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<mods:title id="CA8B29D624B2FBB8DA36D711D5DCF55D">A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)</mods:title>
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<treatment id="03927F0EFFC8606EFCDFFEBD6B2DD44D" ID-DOI="http://doi.org/10.5281/zenodo.5108122" ID-Zenodo-Dep="5108122" LSID="urn:lsid:plazi:treatment:03927F0EFFC8606EFCDFFEBD6B2DD44D" httpUri="http://treatment.plazi.org/id/03927F0EFFC8606EFCDFFEBD6B2DD44D" lastPageNumber="574" pageId="39" pageNumber="574">
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<heading id="D0CC7974FFC8606EFCDFFEBD6A05D2B7" box="[856,1398,370,394]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="574" reason="5">
<taxonomicName id="4C3BB59BFFC8606EFCDFFEBD6A05D2B7" ID-CoL="7BYDH" authority="(Alder &amp; Hancock, 1864)" box="[856,1398,370,394]" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFCDFFEBD6B41D2B5" box="[856,1074,371,394]" italics="true" pageId="39" pageNumber="574">THORDISA VILLOSA</emphasis>
(
<bibRefCitation id="EFAAB3E9FFC8606EFBC6FEBC6A1DD2B7" author="ALDER &amp; HANCOCK" box="[1089,1390,370,394]" firstAuthor="ALDER" pageId="39" pageNumber="574" pagination="113 - 147" refId="ref53398" refString="Alder J, Hancock A. 1864. Notice of a collection of nudibranchiate Mollusca made in India by Water Elliot Esq., with descriptions of several new genera and species. Transactions of the Zoological Society of London 5: 113 - 147, plates 28 - 33." type="journal article" year="1864">ALDER &amp; HANCOCK, 1864</bibRefCitation>
)
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</heading>
</paragraph>
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<subSubSection id="C3219D93FFC8606EFCB4FE556BC7D116" pageId="39" pageNumber="574" type="reference_group">
<paragraph id="8B84CE18FFC8606EFCB4FE556CFAD2F3" blockId="39.[819,1435,410,555]" pageId="39" pageNumber="574">
<treatmentCitationGroup id="AB2BE936FFC8606EFCB4FE556CFAD2F3" pageId="39" pageNumber="574">
<treatmentCitation id="0A9AE809FFC8606EFCB4FE556A34D28D" author="Alder J &amp; Hancock A" box="[819,1351,410,432]" page="120" pageId="39" pageNumber="574" year="1864">
<taxonomicName id="4C3BB59BFFC8606EFCB4FE556A34D28D" authority="Alder &amp; Hancock, 1864: 119 - 120" authorityName="Alder &amp; Hancock" authorityPageNumber="119 - 120" authorityYear="1864" box="[819,1351,410,432]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFCB4FE556CB3D292" box="[819,960,410,432]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
<bibRefCitation id="EFAAB3E9FFC8606EFC4EFE546A34D28D" author="Alder &amp; Hancock" box="[969,1351,410,432]" firstAuthor="Alder" page="119 - 120" pageId="39" pageNumber="574" pagination="113 - 147" refId="ref53398" refString="Alder J, Hancock A. 1864. Notice of a collection of nudibranchiate Mollusca made in India by Water Elliot Esq., with descriptions of several new genera and species. Transactions of the Zoological Society of London 5: 113 - 147, plates 28 - 33." type="journal article" year="1864">Alder &amp; Hancock, 1864: 119120</bibRefCitation>
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, pl. 33, fig. 1.
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</paragraph>
<paragraph id="8B84CE18FFC8606EFCB4FE166B09D131" blockId="39.[819,1435,410,555]" pageId="39" pageNumber="574">
<treatmentCitationGroup id="AB2BE936FFC8606EFCB4FE166B09D131" pageId="39" pageNumber="574">
<treatmentCitation id="0A9AE809FFC8606EFCB4FE166A4CD2D3" author="Bergh R" box="[819,1343,472,494]" page="542" pageId="39" pageNumber="574" year="1877">
<taxonomicName id="4C3BB59BFFC8606EFCB4FE166A4CD2D3" ID-CoL="7BYDK" authority="Bergh, 1877 a: 540 - 542" authorityName="Bergh" authorityPageNumber="540 - 542" authorityYear="1877" box="[819,1343,472,494]" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="maculigera">
<emphasis id="B94F120AFFC8606EFCB4FE166B53D2D0" box="[819,1056,472,493]" italics="true" pageId="39" pageNumber="574">Thordisa maculigera</emphasis>
<bibRefCitation id="EFAAB3E9FFC8606EFBAAFE166A4CD2D3" author="Bergh" box="[1069,1343,472,494]" firstAuthor="Bergh" page="540 - 542" pageId="39" pageNumber="574" pagination="58 - 61" refId="ref53810" refString="Bergh R. 1877 a. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 2, heft 10. Wiesbaden: Kreidel, 495 - 546, plates 58 - 61." type="book chapter" year="1877" yearSuffix="a">Bergh, 1877a: 540542</bibRefCitation>
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, pl. 61, figs 1924, pl. 62, figs 1, 2.
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</paragraph>
<paragraph id="8B84CE18FFC8606EFCB4FDDB6BC7D116" blockId="39.[819,1435,410,555]" box="[819,1204,533,555]" pageId="39" pageNumber="574">
<treatmentCitationGroup id="AB2BE936FFC8606EFCB4FDDB6BC7D116" box="[819,1204,533,555]" pageId="39" pageNumber="574">
<treatmentCitation id="0A9AE809FFC8606EFCB4FDDB6BC3D116" author="Eliot CN" box="[819,1200,533,555]" page="368" pageId="39" pageNumber="574" year="1904">
<taxonomicName id="4C3BB59BFFC8606EFCB4FDDB6BC3D116" ID-CoL="56KL7" authority="Eliot, 1904: 368" authorityName="Eliot" authorityPageNumber="368" authorityYear="1904" box="[819,1200,533,555]" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="stellata">
<emphasis id="B94F120AFFC8606EFCB4FDDB6C82D117" box="[819,1009,533,554]" italics="true" pageId="39" pageNumber="574">Thordisa stellata</emphasis>
<bibRefCitation id="EFAAB3E9FFC8606EFC7EFDDB6BC3D116" author="Eliot" box="[1017,1200,533,555]" firstAuthor="Eliot" page="368" pageId="39" pageNumber="574" pagination="354 - 385" refId="ref55359" refString="Eliot CN. 1904. On some nudibranchs from east Africa and Zanzibar. Part III. Dorididae Cryptobranchiatae. Proceedings of the Zoological Society of London 1903 (2): 354 - 385, plates 32 - 34." type="journal article" year="1904">Eliot, 1904: 368</bibRefCitation>
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.
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<emphasis id="B94F120AFFC8606EFCB4FDBF6CBDD1BB" box="[819,974,625,646]" italics="true" pageId="39" pageNumber="574">Type material</emphasis>
</paragraph>
<paragraph id="8B84CE18FFC8606EFCB4FD5E6B77D041" blockId="39.[819,1435,625,892]" pageId="39" pageNumber="574">
<taxonomicName id="4C3BB59BFFC8606EFCB4FD5E6BE4D19B" authority="Alder &amp; Hancock" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[819,1175,656,678]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFCB4FD5E6CB1D198" box="[819,962,656,677]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
Alder &amp; Hancock
</taxonomicName>
,
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: Madras,
<collectingCountry id="F32C8E88FFC8606EFCB4FD606C01D1F9" box="[819,882,686,708]" name="India" pageId="39" pageNumber="574">India</collectingCountry>
, two specimens,
<quantity id="4CC363FDFFC8606EFBC9FD616BCBD1F9" box="[1102,1208,687,708]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="39" pageNumber="574" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">713 mm</quantity>
preserved length, dried, leg. W. Elliot (
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20, 42).
<taxonomicName id="4C3BB59BFFC8606EFB6FFD036CBCD03F" authority="Bergh" authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="maculigera">
<emphasis id="B94F120AFFC8606EFB6FFD036C00D03C" italics="true" pageId="39" pageNumber="574">Thordisa maculigera</emphasis>
Bergh
</taxonomicName>
,
<typeStatus id="548070BAFFC8606EFC6FFD226B0BD03C" box="[1000,1144,748,769]" pageId="39" pageNumber="574">HOLOTYPE</typeStatus>
(by monotypy):
<collectingRegion id="49FF00FAFFC8606EFADEFD226AE4D03F" box="[1369,1431,748,770]" country="Philippines" name="Cebu" pageId="39" pageNumber="574">Cebu</collectingRegion>
,
<collectingCountry id="F32C8E88FFC8606EFCB4FCC46CC6D01D" box="[819,949,778,800]" name="Philippines" pageId="39" pageNumber="574">Philippines</collectingCountry>
<quantity id="4CC363FDFFC8606EFC43FCC46B2CD01D" box="[964,1119,778,800]" metricMagnitude="0" metricUnit="m" metricValue="1.86415" pageId="39" pageNumber="574" unit="mm" value="1864.15">1864, 15 mm</quantity>
preserved length, leg. C. Semper (
<collectionCode id="ED2A56DDFFC8606EFC18FCE76C9ED003" box="[927,1005,809,830]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="39" pageNumber="574">ZMUC</collectionCode>
<collectionCode id="ED2A56DDFFC8606EFC79FCE76B4DD002" box="[1022,1086,809,831]" pageId="39" pageNumber="574">GAS-</collectionCode>
2102). The type material of
<taxonomicName id="4C3BB59BFFC8606EFCB0FC866B45D063" authority="Eliot" authorityName="Eliot" authorityYear="1904" box="[823,1078,840,862]" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="stellata">
<emphasis id="B94F120AFFC8606EFCB0FC866C85D060" box="[823,1014,840,861]" italics="true" pageId="39" pageNumber="574">Thordisa stellata</emphasis>
Eliot
</taxonomicName>
is untraceable; it could not be located in
<collectionCode id="ED2A56DDFFC8606EFC2CFCA96B73D041" box="[939,1024,871,892]" pageId="39" pageNumber="574">BMNH</collectionCode>
.
</paragraph>
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<subSubSection id="C3219D93FFC8606EFCB4FC0C6B2DD44D" pageId="39" pageNumber="574" type="discussion">
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<emphasis id="B94F120AFFC8606EFCB4FC0C6CE6D0EA" box="[819,917,962,983]" italics="true" pageId="39" pageNumber="574">Remarks</emphasis>
</paragraph>
<paragraph id="8B84CE18FFC8606EFCB4FC2F6BA6D636" blockId="39.[819,1437,962,1904]" pageId="39" pageNumber="574">
<bibRefCitation id="EFAAB3E9FFC8606EFCB4FC2F6B3DD0CA" author="Alder &amp; Hancock" box="[819,1102,993,1015]" firstAuthor="Alder" pageId="39" pageNumber="574" pagination="113 - 147" refId="ref53398" refString="Alder J, Hancock A. 1864. Notice of a collection of nudibranchiate Mollusca made in India by Water Elliot Esq., with descriptions of several new genera and species. Transactions of the Zoological Society of London 5: 113 - 147, plates 28 - 33." type="journal article" year="1864">Alder &amp; Hancock (1864)</bibRefCitation>
described
<taxonomicName id="4C3BB59BFFC8606EFB4AFC2C6A2FD0CB" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1229,1372,993,1015]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFB4AFC2C6A2FD0CB" box="[1229,1372,993,1015]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
</taxonomicName>
from
<collectingCountry id="F32C8E88FFC8606EFCB4FBCE6C01D72B" box="[819,882,1024,1046]" name="India" pageId="39" pageNumber="574">India</collectingCountry>
, based on two specimens having the dorsum covered with processes with filaments. The living animals were ochre yellow to orange with large brown botches surrounding the mantle margin.
<bibRefCitation id="EFAAB3E9FFC8606EFB6EFB926AE8D74F" author="Bergh" box="[1257,1435,1116,1138]" firstAuthor="Bergh" pageId="39" pageNumber="574" pagination="58 - 61" refId="ref53810" refString="Bergh R. 1877 a. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 2, heft 10. Wiesbaden: Kreidel, 495 - 546, plates 58 - 61." type="book chapter" year="1877" yearSuffix="a">Bergh (1877a)</bibRefCitation>
described
<taxonomicName id="4C3BB59BFFC8606EFC2DFBB46BE5D7B2" authorityName="Bergh" authorityYear="1877" box="[938,1174,1146,1167]" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="maculigera">
<emphasis id="B94F120AFFC8606EFC2DFBB46BE5D7B2" box="[938,1174,1146,1167]" italics="true" pageId="39" pageNumber="574">Thordisa maculigera</emphasis>
</taxonomicName>
from the
<collectingCountry id="F32C8E88FFC8606EFA95FBB46AE4D7AD" box="[1298,1431,1146,1168]" name="Philippines" pageId="39" pageNumber="574">Philippines</collectingCountry>
, but provided no information of the colour in the living animal. Years later,
<bibRefCitation id="EFAAB3E9FFC8606EFB91FB766BD9D7F3" author="Bergh" box="[1046,1194,1208,1230]" firstAuthor="Bergh" pageId="39" pageNumber="574" pagination="155 - 218" refId="ref54458" refString="Bergh R. 1902. I. Gastropoda Opisthobranchiata, 12. In: Mortensen T, ed. The Danish expedition to Siam 1899 - 1900. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling. Bogtrykkeri. Copenhagen, 155 - 218, plates 1 - 3." type="book chapter" year="1902">Bergh (1902)</bibRefCitation>
reported this species from the Gulf of
<collectingCountry id="F32C8E88FFC8606EFB86FB186B1AD7D1" box="[1025,1129,1238,1260]" name="Thailand" pageId="39" pageNumber="574">Thailand</collectingCountry>
and recognized that it is probably a synonym of
<taxonomicName id="4C3BB59BFFC8606EFBC5FB386BBED637" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1090,1229,1269,1291]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFBC5FB386BBED637" box="[1090,1229,1269,1291]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFC8606EFCCBFADA6B9AD5A4" blockId="39.[819,1437,962,1904]" pageId="39" pageNumber="574">
<bibRefCitation id="EFAAB3E9FFC8606EFCCBFADA6CA7D617" author="Eliot" box="[844,980,1300,1322]" firstAuthor="Eliot" pageId="39" pageNumber="574" pagination="354 - 385" refId="ref55359" refString="Eliot CN. 1904. On some nudibranchs from east Africa and Zanzibar. Part III. Dorididae Cryptobranchiatae. Proceedings of the Zoological Society of London 1903 (2): 354 - 385, plates 32 - 34." type="journal article" year="1904">Eliot (1904)</bibRefCitation>
redescribed
<taxonomicName id="4C3BB59BFFC8606EFBECFADA6B8AD614" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1131,1273,1300,1321]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFBECFADA6B8AD614" box="[1131,1273,1300,1321]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
</taxonomicName>
based on one specimen from East Africa, which clearly fits with the original description by
<bibRefCitation id="EFAAB3E9FFC8606EFBBEFA9F6A38D65A" author="Alder &amp; Hancock" box="[1081,1355,1361,1383]" firstAuthor="Alder" pageId="39" pageNumber="574" pagination="113 - 147" refId="ref53398" refString="Alder J, Hancock A. 1864. Notice of a collection of nudibranchiate Mollusca made in India by Water Elliot Esq., with descriptions of several new genera and species. Transactions of the Zoological Society of London 5: 113 - 147, plates 28 - 33." type="journal article" year="1864">Alder &amp; Hancock (1864)</bibRefCitation>
. At the same time, he synonymized it with
<taxonomicName id="4C3BB59BFFC8606EFB48FABE6C25D699" authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="maculigera">
<emphasis id="B94F120AFFC8606EFB48FABE6C25D699" italics="true" pageId="39" pageNumber="574">Thordisa maculigera</emphasis>
</taxonomicName>
, with some reservations due to differences in the radular morphology. Later,
<bibRefCitation id="EFAAB3E9FFC8606EFBF9FA636A65D6FE" author="Eliot" box="[1150,1302,1453,1475]" firstAuthor="Eliot" pageId="39" pageNumber="574" pagination="636 - 691" refId="ref55531" refString="Eliot CN. 1906 c. On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder &amp; Hancock preserved in the Hancock Museum at Newcastle-on-Tyne. Proceedings of the Zoological Society of London 1906 (2): 636 - 691. plates 42 - 47." type="journal article" year="1906" yearSuffix="c">Eliot (1906c)</bibRefCitation>
reaffirmed his opinion on the synonymy of
<taxonomicName id="4C3BB59BFFC8606EFB2FFA036A7CD6DC" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1192,1295,1484,1506]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFB2FFA036A7CD6DC" box="[1192,1295,1484,1506]" italics="true" pageId="39" pageNumber="574">D. villosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3BB59BFFC8606EFACCFA036CF1D6C2" authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="maculigera">
<emphasis id="B94F120AFFC8606EFACCFA036CF1D6C2" italics="true" pageId="39" pageNumber="574">T. maculigera</emphasis>
</taxonomicName>
, based on the examination of more specimens. He also examined the type material of
<taxonomicName id="4C3BB59BFFC8606EFA82F9C46AE7D523" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1285,1428,1545,1567]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFA82F9C46AE7D523" box="[1285,1428,1545,1567]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
</taxonomicName>
, but the two
<typeStatus id="548070BAFFC8606EFC41F9E66B58D500" box="[966,1067,1576,1597]" pageId="39" pageNumber="574">syntypes</typeStatus>
had the buccal mass removed, and comparison of the radular morphology was not possible.
<bibRefCitation id="EFAAB3E9FFC8606EFC1BF9AB6B5CD546" author="Eliot" box="[924,1071,1637,1659]" firstAuthor="Eliot" pageId="39" pageNumber="574" pagination="636 - 691" refId="ref55531" refString="Eliot CN. 1906 c. On the nudibranchs of Southern India and Ceylon, with special reference to the drawings by Kelaart and the collections belonging to Alder &amp; Hancock preserved in the Hancock Museum at Newcastle-on-Tyne. Proceedings of the Zoological Society of London 1906 (2): 636 - 691. plates 42 - 47." type="journal article" year="1906" yearSuffix="c">Eliot (1906c)</bibRefCitation>
also regarded
<taxonomicName id="4C3BB59BFFC8606EFB59F9AB6CC3D5A4" authority="Eliot, 1904" authorityName="Eliot" authorityYear="1904" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="stellata">
<emphasis id="B94F120AFFC8606EFB59F9AB6AEED547" box="[1246,1437,1637,1658]" italics="true" pageId="39" pageNumber="574">Thordisa stellata</emphasis>
<bibRefCitation id="EFAAB3E9FFC8606EFCB4F94A6CC3D5A4" author="Eliot" box="[819,944,1668,1690]" firstAuthor="Eliot" pageId="39" pageNumber="574" pagination="354 - 385" refId="ref55359" refString="Eliot CN. 1904. On some nudibranchs from east Africa and Zanzibar. Part III. Dorididae Cryptobranchiatae. Proceedings of the Zoological Society of London 1903 (2): 354 - 385, plates 32 - 34." type="journal article" year="1904">Eliot, 1904</bibRefCitation>
</taxonomicName>
as a synonym of
<taxonomicName id="4C3BB59BFFC8606EFBFBF94B6B92D5A4" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1148,1249,1668,1690]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFBFBF94B6B92D5A4" box="[1148,1249,1668,1690]" italics="true" pageId="39" pageNumber="574">D. villosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B84CE18FFC8606EFCCBF96C6B2DD44D" blockId="39.[819,1437,962,1904]" pageId="39" pageNumber="574">
Unfortunately I was unable to find complete specimens for this study. The
<typeStatus id="548070BAFFC8606EFBD2F90C6BC9D5EA" box="[1109,1210,1730,1751]" pageId="39" pageNumber="574">syntypes</typeStatus>
of
<taxonomicName id="4C3BB59BFFC8606EFB65F90C6A1CD5EB" authorityName="Alder &amp; Hancock" authorityYear="1864" box="[1250,1391,1729,1751]" class="Gastropoda" family="Dorididae" genus="Doris" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="villosa">
<emphasis id="B94F120AFFC8606EFB65F90C6A1CD5EB" box="[1250,1391,1729,1751]" italics="true" pageId="39" pageNumber="574">Doris villosa</emphasis>
</taxonomicName>
are poorly preserved, but I found the radula, mounted on a slide, in the
<collectionCode id="ED2A56DDFFC8606EFC5BF9316B5DD429" box="[988,1070,1791,1812]" pageId="39" pageNumber="574">HMNC</collectionCode>
collections. This radula is very similar to the drawings of the radula of
<taxonomicName id="4C3BB59BFFC8606EFB7DF8D36CF1D46C" authorityName="Bergh" authorityYear="1877" class="Gastropoda" family="Discodorididae" genus="Thordisa" kingdom="Animalia" order="Nudibranchia" pageId="39" pageNumber="574" phylum="Mollusca" rank="species" species="maculigera">
<emphasis id="B94F120AFFC8606EFB7DF8D36CF1D46C" italics="true" pageId="39" pageNumber="574">Thordisa maculigera</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFAAB3E9FFC8606EFC29F8F26B21D46F" author="Bergh" box="[942,1106,1852,1874]" firstAuthor="Bergh" pageId="39" pageNumber="574" pagination="58 - 61" refId="ref53810" refString="Bergh R. 1877 a. Malacologische Untersuchungen. In: Semper C, ed. Reisen im Archipel der Philippinen, theil 2, heft 10. Wiesbaden: Kreidel, 495 - 546, plates 58 - 61." type="book chapter" year="1877" yearSuffix="a">Bergh (1877a)</bibRefCitation>
and there is no doubt these two names are synonyms.
</paragraph>
</subSubSection>
</treatment>
</document>