189 lines
21 KiB
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189 lines
21 KiB
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<document id="32B3CA214AC970A9A49908EF438CED60" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude & Tabuce, Rodolphe" docDate="2021" docId="03923C45FF89FF8030F5FD8AFD6A13A3" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Tethylamna Cappetta & Case 2016" docType="treatment" docVersion="6" lastPageNumber="127" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="127" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="9064FDA2AC3F5840EC8B8B1339275C6E">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="4A359D39F92B9FC34C8931B9ABB143C7">Zouhri, Samir</mods:namePart>
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<mods:namePart id="77FA2BF9CD8F477CEADA51A476FAC3F4">Gingerich, Philip D.</mods:namePart>
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<mods:namePart id="02D1A11DCDAEF3DA44766BC86DE96F72">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="4A6DDAE082CE8C5F5E2FAD3CFD7C4D5E">Bourdon, Estelle</mods:namePart>
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<mods:namePart id="57DABA7F1413CC701A15895C59446563">Adnet, Sylvain</mods:namePart>
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<mods:namePart id="3D4F693C7456A52589E50CEA6CDD2CAD">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="22F446608501F01FBE6F6A30126B53DE">Elboudali, Najia</mods:namePart>
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<mods:namePart id="80ABFE3BADE9F35994023E250E0E75F8">Amane, Ayoub</mods:namePart>
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<mods:namePart id="04DC4A276A2735F5FBD60E894E63285A">Rage, Jean-Claude</mods:namePart>
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<mods:namePart id="A841D98D49F4261742778110F1F9EBA9">Tabuce, Rodolphe</mods:namePart>
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<mods:typeOfResource id="D806552EF55F9AA9A80B9A205BFD4483">text</mods:typeOfResource>
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<mods:title id="6EA8E3E3E97E0FADD32ACB7991CF7DC5">Geodiversitas</mods:title>
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<mods:part id="84B283E8294381DCC57537044A86234E">
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<mods:date id="91803C8E7F95EBFA31D3B9975510C710">2021</mods:date>
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<mods:number id="54C8E3AD5CCBE44BF85FDCB850C90CBC">2021-03-11</mods:number>
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<mods:number id="F280B9141B5D3566172EFD6449E14EF7">43</mods:number>
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<mods:start id="6059731139F106B2451E6D7583EBF8CA">121</mods:start>
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<mods:identifier id="9C9D0EEE098D104AB9117CC2565CF97C" type="DOI">10.5252/geodiversitas2021v43a5</mods:identifier>
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<mods:identifier id="0D6FEE460C33C5005004251578F69A9C" type="GBIF-Dataset">e42658a2-0e15-4e90-9bc5-0d00cbcbd781</mods:identifier>
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<mods:identifier id="A7702E291F8F48A4534A84CB8B21CFD8" type="ISSN">1638-9395</mods:identifier>
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<treatment id="03923C45FF89FF8030F5FD8AFD6A13A3" ID-DOI="http://doi.org/10.5281/zenodo.4606615" ID-GBIF-Taxon="180569482" ID-Zenodo-Dep="4606615" LSID="urn:lsid:plazi:treatment:03923C45FF89FF8030F5FD8AFD6A13A3" httpUri="http://treatment.plazi.org/id/03923C45FF89FF8030F5FD8AFD6A13A3" lastPageNumber="127" pageId="8" pageNumber="127">
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<subSubSection id="C321DED8FF89FF8030F5FD8AFD0B141E" box="[237,670,597,624]" pageId="8" pageNumber="127" type="nomenclature">
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<paragraph id="8B848D53FF89FF8030F5FD8AFD0B141E" blockId="8.[237,670,597,624]" box="[237,670,597,624]" pageId="8" pageNumber="127">
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<heading id="D0CC3A3FFF89FF8030F5FD8AFD0B141E" box="[237,670,597,624]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="127" reason="2">
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<taxonomicName id="4C3BF6D0FF89FF8030F5FD8AFEE5141E" authorityName="Cappetta & Case" authorityYear="2016" box="[237,368,597,624]" class="Elasmobranchii" family="Odontaspididae" genus="Tethylamna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="8" pageNumber="127" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF89FF8030F5FD8AFEE5141E" bold="true" box="[237,368,597,624]" italics="true" pageId="8" pageNumber="127">Tethylamna</emphasis>
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</taxonomicName>
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cf.
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<emphasis id="B94F5141FF89FF803180FD89FD99141E" bold="true" box="[408,524,598,624]" italics="true" pageId="8" pageNumber="127">twiggsensis</emphasis>
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(
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<bibRefCitation id="EFAAF0A2FF89FF803203FD89FD03141E" author="CASE G. R." box="[539,662,598,624]" pageId="8" pageNumber="127" pagination="52 - 79" refId="ref20388" refString="CASE G. R. 1981. - Late Eocene selachians from South-central Georgia. Palaeontographica Abteilung A 176 (1 - 3): 52 - 79." type="journal article" year="1981">Case, 1981</bibRefCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF89FF80309CFD50FDDF14AF" pageId="8" pageNumber="127" type="materials_examined">
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<paragraph id="8B848D53FF89FF80309CFD50FDDF14AF" blockId="8.[131,775,655,705]" pageId="8" pageNumber="127">EXAMINED MATERIAL. — 50 isolated teeth, figured material includes FSAC Bouj-331, 332, 333, 334, 335 and 336.</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF89FF80309CFD29FE9D15E1" pageId="8" pageNumber="127" type="description">
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<paragraph id="8B848D53FF89FF80309CFD29FE801560" blockId="8.[132,776,758,911]" box="[132,277,758,784]" pageId="8" pageNumber="127">DESCRIPTION</paragraph>
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<paragraph id="8B848D53FF89FF80309CFCC9FE9D15E1" blockId="8.[132,776,758,911]" pageId="8" pageNumber="127">
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Numerous teeth are attributed to odontaspidid
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<taxonomicName id="4C3BF6D0FF89FF803297FCCAFC9D1541" authorityName="Cappetta & Case" authorityYear="2016" box="[655,776,789,815]" class="Elasmobranchii" family="Odontaspididae" genus="Tethylamna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="8" pageNumber="127" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF89FF803297FCCAFC9D1541" box="[655,776,789,815]" italics="true" pageId="8" pageNumber="127">Tethylamna</emphasis>
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</taxonomicName>
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cf.
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<emphasis id="B94F5141FF89FF8030BDFCE9FE871521" box="[165,274,822,847]" italics="true" pageId="8" pageNumber="127">twiggsensis</emphasis>
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. This species is easily recognizable by a pair of double flat cusplets on anterior (
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<figureCitation id="130091D6FF89FF8031F8FC8AFDD9151E" box="[480,588,853,880]" captionStart="FIG" captionStartId="9.[132,143,1751,1768]" captionTargetBox="[132,1455,215,1708]" captionTargetId="figure-89@9.[698,1267,774,1335]" captionTargetPageId="9" captionText="FIG. 3. — Elasmobranchii from Sabkha of Gueran collected around the Archaeocete whale carcasses.(A-F) Tethylamna cf.twiggsensis: A, FSAC Bouj-331 anterior lower tooth in labial view; B, FSAC Bouj-332 anterolateral lower tooth in labial view; C, FSAC Bouj-333 lateral lower tooth in (C1) labial and (C2) lingual views; D, FSAC Bouj-334 posterior lower tooth in labial view; E, FSAC Bouj-335 lateral lower tooth in (E1) labial and (E2) lingual views; F, FSAC Bouj-336 anterior upper tooth in (F1) labial and (F2) lingual views. G-I, Galeocerdo cf. eaglesomei; G, FSAC Bouj-337 anterior tooth in (G1) labial and (G2) lingual views; H, FSAC Bouj- 338 anterolateral tooth in lingual view; I, FSAC Bouj-339 lateral tooth in (I1) labial and (I2) lingual views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605969" httpUri="https://zenodo.org/record/4605969/files/figure.png" pageId="8" pageNumber="127">Fig. 3A, F</figureCitation>
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) and lateral teeth (
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<figureCitation id="130091D6FF89FF803095FCAAFF5E15E1" box="[141,203,885,911]" captionStart="FIG" captionStartId="9.[132,143,1751,1768]" captionTargetBox="[132,1455,215,1708]" captionTargetId="figure-89@9.[698,1267,774,1335]" captionTargetPageId="9" captionText="FIG. 3. — Elasmobranchii from Sabkha of Gueran collected around the Archaeocete whale carcasses.(A-F) Tethylamna cf.twiggsensis: A, FSAC Bouj-331 anterior lower tooth in labial view; B, FSAC Bouj-332 anterolateral lower tooth in labial view; C, FSAC Bouj-333 lateral lower tooth in (C1) labial and (C2) lingual views; D, FSAC Bouj-334 posterior lower tooth in labial view; E, FSAC Bouj-335 lateral lower tooth in (E1) labial and (E2) lingual views; F, FSAC Bouj-336 anterior upper tooth in (F1) labial and (F2) lingual views. G-I, Galeocerdo cf. eaglesomei; G, FSAC Bouj-337 anterior tooth in (G1) labial and (G2) lingual views; H, FSAC Bouj- 338 anterolateral tooth in lingual view; I, FSAC Bouj-339 lateral tooth in (I1) labial and (I2) lingual views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605969" httpUri="https://zenodo.org/record/4605969/files/figure.png" pageId="8" pageNumber="127">Fig. 3</figureCitation>
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C-E).
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF89FF80309CFC6AFD6A13A3" pageId="8" pageNumber="127" type="discussion">
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<paragraph id="8B848D53FF89FF80309CFC6AFF7D15A3" blockId="8.[130,782,949,1485]" box="[132,232,949,975]" pageId="8" pageNumber="127">REMARKS</paragraph>
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<paragraph id="8B848D53FF89FF80309CFC0BFD6A13A3" blockId="8.[130,782,949,1485]" pageId="8" pageNumber="127">
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The range of this species is currently restricted to the latest Lutetian-late Priabonian and its geographic distribution extends to paleotropical seas between tropical eastern Pacific, Caribbean and oriental Neotethys (
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<bibRefCitation id="EFAAF0A2FF89FF8031AEFBEBFDA11220" author="CASIER E." box="[438,564,1076,1102]" pageId="8" pageNumber="127" pagination="1 - 9" refId="ref20498" refString="CASIER E. 1971. - Sur un materiel ichthyologique des ' Midra (and Saila) shales' du Qatar (Golfe Persique). Bulletin de l'Institut royal des Sciences naturelles de Belgique 47 (2): 1 - 9" type="journal article" year="1971">Casier 1971</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF803258FBEBFD3B1220" author="CASE G. R." box="[576,686,1076,1102]" pageId="8" pageNumber="127" pagination="52 - 79" refId="ref20388" refString="CASE G. R. 1981. - Late Eocene selachians from South-central Georgia. Palaeontographica Abteilung A 176 (1 - 3): 52 - 79." type="journal article" year="1981">Case 1981</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF8032A1FBEBFE8E1200" author="CASE G. R. & BORODIN P. D." pageId="8" pageNumber="127" pagination="5 - 16" refId="ref20417" refString="CASE G. R. & BORODIN P. D. 2000. - Late Eocene selachian from the Irwinton Sand Member of the Barnwell Formation (Jacksonian), WKA mines, Gordon, Wilkinson County, Georgy. Munchner Geowissenschaftliche Abhandlungen 39: 5 - 16." type="journal article" year="2000">Case & Borodin 2000</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF80313FFB8BFD891200" author="CASE G. R. & CAPPETTA H." box="[295,540,1108,1134]" pageId="8" pageNumber="127" pagination="1 - 30" refId="ref20466" refString="CASE G. R. & CAPPETTA H. 1990. - The Eocene Selachians Fauna from the Fayum Depression in Egypt. Palaeontographica Abteilung A 212: 1 - 30." type="journal article" year="1990">Case & Cappetta 1990</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF803230FB8BFC951200" author="WARD J. W. & WIEST R. L." box="[552,768,1108,1134]" pageId="8" pageNumber="127" pagination="81 - 88" refId="ref26141" refString="WARD J. W. & WIEST R. L. 1990. - A checklist of Palaeocene and Eocene sharks and rays (Chondrichthyes) from the Pamunkey Group, Maryland and Virginia, USA. Tertiary Research 12: 81 - 88." type="journal article" year="1990">Ward & Wiest 1990</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF80309CFBABFEA912E0" author="ADNET S. & ANTOINE P. O. & HASSAN BAQRIS & CROCHET J. Y. & MARIVAUX L. & WELCOMME J. L. & METAIS G." box="[132,316,1140,1166]" pageId="8" pageNumber="127" pagination="303 - 323" refId="ref18851" refString="ADNET S., ANTOINE P. O., HASSAN BAQRIS. R., CROCHET J. Y., MARIVAUX L., WELCOMME J. L. & METAIS G. 2007. - On the first modern and tropical Selachian association (Chondrichthyes) from the late Eocene-early Oligocene of Baluchistan, Pakistan. Dating interests, paleoenvironment and paleobiogeography. Journal of Asian Earth Sciences 30: 303 - 323. https: // doi. org / 10.1017 / S 0016756810000348" type="journal article" year="2007">
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Adnet
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<emphasis id="B94F5141FF89FF8030D4FBAAFF6A12E0" box="[204,255,1140,1166]" italics="true" pageId="8" pageNumber="127">et al.</emphasis>
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2007
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</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF803151FBABFDD412E0" author="UNDERWOOD C. J. & WARD D. J. & KING C. & ANTAR S. M. & MOUT I. S. & GINGERICH P. D." box="[329,577,1140,1166]" pageId="8" pageNumber="127" pagination="47 - 66" refId="ref26062" refString="UNDERWOOD C. J., WARD D. J., KING C., ANTAR S. M., ZAL- MOUT I. S. & GINGERICH P. D. 2011. - Shark and ray faunas in the middle and late Eocene of the Fayum area, Egypt. Proceedings of the Geologists' Association 122: 47 - 66. https: // doi. org / 10.1016 / j. pgeola. 20" type="journal article" year="2011">
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Underwood
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<emphasis id="B94F5141FF89FF8031C9FBAAFD9612E0" box="[465,515,1140,1166]" italics="true" pageId="8" pageNumber="127">et al.</emphasis>
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2011
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</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF89FF803256FBABFF2E12C0" author="CAPPETTA H. & CASE G. R." pageId="8" pageNumber="127" pagination="43 - 103" refId="ref20339" refString="CAPPETTA H. & CASE G. R. 2016. - A Selachian Fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA. Palaeontographica Abteilung A 307 (1 - 6): 43 - 103." type="journal article" year="2016">Cappetta & Case 2016</bibRefCitation>
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). Originally described from the late Eocene of
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<collectingCountry id="F32CCDC3FF89FF8032B2FB4BFC9612C0" box="[682,771,1172,1198]" name="Georgia" pageId="8" pageNumber="127">Georgia</collectingCountry>
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,
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<collectingCountry id="F32CCDC3FF89FF80309BFB6BFE8012A0" box="[131,277,1204,1230]" name="United States of America" pageId="8" pageNumber="127">United States</collectingCountry>
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(
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<bibRefCitation id="EFAAF0A2FF89FF80313FFB6BFE0E12A0" author="CASE G. R." box="[295,411,1204,1230]" pageId="8" pageNumber="127" pagination="52 - 79" refId="ref20388" refString="CASE G. R. 1981. - Late Eocene selachians from South-central Georgia. Palaeontographica Abteilung A 176 (1 - 3): 52 - 79." type="journal article" year="1981">Case 1981</bibRefCitation>
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), distinction between the middle and the late Eocene representatives are sometimes controversial. The Bartonian teeth have lateral cusplets less pronounced compared to those from Priabonian (
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<bibRefCitation id="EFAAF0A2FF89FF803212FACCFD6B1340" author="UNDERWOOD C. J. & WARD D. J. & KING C. & ANTAR S. M. & MOUT I. S. & GINGERICH P. D." box="[522,766,1299,1326]" pageId="8" pageNumber="127" pagination="47 - 66" refId="ref26062" refString="UNDERWOOD C. J., WARD D. J., KING C., ANTAR S. M., ZAL- MOUT I. S. & GINGERICH P. D. 2011. - Shark and ray faunas in the middle and late Eocene of the Fayum area, Egypt. Proceedings of the Geologists' Association 122: 47 - 66. https: // doi. org / 10.1016 / j. pgeola. 20" type="journal article" year="2011">
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Underwood
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<emphasis id="B94F5141FF89FF803288FACBFD571343" box="[656,706,1299,1325]" italics="true" pageId="8" pageNumber="127">et al.</emphasis>
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2011
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</bibRefCitation>
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) explaining why they are often referred to confer
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<emphasis id="B94F5141FF89FF80328EFAEBFC951323" box="[662,768,1332,1357]" italics="true" pageId="8" pageNumber="127">twigssensis</emphasis>
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. These Bartonian samples could corresponds to intergradual change from the possible ancestor
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<taxonomicName id="4C3BF6D0FF89FF8031F8FAABFDA213E3" authorityName="Cappetta & Case" authorityYear="2016" box="[480,567,1395,1421]" class="Elasmobranchii" family="Odontaspididae" genus="Tethylamna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="8" pageNumber="127" phylum="Chordata" rank="species" species="dunni">
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<emphasis id="B94F5141FF89FF8031F8FAABFDA213E3" box="[480,567,1395,1421]" italics="true" pageId="8" pageNumber="127">T. dunni</emphasis>
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</taxonomicName>
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of
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<bibRefCitation id="EFAAF0A2FF89FF80324DFAACFF5B13C0" author="CAPPETTA H. & CASE G. R." pageId="8" pageNumber="127" pagination="43 - 103" refId="ref20339" refString="CAPPETTA H. & CASE G. R. 2016. - A Selachian Fauna from the middle Eocene (Lutetian, Lisbon Formation) of Andalusia, Covington County, Alabama, USA. Palaeontographica Abteilung A 307 (1 - 6): 43 - 103." type="journal article" year="2016">Cappetta & Case (2016)</bibRefCitation>
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recovered from the Lutetian of
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<collectingRegion id="49FF43B1FF89FF803201FA4CFDEF13C3" box="[537,634,1427,1453]" country="United States of America" name="Alabama" pageId="8" pageNumber="127">Alabama</collectingRegion>
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,
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<collectingCountry id="F32CCDC3FF89FF80329BFA4CFD2213C3" box="[643,695,1427,1453]" name="United States of America" pageId="8" pageNumber="127">USA</collectingCountry>
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toward those of Priabonian, including
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<typeStatus id="548033F1FF89FF8031C9FA6CFE6A13A3" box="[465,511,1459,1485]" pageId="8" pageNumber="127">type</typeStatus>
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of species (
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<bibRefCitation id="EFAAF0A2FF89FF803265FA6CFD7A13A3" author="CASE G. R." box="[637,751,1459,1485]" pageId="8" pageNumber="127" pagination="52 - 79" refId="ref20388" refString="CASE G. R. 1981. - Late Eocene selachians from South-central Georgia. Palaeontographica Abteilung A 176 (1 - 3): 52 - 79." type="journal article" year="1981">Case 1981</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |