|
||
The new species is a member of the
|
||
<taxonomicName id="4C3B4D04EA582F7041C4F93BFCC7F958" authorityName="Illies" authorityYear="1954" box="[751,880,1756,1779]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="tripartita">
|
||
<emphasis id="B94FEA95EA582F7041C4F93BFCC7F958" box="[751,880,1756,1779]" italics="true" pageId="5" pageNumber="176">I. tripartita</emphasis>
|
||
</taxonomicName>
|
||
group based on characters defined by
|
||
<bibRefCitation id="EFAA4B76EA582F704604F93CFF56F8B3" author="Consiglio" pageId="5" pageNumber="176" refString="Consiglio, C. (1967) Una nUova specie Italiana e considerazioni sUi grUppi di specie nel genere Isoperla (Plecoptera, Perlodidae). Fragmenta Entomologica, 5, 67 - 75." type="journal article" year="1967">Consiglio (1967)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAA4B76EA582F704233F918FE7AF8BC" author="Muranyi" box="[280,461,1791,1816]" pageId="5" pageNumber="176" refString="MUranyi, D. (2011) Balkanian species of the genUs Isoperla Banks, 1906 (Plecoptera: Perlodidae). Zootaxa, 3049, 1 - 46." type="journal article" year="2011">Murányi (2011)</bibRefCitation>
|
||
. The aedeagus of the new species possesses the typical divided medial penial armature located on the ventral lobe of the penis; this armature may be divided into a distal and proximal portion where usually one of these is at least partly divided into a left and a right half. Lateral penial armatures are usually present. The following species are currently considered members of this group:
|
||
<emphasis id="B94FEA95EA582F7040E7F88CFAA5F828" box="[972,1298,1899,1923]" italics="true" pageId="5" pageNumber="176">
|
||
<taxonomicName id="4C3B4D04EA582F7040E7F88CFABBF828" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[972,1292,1899,1923]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">Isoperla tripartita tripartita</taxonomicName>
|
||
;
|
||
</emphasis>
|
||
<taxonomicName id="4C3B4D04EA582F704637F88BFEDCF80C" authority="Zwick, 1978" authorityName="Zwick" authorityYear="1978" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="recta">
|
||
<emphasis id="B94FEA95EA582F704637F88BFF67F80C" italics="true" pageId="5" pageNumber="176">I. tripartita recta</emphasis>
|
||
<bibRefCitation id="EFAA4B76EA582F7043F2F868FEDCF80C" author="Zwick" box="[217,363,1935,1960]" pageId="5" pageNumber="176" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aUs Griechenland Und benachbarten Landern - 2. Teil. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 51, 213 - 239." type="journal article" year="1978">Zwick, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C3B4D04EA582F704252F877FDA7F80C" authorityName="Muranyi" authorityYear="2011" box="[377,528,1935,1959]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="B94FEA95EA582F704252F877FDA7F80C" box="[377,528,1935,1959]" italics="true" pageId="5" pageNumber="176">I. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA582F70410BF877FCF2F80C" authority="Tabacaru, 1971" authorityName="Tabacaru" authorityYear="1971" box="[544,837,1935,1960]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="illyrica">
|
||
<emphasis id="B94FEA95EA582F70410BF877FD3CF80C" box="[544,651,1935,1959]" italics="true" pageId="5" pageNumber="176">I. illyrica</emphasis>
|
||
Tabacaru, 1971
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C3B4D04EA582F704078F877FBE8F80C" authority="Zwick, 1978" authorityName="Zwick" authorityYear="1978" box="[851,1119,1935,1960]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="obliqua">
|
||
<emphasis id="B94FEA95EA582F704078F877FC74F80C" box="[851,963,1935,1959]" italics="true" pageId="5" pageNumber="176">I. obliqua</emphasis>
|
||
<bibRefCitation id="EFAA4B76EA582F7040E7F868FBE8F80C" author="Zwick" box="[972,1119,1935,1960]" pageId="5" pageNumber="176" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aUs Griechenland Und benachbarten Landern - 2. Teil. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 51, 213 - 239." type="journal article" year="1978">Zwick, 1978</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C3B4D04EA582F7047B5F877FB4EF80C" authorityName="Muranyi" authorityYear="2011" box="[1182,1273,1936,1959]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="pesici">
|
||
<emphasis id="B94FEA95EA582F7047B5F877FB4EF80C" box="[1182,1273,1936,1959]" italics="true" pageId="5" pageNumber="176">I. pesici</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAA4B76EA582F704621F868FA2FF80C" author="Zwick" box="[1290,1432,1935,1960]" pageId="5" pageNumber="176" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aUs Griechenland Und benachbarten Landern - 2. Teil. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 51, 213 - 239." type="journal article" year="1978">Zwick 1978</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFAA4B76EA582F7043BCF854FE8DF860" author="Muranyi" box="[151,314,1971,1996]" pageId="5" pageNumber="176" refString="MUranyi, D. (2011) Balkanian species of the genUs Isoperla Banks, 1906 (Plecoptera: Perlodidae). Zootaxa, 3049, 1 - 46." type="journal article" year="2011">Murányi 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF44660FEA5B2F7343BCFB7DFA3FFB1B" box="[151,1416,1178,1200]" httpUri="https://zenodo.org/record/1143931/files/figure.png" pageId="6" pageNumber="177" targetBox="[151,1436,313,1157]" targetPageId="6">
|
||
<paragraph id="8B843687EA5B2F7343BCFB7DFA3FFB1B" blockId="6.[151,1416,1178,1200]" box="[151,1416,1178,1200]" pageId="6" pageNumber="177">
|
||
<emphasis id="B94FEA95EA5B2F7343BCFB7DFEA8FB04" bold="true" box="[151,287,1178,1200]" pageId="6" pageNumber="177">FIGURE 19.</emphasis>
|
||
Sampling locality of the type material series at the river Vjosa near KUtë. Copyrighted image by Gregor ŠUbic.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B843687EA5B2F7343ECFB38FAC9FA07" blockId="6.[151,1436,1247,2029]" pageId="6" pageNumber="177">
|
||
Following
|
||
<bibRefCitation id="EFAA4B76EA5B2F734268FB38FE40FB53" author="Muranyi" box="[323,503,1247,1272]" pageId="6" pageNumber="177" refString="MUranyi, D. (2011) Balkanian species of the genUs Isoperla Banks, 1906 (Plecoptera: Perlodidae). Zootaxa, 3049, 1 - 46." type="journal article" year="2011">Murányi (2011)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAA4B76EA5B2F734119FB38FC9CFB53" author="Zwick" box="[562,811,1247,1272]" pageId="6" pageNumber="177" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aUs Griechenland Und benachbarten Landern - 2. Teil. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 51, 213 - 239." type="journal article" year="1978">Zwick (1978, fig. 72)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5B2F734012FB06FC2DFB5C" authorityName="Graf & Pauls & Vitecek" authorityYear="2018" box="[825,922,1248,1272]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5B2F734012FB06FC2DFB5C" box="[825,922,1248,1272]" italics="true" pageId="6" pageNumber="177">I. vjosae</emphasis>
|
||
</taxonomicName>
|
||
can be easily separated from all taxa of this group except
|
||
<taxonomicName id="4C3B4D04EA5B2F734205FAE2FE21FAB7" authorityName="Tabacaru" authorityYear="1971" box="[302,406,1284,1308]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="illyrica">
|
||
<emphasis id="B94FEA95EA5B2F734205FAE2FE21FAB7" box="[302,406,1284,1308]" italics="true" pageId="6" pageNumber="177">I. illyrica</emphasis>
|
||
</taxonomicName>
|
||
by the distinct medial gap between the proximal medial armatures (
|
||
<figureCitation id="13002A02EA5B2F7347A6FAE3FB41FAB7" box="[1165,1270,1284,1309]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="6" pageNumber="177">Figs. 7, 8</figureCitation>
|
||
) that are fused in
|
||
<taxonomicName id="4C3B4D04EA5B2F73439FFACEFE14FA94" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" box="[180,419,1320,1344]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94FEA95EA5B2F73439FFACEFE14FA94" box="[180,419,1320,1344]" italics="true" pageId="6" pageNumber="177">I. tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
(Fig. 15),
|
||
<taxonomicName id="4C3B4D04EA5B2F734137FACEFD6BFAEB" authorityName="Zwick" authorityYear="1978" box="[540,732,1320,1344]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="recta">
|
||
<emphasis id="B94FEA95EA5B2F734137FACEFD6BFAEB" box="[540,732,1320,1344]" italics="true" pageId="6" pageNumber="177">I. tripartita recta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5B2F7341C0FACEFCEFFAEB" authorityName="Zwick" authorityYear="1978" box="[747,856,1320,1344]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="obliqua">
|
||
<emphasis id="B94FEA95EA5B2F7341C0FACEFCEFFAEB" box="[747,856,1320,1344]" italics="true" pageId="6" pageNumber="177">I. obliqua</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5B2F734043FACEFC76FA94" authorityName="Muranyi" authorityYear="2011" box="[872,961,1320,1344]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="pesici">
|
||
<emphasis id="B94FEA95EA5B2F734043FACEFC76FA94" box="[872,961,1320,1344]" italics="true" pageId="6" pageNumber="177">I. pesici</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D04EA5B2F7340D1FACEFB39FAEB" authorityName="Muranyi" authorityYear="2011" box="[1018,1166,1320,1344]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="B94FEA95EA5B2F7340D1FACEFB39FAEB" box="[1018,1166,1320,1344]" italics="true" pageId="6" pageNumber="177">I. autumnalis</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C3B4D04EA5B2F734796FACEFAA1FA94" authorityName="Muranyi" authorityYear="2011" box="[1213,1302,1320,1344]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="pesici">
|
||
<emphasis id="B94FEA95EA5B2F734796FACEFB71FAEB" box="[1213,1222,1321,1344]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F7347FEFACFFAA1FA94" box="[1237,1302,1320,1343]" italics="true" pageId="6" pageNumber="177">pesici</emphasis>
|
||
</taxonomicName>
|
||
, the medial section of the proximal medial armature can be constricted in some populations, but is never as clearly separated as in
|
||
<taxonomicName id="4C3B4D04EA5B2F73439EFA96FE88FA2C" authority="sp." authorityName="Sp." box="[181,319,1392,1416]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5B2F73439EFA96FF09FA23" box="[181,190,1393,1416]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F7343E4FA97FEA1FA2C" box="[207,278,1392,1415]" italics="true" pageId="6" pageNumber="177">vjosae</emphasis>
|
||
<taxonomicNameLabel id="A27C57EEEA5B2F734234FA97FE88FA2C" box="[287,319,1392,1415]" pageId="6" pageNumber="177" rank="species">sp.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
n. (pers. comm. D. Murányi). In
|
||
<taxonomicName id="4C3B4D04EA5B2F7341EAFA96FC9BFA23" authorityName="Tabacaru" authorityYear="1971" box="[705,812,1392,1416]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="illyrica">
|
||
<emphasis id="B94FEA95EA5B2F7341EAFA96FC9BFA23" box="[705,812,1392,1416]" italics="true" pageId="6" pageNumber="177">I. illyrica</emphasis>
|
||
</taxonomicName>
|
||
the distal medial armatures are not divided while in
|
||
<taxonomicName id="4C3B4D04EA5B2F7346A7FA96FEB2FA07" authority="sp." authorityName="Sp." class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5B2F7346A7FA96FA22FA23" box="[1420,1429,1393,1416]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F7343BCFA72FF68FA07" box="[151,223,1429,1452]" italics="true" pageId="6" pageNumber="177">vjosae</emphasis>
|
||
<taxonomicNameLabel id="A27C57EEEA5B2F7343CEFA72FEB2FA07" box="[229,261,1429,1452]" pageId="6" pageNumber="177" rank="species">sp.</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
n. left and right portions of both distal and proximal parts are well separated (
|
||
<bibRefCitation id="EFAA4B76EA5B2F734744FA73FAC6FA07" author="Zwick" box="[1135,1393,1428,1453]" pageId="6" pageNumber="177" refString="Zwick, P. (1978) Steinfliegen (Plecoptera) aUs Griechenland Und benachbarten Landern - 2. Teil. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 51, 213 - 239." type="journal article" year="1978" yearSuffix="a">Zwick 1978; Abb. 72a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5B2F7343ECFA50FAABF997" blockId="6.[151,1436,1247,2029]" pageId="6" pageNumber="177">
|
||
Additionally the new species differs from
|
||
<taxonomicName id="4C3B4D04EA5B2F73418CFA5EFCDFFA7B" authorityName="Zwick" authorityYear="1978" box="[679,872,1464,1488]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="recta">
|
||
<emphasis id="B94FEA95EA5B2F73418CFA5EFCDFFA7B" box="[679,872,1464,1488]" italics="true" pageId="6" pageNumber="177">I. tripartita recta</emphasis>
|
||
</taxonomicName>
|
||
by having divergent left and right portions of the distal medial penial armature (
|
||
<figureCitation id="13002A02EA5B2F7342D3FA3BFDD0FA5F" box="[504,615,1500,1525]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="6" pageNumber="177">Figs. 7, 8</figureCitation>
|
||
) instead of parallel ones.
|
||
<taxonomicName id="4C3B4D04EA5B2F7340BCFA3BFBE2FA5F" authorityName="Zwick" authorityYear="1978" box="[919,1109,1500,1524]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="obliqua">
|
||
<emphasis id="B94FEA95EA5B2F7340BCFA3BFBE2FA5F" box="[919,1109,1500,1524]" italics="true" pageId="6" pageNumber="177">Isoperla obliqua</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3B4D04EA5B2F7347BAFA3AFB5BFA5F" authorityName="Muranyi" authorityYear="2011" box="[1169,1260,1501,1524]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="pesici">
|
||
<emphasis id="B94FEA95EA5B2F7347BAFA3AFB5BFA5F" box="[1169,1260,1501,1524]" italics="true" pageId="6" pageNumber="177">I. pesici</emphasis>
|
||
</taxonomicName>
|
||
have teardropshaped or rounded distal medial penial armatures, respectively (Fig. 18), which are elongated in
|
||
<taxonomicName id="4C3B4D04EA5B2F7347F2F9E6FAD5F9B3" authority="Sp." authorityName="Sp." box="[1241,1378,1535,1560]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5B2F7347F2F9E6FB55F9B3" box="[1241,1250,1537,1560]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F7347DAF9E7FA8FF9BC" box="[1265,1336,1536,1559]" italics="true" pageId="6" pageNumber="177">vjosae</emphasis>
|
||
<emphasis id="B94FEA95EA5B2F73466BFA18FAD5F9B3" bold="true" box="[1344,1378,1535,1560]" pageId="6" pageNumber="177">
|
||
<taxonomicNameLabel id="A27C57EEEA5B2F73466BFA18FAD5F9B3" box="[1344,1378,1535,1560]" pageId="6" pageNumber="177" rank="species">Sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94FEA95EA5B2F734642FA18FA2BF9B3" bold="true" box="[1385,1436,1535,1560]" pageId="6" pageNumber="177">nov.</emphasis>
|
||
(
|
||
<figureCitation id="13002A02EA5B2F7343B4F9C3FF53F997" box="[159,228,1572,1597]" captionStart="FIGURES 2–13" captionStartId="4.[151,264,1895,1918]" captionTargetBox="[184,1394,194,1867]" captionTargetId="figure@4.[183,1395,193,1868]" captionTargetPageId="4" captionText="FIGURES 2–13: Isoperla vjosae sp. nov.; 2: male habitUs; 3: male head and pronotUm, dorsal; 4: male meso- and metasternUm; 5: male abdominal tergites; 6: vesicle of sternite VIII; 7: everted male copUlatory organ with medial penial armatUres; 8: distal and proximal medial armatUres, arrow=lateral penial armatUres; 9: female sUbgenital plate; 10: larval habitUs; 11: larval headand pronotUm; 12:larval meso-and metathorax; 13: larval abdominal tergites.Scale bar represents 500 µm." httpUri="https://zenodo.org/record/1143925/files/figure.png" pageId="6" pageNumber="177">Fig. 8</figureCitation>
|
||
). Additionally,
|
||
<taxonomicName id="4C3B4D04EA5B2F7342B8F9C2FD90F997" authorityName="Muranyi" authorityYear="2011" box="[403,551,1572,1596]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="autumnalis">
|
||
<emphasis id="B94FEA95EA5B2F7342B8F9C2FE2BF997" box="[403,412,1573,1596]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F734281F9C3FD90F997" box="[426,551,1572,1596]" italics="true" pageId="6" pageNumber="177">autumnalis</emphasis>
|
||
</taxonomicName>
|
||
has an uncoloured and undivided proximal medial penial armature.
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5B2F7343ECF9A0FEC0F8BE" blockId="6.[151,1436,1247,2029]" pageId="6" pageNumber="177">
|
||
The colour patterns of both adults and larvae of the new species seem to deviate from colour patterns of other species of
|
||
<taxonomicName id="4C3B4D04EA5B2F734233F98BFE1AF92F" authorityName="tripartita Illies" authorityYear="1954" box="[280,429,1644,1669]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
the
|
||
<emphasis id="B94FEA95EA5B2F73426DF98AFE1AF92F" box="[326,429,1645,1668]" italics="true" pageId="6" pageNumber="177">tripartita</emphasis>
|
||
</taxonomicName>
|
||
group. However, colouration may vary specifically (e.g., as observed in
|
||
<taxonomicName id="4C3B4D04EA5B2F734633F98AFF49F90C" authority="tripartita" authorityName="tripartita Illies" authorityYear="1954" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
<emphasis id="B94FEA95EA5B2F734633F98AFA96F92F" box="[1304,1313,1645,1668]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F73461EF98AFF49F90C" italics="true" pageId="6" pageNumber="177">tripartita tripartita</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFAA4B76EA5B2F734227F968FE3FF903" author="Illies" box="[268,392,1679,1704]" pageId="6" pageNumber="177" refString="Illies, J. (1955) Isoperla tripartita n. sp., eine neUe Plecoptere aUs dem Wienerwald. Osterreichische Zoologische Zeitschrift, 5, 118 - 122." type="journal article" year="1955">Illies 1955</bibRefCitation>
|
||
, Murányi
|
||
<emphasis id="B94FEA95EA5B2F7342D1F976FD9CF903" box="[506,555,1680,1704]" italics="true" pageId="6" pageNumber="177">et al</emphasis>
|
||
. 2011), and we observed some variation in both larvae and adults although we did not extensively study this character set. Yet, overlapping colouration patterns within species might be indicative of on-going speciation processes, and could potentially be used as additional character to discern lineages within
|
||
<taxonomicName id="4C3B4D04EA5B2F7343BCF91BFE90F8BF" authorityName="tripartita Illies" authorityYear="1954" box="[151,295,1788,1813]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="171" phylum="Arthropoda" rank="subSpecies" species="tripartita" subSpecies="tripartita">
|
||
the
|
||
<emphasis id="B94FEA95EA5B2F7343EBF91AFE90F8BF" box="[192,295,1789,1812]" italics="true" pageId="6" pageNumber="177">tripartita</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5B2F7343ECF8F8FA2BF847" blockId="6.[151,1436,1247,2029]" pageId="6" pageNumber="177">
|
||
<emphasis id="B94FEA95EA5B2F7343ECF8F8FE4FF893" bold="true" box="[199,504,1823,1848]" pageId="6" pageNumber="177">ECology and diStribution.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="4C3B4D04EA5B2F734158F8C6FD64F89C" box="[627,723,1824,1848]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5B2F734158F8C6FDCBF893" box="[627,636,1825,1848]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F7341A7F8C7FD64F89C" box="[652,723,1824,1847]" italics="true" pageId="6" pageNumber="177">vjosae</emphasis>
|
||
</taxonomicName>
|
||
were collected along the banks of the Vjosa River (
|
||
<figureCitation id="13002A02EA5B2F73460EF8F8FACEF893" box="[1317,1401,1823,1848]" captionStart="FIGURE 19" captionStartId="6.[151,250,1178,1200]" captionTargetBox="[151,1436,313,1157]" captionTargetId="figure@6.[151,1436,193,1157]" captionTargetPageId="6" captionText="FIGURE 19. Sampling locality of the type material series at the river Vjosa near KUtë.Copyrighted image byGregor ŠUbic." httpUri="https://zenodo.org/record/1143931/files/figure.png" pageId="6" pageNumber="177">Fig. 19</figureCitation>
|
||
), a large, wide, shifting gravel, low altitude (
|
||
<quantity id="4CC39B62EA5B2F734151F8A3FD00F8F7" box="[634,695,1860,1885]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="177" unit="m" value="50.0">50 m</quantity>
|
||
a.s.l) river, a habitat atypical for other
|
||
<taxonomicName id="4C3B4D04EA5B2F7347ACF8A2FB59F8F7" authorityName="Illies" authorityYear="1954" box="[1159,1262,1861,1884]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="tripartita">
|
||
<emphasis id="B94FEA95EA5B2F7347ACF8A2FB59F8F7" box="[1159,1262,1861,1884]" italics="true" pageId="6" pageNumber="177">tripartita</emphasis>
|
||
</taxonomicName>
|
||
group species. Usually, taxa of this group occur in smaller submontane and montane streams and brooks (
|
||
<bibRefCitation id="EFAA4B76EA5B2F7347A6F880FA86F82B" author="Muranyi" box="[1165,1329,1895,1920]" pageId="6" pageNumber="177" refString="MUranyi, D. (2011) Balkanian species of the genUs Isoperla Banks, 1906 (Plecoptera: Perlodidae). Zootaxa, 3049, 1 - 46." type="journal article" year="2011">Murányi 2011</bibRefCitation>
|
||
, Murányi et al. 2016). Possibly, the high river-bedload dynamics of the Vjosa River create habitat conditions that are similar to conditions prevalent at higher elevations in regard to dissolved oxygen, sediment particle size distribution and bedload. As other members of the genus
|
||
<taxonomicName id="4C3B4D04EA5B2F73414CF833FD73F847" authorityName="Banks" authorityYear="1906" box="[615,708,2004,2028]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94FEA95EA5B2F73414CF833FD73F847" box="[615,708,2004,2028]" italics="true" pageId="6" pageNumber="177">Isoperla</emphasis>
|
||
</taxonomicName>
|
||
, larvae of
|
||
<taxonomicName id="4C3B4D04EA5B2F734014F832FC29F847" box="[831,926,2005,2028]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5B2F734014F832FCFFF847" box="[831,840,2005,2028]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
<emphasis id="B94FEA95EA5B2F73407CF832FC29F847" box="[855,926,2005,2028]" italics="true" pageId="6" pageNumber="177">vjosae</emphasis>
|
||
</taxonomicName>
|
||
are likely predaceous. At the
|
||
<typeStatus id="54808825EA5B2F7347D8F832FA94F846" box="[1267,1315,2005,2029]" pageId="6" pageNumber="177">type</typeStatus>
|
||
locality,
|
||
<emphasis id="B94FEA95EA5B2F7346A7F832FA22F847" box="[1420,1429,2005,2028]" italics="true" pageId="6" pageNumber="177">I</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C321650CEA5A2F7243BCFF7FFE57FD96" pageId="7" pageNumber="178" type="description">
|
||
<paragraph id="8B843687EA5A2F7243BCFF7FFBBCFE64" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="178">
|
||
<taxonomicName id="4C3B4D04EA5A2F7243BCFF7FFF68FF04" box="[151,223,152,175]" class="Insecta" family="Perlodidae" genus="Isoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vjosae">
|
||
<emphasis id="B94FEA95EA5A2F7243BCFF7FFF68FF04" box="[151,223,152,175]" italics="true" pageId="7" pageNumber="178">vjosae</emphasis>
|
||
</taxonomicName>
|
||
occurred sympatrically with the plecopterans
|
||
<taxonomicName id="4C3B4D04EA5A2F72403BFF7FFB0EFF1B" authority="(Burmeister)" authorityName="Burmeister" baseAuthorityName="Burmeister" baseAuthorityYear="1839" box="[784,1209,151,176]" class="Insecta" family="Perlidae" genus="Marthamea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vitripennis">
|
||
<emphasis id="B94FEA95EA5A2F72403BFF7FFBAEFF04" box="[784,1049,152,176]" italics="true" pageId="7" pageNumber="178">Marthamea vitripennis</emphasis>
|
||
(Burmeister)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F7247E0FF7FFF47FF7E" authority="(Kolbe)" authorityName="Kolbe" baseAuthorityName="Kolbe" baseAuthorityYear="1885" class="Insecta" family="Perlidae" genus="Eoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="ochracea">
|
||
<emphasis id="B94FEA95EA5A2F7247E0FF7FFA2BFF1B" box="[1227,1436,152,176]" italics="true" pageId="7" pageNumber="178">Eoperla ochracea</emphasis>
|
||
(Kolbe)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F72422BFF5BFDC1FF7F" authority="(Newman)" authorityName="Newman" baseAuthorityName="Newman" baseAuthorityYear="1836" box="[256,630,188,212]" class="Insecta" family="Chloroperlidae" genus="Xanthoperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis id="B94FEA95EA5A2F72422BFF5BFE47FF7F" box="[256,496,188,212]" italics="true" pageId="7" pageNumber="178">Xanthoperla apicalis</emphasis>
|
||
(Newman)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F7241AEFF5BFBBAFF7E" authority="(Scopoli)" authorityName="Scopoli" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[645,1037,188,213]" class="Insecta" family="Chloroperlidae" genus="Chloroperla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="tripunctata">
|
||
<emphasis id="B94FEA95EA5A2F7241AEFF5BFC20FF7F" box="[645,919,188,212]" italics="true" pageId="7" pageNumber="178">Chloroperla tripunctata</emphasis>
|
||
(Scopoli)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F724737FF5BFB37FF7F" authorityName="Banks" authorityYear="1903" box="[1052,1152,188,212]" class="Insecta" family="Perlodidae" genus="Perlodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94FEA95EA5A2F724737FF5BFB37FF7F" box="[1052,1152,188,212]" italics="true" pageId="7" pageNumber="178">Perlodes</emphasis>
|
||
</taxonomicName>
|
||
cf.
|
||
<emphasis id="B94FEA95EA5A2F724798FF5BFABDFF7F" box="[1203,1290,188,212]" italics="true" pageId="7" pageNumber="178">floridus</emphasis>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F724636FF5AFAC3FF7F" box="[1309,1396,189,212]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94FEA95EA5A2F724636FF5AFAC3FF7F" box="[1309,1396,189,212]" italics="true" pageId="7" pageNumber="178">Leuctra</emphasis>
|
||
</taxonomicName>
|
||
cf.
|
||
<emphasis id="B94FEA95EA5A2F7243BCFF07FEA2FF53" box="[151,277,224,248]" italics="true" pageId="7" pageNumber="178">leptogaster</emphasis>
|
||
and
|
||
<taxonomicName id="4C3B4D04EA5A2F724278FF06FDCFFF53" authority="(Linnaeus)" authorityName="Linnaeus" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[339,632,223,248]" class="Insecta" family="Leuctridae" genus="Leuctra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="fusca">
|
||
<emphasis id="B94FEA95EA5A2F724278FF06FE47FF53" box="[339,496,224,248]" italics="true" pageId="7" pageNumber="178">Leuctra fusca</emphasis>
|
||
(Linnaeus)
|
||
</taxonomicName>
|
||
, the
|
||
<taxonomicName id="4C3B4D04EA5A2F72419FFF38FC81FF53" box="[692,822,223,248]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
|
||
species
|
||
<taxonomicName id="4C3B4D04EA5A2F7240B6FF07FA2BFF5C" authority="Malicky, Agapetus" authorityName="Malicky, Agapetus" box="[925,1436,223,248]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="diakoftensis">
|
||
<emphasis id="B94FEA95EA5A2F7240B6FF07FB0CFF53" box="[925,1211,224,248]" italics="true" pageId="7" pageNumber="178">Rhyacophila diakoftensis</emphasis>
|
||
Malicky,
|
||
<emphasis id="B94FEA95EA5A2F72461FFF07FA2BFF5C" box="[1332,1436,224,247]" italics="true" pageId="7" pageNumber="178">Agapetus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94FEA95EA5A2F7243BCFEE3FF5FFEB7" box="[151,232,260,284]" italics="true" pageId="7" pageNumber="178">laniger</emphasis>
|
||
Pictet,
|
||
<taxonomicName id="4C3B4D04EA5A2F72427BFEE3FD01FEB6" authority="Raciecka" authorityName="Raciecka" authorityYear="1937" box="[336,694,260,285]" class="Insecta" family="Hydroptilidae" genus="Allotrichia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="vilnensis">
|
||
<emphasis id="B94FEA95EA5A2F72427BFEE3FD8AFEB7" box="[336,573,260,284]" italics="true" pageId="7" pageNumber="178">Allotrichia vilnensis</emphasis>
|
||
Raciecka
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F7241E7FEE3FBB4FEB7" authority="Morton" authorityName="Morton" authorityYear="1893" box="[716,1027,260,284]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="falcata">
|
||
<emphasis id="B94FEA95EA5A2F7241E7FEE3FC2CFEB7" box="[716,923,260,284]" italics="true" pageId="7" pageNumber="178">Oxyethira falcata</emphasis>
|
||
Morton
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F72473CFEE3FADBFEB6" authority="Mosely" authorityName="Mosely" authorityYear="1920" box="[1047,1388,260,285]" class="Insecta" family="Hydroptilidae" genus="Hydroptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="simulans">
|
||
<emphasis id="B94FEA95EA5A2F72473CFEE3FAB1FEB7" box="[1047,1286,260,284]" italics="true" pageId="7" pageNumber="178">Hydroptila simulans</emphasis>
|
||
Mosely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F7246AAFEE2FED3FEEB" authority="Mosely" authorityName="Mosely" authorityYear="1939" class="Insecta" family="Hydroptilidae" genus="Hydroptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="angustata">
|
||
<emphasis id="B94FEA95EA5A2F7246AAFEE2FEB2FEEB" italics="true" pageId="7" pageNumber="178">H. angustata</emphasis>
|
||
Mosely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F724258FECEFDD5FEEB" authority="Malicky" authorityName="Malicky" authorityYear="1996" box="[371,610,295,320]" class="Insecta" family="Hydroptilidae" genus="Hydroptila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="brissaga">
|
||
<emphasis id="B94FEA95EA5A2F724258FECEFE4EFEEB" box="[371,505,296,320]" italics="true" pageId="7" pageNumber="178">H. brissaga</emphasis>
|
||
Malicky
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F724144FECFFCC0FEEB" authority="Felber" authorityName="Felber" box="[623,887,295,320]" class="Insecta" family="Hydroptilidae" genus="Stactobiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="risi">
|
||
<emphasis id="B94FEA95EA5A2F724144FECFFC94FE94" box="[623,803,296,320]" italics="true" pageId="7" pageNumber="178">Stactobiella risi</emphasis>
|
||
Felber
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3B4D04EA5A2F7240AEFECFFB26FEEB" authority="Ulmer" authorityName="Ulmer" authorityYear="1907" box="[901,1169,295,320]" class="Insecta" family="Hydroptilidae" genus="Adicella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="syriaca">
|
||
<emphasis id="B94FEA95EA5A2F7240AEFECFFBF7FE94" box="[901,1088,296,320]" italics="true" pageId="7" pageNumber="178">Adicella syriaca</emphasis>
|
||
Ulmer
|
||
</taxonomicName>
|
||
and among others, the ephemeropteran,
|
||
<taxonomicName id="4C3B4D04EA5A2F724270FEAAFD6DFECE" authority="Muller" authorityName="Muller" box="[347,730,332,357]" class="Insecta" family="Prosopistomatidae" genus="Prosopistoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="178" phylum="Arthropoda" rank="species" species="pennigerum">
|
||
<emphasis id="B94FEA95EA5A2F724270FEAAFD32FECF" box="[347,645,333,356]" italics="true" pageId="7" pageNumber="178">Prosopistoma pennigerum</emphasis>
|
||
Müller
|
||
</taxonomicName>
|
||
- a community that clearly reflects a highly dynamic habitat. Many of these species are endangered or extirpated in Central Europe due to their reliance on no longer existing highly dynamic gravel reaches of large rivers. Despite repeated sampling efforts, the species could not be collected in July, September or October, indicating a short flight period in early summer.
|
||
</paragraph>
|
||
<paragraph id="8B843687EA5A2F7243ECFE3BFE57FD96" blockId="7.[151,1437,151,573]" pageId="7" pageNumber="178">
|
||
<emphasis id="B94FEA95EA5A2F7243ECFE3BFDB3FE5E" bold="true" box="[199,516,476,501]" pageId="7" pageNumber="178">RemarKS on ConServation.</emphasis>
|
||
The
|
||
<typeStatus id="54808825EA5A2F72416AFE3AFDC6FE5E" box="[577,625,477,501]" pageId="7" pageNumber="178">type</typeStatus>
|
||
locality of this species has been selected as a site for a dam and large hydropower plant. Construction of this hydropower plant will likely extirpate the characteristic benthic community currently inhabiting the river.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |