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<document id="ADECC3B4825A2E0298BF0F657023B575" ID-DOI="10.1046/j.1096-3642.2002.00039.x" ID-GBIF-Dataset="8acc9095-eaff-47d7-b3da-91b6c2fb636e" ID-ISSN="0024-4082" ID-Zenodo-Dep="4634200" IM.tables_requiresApprovalFor="existingObjects" IM.treatments_approvedBy="admin" checkinTime="1616618193905" checkinUser="carolina" docAuthor="Valdés, Ángel" docDate="2002" docId="03927F0EFFDD607AFCE2FA4B6BBAD002" docLanguage="en" docName="ZoolJLinnSoc.136.535-636.pdf" docOrigin="Zoological Journal of the Linnean Society 136 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1046/j.1096-3642.2002.00039.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.1:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="1" docTitle="Peltodoris atromaculata BERGH 1880" docType="treatment" docVersion="7" lastPageNumber="586" masterDocId="FFAB0776FFEF6049FF87FFCE6F73D33D" masterDocTitle="A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)" masterLastPageNumber="636" masterPageNumber="535" pageNumber="585" updateTime="1692293512987" updateUser="admin" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6ACDA9ACF028A8E501D2D834134DD530">A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)</mods:title>
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<mods:namePart id="14556ECC3ADB75792E41BF57BD045B2E">Valdés, Ángel</mods:namePart>
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<mods:title id="FD7A9E3E84AE93DB06200D2E1D0796F1">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="A395F746F91A1C39B934C9AD917EC8CD">2002</mods:date>
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<mods:number id="E6ACD73DB55A82886E37AB6E52AA8A99">2002-12-31</mods:number>
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<mods:classification id="6F3E64DF4160F882C892D1DAB239E2F9">journal article</mods:classification>
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<treatment id="03927F0EFFDD607AFCE2FA4B6BBAD002" ID-DOI="http://doi.org/10.5281/zenodo.5110237" ID-GBIF-Taxon="183550727" ID-Zenodo-Dep="5110237" LSID="urn:lsid:plazi:treatment:03927F0EFFDD607AFCE2FA4B6BBAD002" httpUri="http://treatment.plazi.org/id/03927F0EFFDD607AFCE2FA4B6BBAD002" lastPageId="51" lastPageNumber="586" pageId="50" pageNumber="585">
<subSubSection id="C3219D93FFDD607BFCE2FA4B6A3CD6A1" box="[869,1359,1412,1436]" pageId="50" pageNumber="585" type="nomenclature">
<paragraph id="8B84CE18FFDD607BFCE2FA4B6A3CD6A1" blockId="50.[869,1359,1412,1468]" box="[869,1359,1412,1436]" pageId="50" pageNumber="585">
<heading id="D0CC7974FFDD607BFCE2FA4B6A3CD6A1" box="[869,1359,1412,1436]" centered="true" fontSize="9" level="2" pageId="50" pageNumber="585" reason="5">
<taxonomicName id="4C3BB59BFFDD607BFCE2FA4B6A3CD6A1" ID-CoL="76GDY" ID-ENA="120394" authority="BERGH, 1880" box="[869,1359,1412,1436]" class="Gastropoda" family="Discodorididae" genus="Peltodoris" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="585" phylum="Mollusca" rank="species" species="atromaculata">
<emphasis id="B94F120AFFDD607BFCE2FA4B6BDCD6A6" box="[869,1199,1413,1436]" italics="true" pageId="50" pageNumber="585">PELTODORIS ATROMACULATA</emphasis>
<bibRefCitation id="EFAAB3E9FFDD607BFB3FFA4A6A3CD6A1" author="Bergh R" box="[1208,1359,1412,1436]" pageId="50" pageNumber="585" pagination="1 - 78" refId="ref53977" refString="Bergh R. 1880. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 4, heft 2. Wiesbaden: Kreidel, 1 - 78, plates A - F." type="book chapter" year="1880">BERGH, 1880</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFDD607BFC71FA6A6BCDD681" box="[1014,1214,1444,1468]" pageId="50" pageNumber="585" type="description">
<paragraph id="8B84CE18FFDD607BFC71FA6A6BCDD681" blockId="50.[869,1359,1412,1468]" box="[1014,1214,1444,1468]" pageId="50" pageNumber="585">
(
<figureCitation id="1300D29DFFDD607BFC78FA6A6B11D681" box="[1023,1122,1444,1468]" captionStart="Figure 4" captionStartId="12.[142,220,1147,1166]" captionTargetBox="[142,1422,195,1115]" captionTargetId="figure-362@12.[142,1422,195,1115]" captionTargetPageId="12" captionText="Figure 4. Living animals. A, Doris pseudoargus (CASIZ 121105). B, Doris immonda (CASIZ 089023), photo by T. M. Gosliner. C, Doris granulosa (CASIZ 073536), photo by T. M. Gosliner. D, Discodoris boholiensis (CASIZ 083654), photo by T. M. Gosliner. E, Discodoris ketos, San Pedrillo, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, Thordisa rubescens (CASIZ 015860), photo by T. M. Gosliner. G, Aphelodoris antillensis (CASIZ 077289), photo by T. M. Gosliner. H, Peltodoris atromaculata (CASIZ 119474). I, Peltodoris nobilis, Monterey Bay, California, photo by A. Smith." figureDoi="http://doi.org/10.5281/zenodo.4634212" httpUri="https://zenodo.org/record/4634212/files/figure.png" pageId="50" pageNumber="585">FIGS 4H</figureCitation>
,
<figureCitation id="1300D29DFFDD607BFBE9FA6A6BFED681" box="[1134,1165,1444,1468]" captionStart="Figure 28" captionStartId="52.[142,220,1731,1750]" captionTargetBox="[142,1422,194,1696]" captionTargetId="figure-78@52.[142,1422,194,1698]" captionTargetPageId="52" captionText="Figure 28. Peltodoris atromaculata (CASIZ 072584), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 250 Mm. C, outer lateral teeth; scale bar = 150 Mm. D, dorsal tubercles; scale bar = 250 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634290" httpUri="https://zenodo.org/record/4634290/files/figure.png" pageId="50" pageNumber="585">28</figureCitation>
,
<figureCitation id="1300D29DFFDD607BFB1DFA6A6BC4D681" box="[1178,1207,1444,1468]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="50" pageNumber="585">29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFDD607BFCA1FA036A6CD6DE" box="[806,1311,1485,1507]" pageId="50" pageNumber="585" type="reference_group">
<paragraph id="8B84CE18FFDD607BFCA1FA036A6CD6DE" blockId="50.[806,1311,1485,1507]" box="[806,1311,1485,1507]" pageId="50" pageNumber="585">
<treatmentCitationGroup id="AB2BE936FFDD607BFCA1FA036A6CD6DE" box="[806,1311,1485,1507]" pageId="50" pageNumber="585">
<treatmentCitation id="0A9AE809FFDD607BFCA1FA036A68D6DE" author="Bergh R" box="[806,1307,1485,1507]" page="46" pageId="50" pageNumber="585" year="1880">
<taxonomicName id="4C3BB59BFFDD607BFCA1FA036A68D6DE" ID-CoL="76GDY" authority="Bergh, 1880: 45 - 46" authorityName="Bergh" authorityPageNumber="45 - 46" authorityYear="1880" box="[806,1307,1485,1507]" class="Gastropoda" family="Discodorididae" genus="Peltodoris" kingdom="Animalia" order="Nudibranchia" pageId="50" pageNumber="585" phylum="Mollusca" rank="species" species="atromaculata">
<emphasis id="B94F120AFFDD607BFCA1FA036B42D6DF" box="[806,1073,1485,1506]" italics="true" pageId="50" pageNumber="585">Peltodoris atromaculata</emphasis>
<bibRefCitation id="EFAAB3E9FFDD607BFBBDFA036A68D6DE" author="Bergh R" box="[1082,1307,1485,1507]" pageId="50" pageNumber="585" pagination="1 - 78" refId="ref53977" refString="Bergh R. 1880. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 4, heft 2. Wiesbaden: Kreidel, 1 - 78, plates A - F." type="book chapter" year="1880">Bergh, 1880: 45–46</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3219D93FFDD607AFCA1F9E76BBAD002" lastPageId="51" lastPageNumber="586" pageId="50" pageNumber="585" type="materials_examined">
<paragraph id="8B84CE18FFDD607BFCA1F9E76CB2D503" blockId="50.[806,1422,1577,1660]" box="[806,961,1577,1598]" pageId="50" pageNumber="585">
<emphasis id="B94F120AFFDD607BFCA1F9E76CB2D503" box="[806,961,1577,1598]" italics="true" pageId="50" pageNumber="585">Type material</emphasis>
</paragraph>
<paragraph id="8B84CE18FFDD607BFCA1F9866A2DD541" blockId="50.[806,1422,1577,1660]" pageId="50" pageNumber="585">
<materialsCitation id="3B53C445FFDD607BFCA1F9866A2DD541" ID-GBIF-Occurrence="3323414316" collectionCode="ZMUC" country="Italy" location="Naples" pageId="50" pageNumber="585" specimenCode="GAS-2054" specimenCount="1" typeStatus="syntype">
<typeStatus id="548070BAFFDD607BFCA1F9866CD1D560" box="[806,930,1608,1629]" pageId="50" pageNumber="585">SYNTYPE</typeStatus>
:
<location id="8EE498C3FFDD607BFC35F9866B70D563" LSID="urn:lsid:plazi:treatment:03927F0EFFDD607AFCE2FA4B6BBAD002:8EE498C3FFDD607BFC35F9866B70D563" box="[946,1027,1608,1630]" country="Italy" name="Naples" pageId="50" pageNumber="585">Naples</location>
,
<collectingCountry id="F32C8E88FFDD607BFB93F9866B3AD563" box="[1044,1097,1608,1630]" name="Italy" pageId="50" pageNumber="585">Italy</collectingCountry>
, spring of 1880, one specimen,
<quantity id="4CC363FDFFDD607BFCE1F9A96CC7D541" box="[870,948,1639,1660]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="50" pageNumber="585" unit="mm" value="34.0">34 mm</quantity>
preserved length (
<collectionCode id="ED2A56DDFFDD607BFB0AF9A96BA8D541" box="[1165,1243,1639,1660]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="50" pageNumber="585">ZMUC</collectionCode>
<specimenCode id="DB9D6663FFDD607BFB64F9A86A24D541" box="[1251,1367,1638,1660]" collectionCode="GAS-" pageId="50" pageNumber="585">GAS-2054</specimenCode>
)
</materialsCitation>
</paragraph>
<paragraph id="8B84CE18FFDD607BFCA1F90C6B77D5EA" blockId="50.[806,1423,1730,1906]" box="[806,1028,1730,1751]" pageId="50" pageNumber="585">
<emphasis id="B94F120AFFDD607BFCA1F90C6B77D5EA" box="[806,1028,1730,1751]" italics="true" pageId="50" pageNumber="585">Additional material</emphasis>
</paragraph>
<paragraph id="8B84CE18FFDD607AFCA1F92F6C71D3C5" blockId="50.[806,1423,1730,1906]" lastBlockId="51.[155,771,196,248]" lastPageId="51" lastPageNumber="586" pageId="50" pageNumber="585">
Islas Medas, La Escala, west coast of Gerona,
<collectingCountry id="F32C8E88FFDD607BFAC2F92F6AF9D5CA" box="[1349,1418,1761,1783]" name="Spain" pageId="50" pageNumber="585">Spain</collectingCountry>
, three specimens,
<quantity id="4CC363FDFFDD607BFC79F8CE6B05D428" box="[1022,1142,1792,1814]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="50" pageNumber="585" unit="mm" value="29.5" valueMax="34.0" valueMin="25.0">25–34 mm</quantity>
preserved length, leg.
<collectionCode id="ED2A56DDFFDD607BFCA1F8D16C45D409" box="[806,822,1823,1844]" name="Tavera, Department of Geology and Geophysics" pageId="50" pageNumber="585">T</collectionCode>
. M. Gosliner (
<collectionCode id="ED2A56DDFFDD607BFC63F8D16B42D409" box="[996,1073,1823,1844]" pageId="50" pageNumber="585">CASIZ</collectionCode>
099147). Cala Salada, Ibiza,
<collectingCountry id="F32C8E88FFDD607BFCA1F8F36C19D46E" box="[806,874,1853,1875]" name="Spain" pageId="50" pageNumber="585">Spain</collectingCountry>
, one specimen,
<quantity id="4CC363FDFFDD607BFBA7F8F36B1DD46E" box="[1056,1134,1853,1875]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="50" pageNumber="585" unit="mm" value="49.0">49 mm</quantity>
preserved length, leg. A. Valdés (
<collectionCode id="ED2A56DDFFDD607BFCF8F8926CBFD44C" box="[895,972,1884,1905]" pageId="50" pageNumber="585">CASIZ</collectionCode>
119474).
<quantity id="4CC363FDFFDD607BFBBEF8926B01D44F" box="[1081,1138,1884,1906]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="50" pageNumber="585" unit="km" value="1.0">1 km</quantity>
east of Caloura, Ilha São Miguel,
<collectingRegion id="49FF00FAFFDC607AFF7FFF0A6E35D3E4" box="[248,326,196,217]" country="Portugal" name="Acores" pageId="51" pageNumber="586">Azores</collectingRegion>
,
<collectingCountry id="F32C8E88FFDC607AFED4FF0A6EC4D3E7" box="[339,439,196,218]" name="Portugal" pageId="51" pageNumber="586">Portugal</collectingCountry>
, eight specimens,
<quantity id="4CC363FDFFDC607AFD0CFF0A6C70D3E4" box="[651,771,196,218]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="6.7" metricValueMin="4.4" pageId="51" pageNumber="586" unit="mm" value="55.5" valueMax="67.0" valueMin="44.0">44–67 mm</quantity>
preserved length, leg.
<collectionCode id="ED2A56DDFFDC607AFE1EFF2D6EDAD3C5" box="[409,425,227,248]" name="Tavera, Department of Geology and Geophysics" pageId="51" pageNumber="586">T</collectionCode>
. M. Gosliner (
<collectionCode id="ED2A56DDFFDC607AFDCDFF2D6DE4D3C5" box="[586,663,227,248]" pageId="51" pageNumber="586">CASIZ</collectionCode>
072584).
</paragraph>
<paragraph id="8B84CE18FFDC607AFF1CFEF06EF0D26E" blockId="51.[155,771,318,923]" box="[155,387,318,339]" pageId="51" pageNumber="586">
<emphasis id="B94F120AFFDC607AFF1CFEF06EF0D26E" box="[155,387,318,339]" italics="true" pageId="51" pageNumber="586">External morphology</emphasis>
</paragraph>
<paragraph id="8B84CE18FFDC607AFF1CFE936EA5D002" blockId="51.[155,771,318,923]" pageId="51" pageNumber="586">
The general colour of the living animals is whitish to pale cream (
<figureCitation id="1300D29DFFDC607AFEA2FEB26E0FD2AF" box="[293,380,380,402]" captionStart="Figure 4" captionStartId="12.[142,220,1147,1166]" captionTargetBox="[142,1422,195,1115]" captionTargetId="figure-362@12.[142,1422,195,1115]" captionTargetPageId="12" captionText="Figure 4. Living animals. A, Doris pseudoargus (CASIZ 121105). B, Doris immonda (CASIZ 089023), photo by T. M. Gosliner. C, Doris granulosa (CASIZ 073536), photo by T. M. Gosliner. D, Discodoris boholiensis (CASIZ 083654), photo by T. M. Gosliner. E, Discodoris ketos, San Pedrillo, Puntarenas, Costa Rica, photo by T. M. Gosliner. F, Thordisa rubescens (CASIZ 015860), photo by T. M. Gosliner. G, Aphelodoris antillensis (CASIZ 077289), photo by T. M. Gosliner. H, Peltodoris atromaculata (CASIZ 119474). I, Peltodoris nobilis, Monterey Bay, California, photo by A. Smith." figureDoi="http://doi.org/10.5281/zenodo.4634212" httpUri="https://zenodo.org/record/4634212/files/figure.png" pageId="51" pageNumber="586">Fig. 4H</figureCitation>
). There is a number of dark brown or black large patches distributed on the dorsum, varying in shape and size. The rhinophores and gill are white or pale cream. The branchial leaves have some small dark brown or black spots. The whole dorsum is covered with small, conical tubercles, which have spicules protruding on their dorsal surface (
<figureCitation id="1300D29DFFDC607AFF24FD9C6E75D154" box="[163,262,594,617]" captionStart="Figure 28" captionStartId="52.[142,220,1731,1750]" captionTargetBox="[142,1422,194,1696]" captionTargetId="figure-78@52.[142,1422,194,1698]" captionTargetPageId="52" captionText="Figure 28. Peltodoris atromaculata (CASIZ 072584), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 250 Mm. C, outer lateral teeth; scale bar = 150 Mm. D, dorsal tubercles; scale bar = 250 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634290" httpUri="https://zenodo.org/record/4634290/files/figure.png" pageId="51" pageNumber="586">Fig. 28D</figureCitation>
). The largest tubercles are situated in the central region of the body. The rhinophoral and branchial sheaths have tubercles similar to those of the rest of the dorsum. There are six tripinnate branchial leaves, forming a circle. The anal papilla is situated in the centre of the branchial circle of leaves. The rhinophores are elongate, having 22 lamellae in a 52-mm preserved length specimen.
</paragraph>
<paragraph id="8B84CE18FFDC607AFF33FC866EE3D0A6" blockId="51.[155,771,318,923]" pageId="51" pageNumber="586">
Ventrally there are two short and conical oral tentacles (
<figureCitation id="1300D29DFFDC607AFF75FCA86E21D040" box="[242,338,870,893]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29F</figureCitation>
). The anterior border of the foot is grooved and notched.
</paragraph>
<paragraph id="8B84CE18FFDC607AFF1CFC2F6F8DD0CB" blockId="51.[155,771,993,1904]" box="[155,254,993,1014]" pageId="51" pageNumber="586">
<emphasis id="B94F120AFFDC607AFF1CFC2F6F8DD0CB" box="[155,254,993,1014]" italics="true" pageId="51" pageNumber="586">Anatomy</emphasis>
</paragraph>
<paragraph id="8B84CE18FFDC607AFF1CFBCE6D1BD522" blockId="51.[155,771,993,1904]" pageId="51" pageNumber="586">
The posterior end of the glandular portion of the oral tube has six strong retractor muscles (
<figureCitation id="1300D29DFFDC607AFDC8FBD06DC2D708" box="[591,689,1054,1077]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29D</figureCitation>
) which attach to the body wall. The oval, muscular buccal bulb has two additional muscles attached; two long salivary glands connect with it at each side of the oesophageal junction. The buccal bulb is longer than the glandular portion of the oral tube. The labial cuticle is smooth. The radular formula is 22 ¥ 48.0.
<quantity id="4CC363FDFFDC607AFD33FB196D9DD7D1" box="[692,750,1238,1260]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" pageId="51" pageNumber="586" unit="in" value="48.0">48 in</quantity>
a 54-mm preserved length specimen. Rachidian teeth are absent. The inner lateral teeth are short, having a long, curved cusp and lacking denticles (
<figureCitation id="1300D29DFFDC607AFD13FAFC6D84D674" box="[660,759,1330,1353]" captionStart="Figure 28" captionStartId="52.[142,220,1731,1750]" captionTargetBox="[142,1422,194,1696]" captionTargetId="figure-78@52.[142,1422,194,1698]" captionTargetPageId="52" captionText="Figure 28. Peltodoris atromaculata (CASIZ 072584), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 250 Mm. C, outer lateral teeth; scale bar = 150 Mm. D, dorsal tubercles; scale bar = 250 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634290" httpUri="https://zenodo.org/record/4634290/files/figure.png" pageId="51" pageNumber="586">Fig. 28A</figureCitation>
). They also have a secondary, short and blunt cusp situated behind the main cusp. The teeth from the middle portion of the half-row are hamate, long and larger than those closer to the medial portion of the radula (
<figureCitation id="1300D29DFFDC607AFF24FA026E77D6DF" box="[163,260,1484,1506]" captionStart="Figure 28" captionStartId="52.[142,220,1731,1750]" captionTargetBox="[142,1422,194,1696]" captionTargetId="figure-78@52.[142,1422,194,1698]" captionTargetPageId="52" captionText="Figure 28. Peltodoris atromaculata (CASIZ 072584), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 250 Mm. C, outer lateral teeth; scale bar = 150 Mm. D, dorsal tubercles; scale bar = 250 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634290" httpUri="https://zenodo.org/record/4634290/files/figure.png" pageId="51" pageNumber="586">Fig. 28B</figureCitation>
). The outermost teeth are smaller and also smooth (
<figureCitation id="1300D29DFFDC607AFE87FA246E11D53C" box="[256,354,1514,1537]" captionStart="Figure 28" captionStartId="52.[142,220,1731,1750]" captionTargetBox="[142,1422,194,1696]" captionTargetId="figure-78@52.[142,1422,194,1698]" captionTargetPageId="52" captionText="Figure 28. Peltodoris atromaculata (CASIZ 072584), SEM images of the radula and dorsal tubercles. A, inner lateral teeth; scale bar = 75 Mm. B, mid-lateral teeth; scale bar = 250 Mm. C, outer lateral teeth; scale bar = 150 Mm. D, dorsal tubercles; scale bar = 250 Mm." figureDoi="http://doi.org/10.5281/zenodo.4634290" httpUri="https://zenodo.org/record/4634290/files/figure.png" pageId="51" pageNumber="586">Fig. 28C</figureCitation>
). The oesophagus is short and connects directly to the stomach (
<figureCitation id="1300D29DFFDC607AFE7EF9C76D2FD522" box="[505,604,1545,1567]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29A</figureCitation>
).
</paragraph>
<paragraph id="8B84CE18FFDC607AFF33F9E66B4ED20B" blockId="51.[155,771,993,1904]" lastBlockId="51.[819,1435,196,831]" pageId="51" pageNumber="586">
The ampulla is long and thin, and branches into a short oviduct and the prostate (
<figureCitation id="1300D29DFFDC607AFD81F9886D1AD560" box="[518,617,1606,1629]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29C</figureCitation>
). The oviduct enters the female gland mass near to its centre. The prostate is flattened, long, folded and granular (
<figureCitation id="1300D29DFFDC607AFF24F96C6E77D584" box="[163,260,1698,1721]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29B</figureCitation>
), with two differentiated portions distinguishable by their colour and texture. It connects with a long duct that narrows and expands again into the small ejaculatory portion of the deferent duct. The muscular deferent duct opens into a common atrium with the vagina. The vagina is long. Near to its proximal end it joins the bursa copulatrix. From the bursa copulatrix leads another duct that connects to the seminal receptacle and the uterine duct. The bursa copulatrix is oval in shape, about 10 times as large as the seminal receptacle.
</paragraph>
<paragraph id="8B84CE18FFDC607AFCCBFEF16A66D1DE" blockId="51.[819,1435,196,831]" pageId="51" pageNumber="586">
In the central nervous system (
<figureCitation id="1300D29DFFDC607AFB46FEF06A51D268" box="[1217,1314,318,341]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29E</figureCitation>
) the cerebral and pleural ganglia are fused and distinct from the pedal ganglia. There are four cerebral nerves leading from the left cerebral ganglion and three from the right one, and three pleural nerves leading from each pleural ganglion. There is a separate abdominal ganglion on the right side of the visceral loop. The buccal ganglia are near to the rest of the central nervous system, joined to the cerebral ganglia by two relatively long nerves. Gastro-oesophageal, rhinophoral and optical ganglia are present. The pedal ganglia are clearly separated, having four nerves leading from each one. The pedal and parapedal commissures are enveloped together with the visceral loop.
</paragraph>
<paragraph id="8B84CE18FFDC607AFCCBFD226BBAD002" blockId="51.[819,1435,196,831]" pageId="51" pageNumber="586">
The circulatory system (
<figureCitation id="1300D29DFFDC607AFBE7FD226BB2D03F" box="[1120,1217,748,770]" captionStart="Figure 29" captionStartId="53.[155,233,1819,1838]" captionTargetBox="[178,1407,201,1773]" captionTargetId="figure-92@53.[149,822,172,1047]" captionTargetPageId="53" captionText="Figure 29. Peltodoris atromaculata (CASIZ 072584). A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of several reproductive organs; scale bar = 1 mm. D, lateral view of the buccal bulb; scale bar = 1 mm. E, central nervous system; scale bar = 0.5 mm. F, ventral view of the mouth area; scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4634294" httpUri="https://zenodo.org/record/4634294/files/figure.png" pageId="51" pageNumber="586">Fig. 29A</figureCitation>
) consists of a large heart and a two blood glands situated in front of and behind the central nervous system.
</paragraph>
</subSubSection>
</treatment>
</document>
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