259 lines
30 KiB
XML
259 lines
30 KiB
XML
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<mods:title id="2001D66AF263E9068644BA552B0BFA92">Description of a new species of the genus Nikkoaspis Kuwana, 1928 (Hemiptera Coccomorpha: Diaspididae) from China</mods:title>
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<mods:namePart id="A96C7F30B4D5D7629E28B8E12DF97D46">Tian, Feng</mods:namePart>
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<mods:affiliation id="C76E3ECD2D1152B7A7B40B288EAD2E11">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China & tianfeng 550369926 @ 126. com; https: // orcid. org / 0000 - 0002 - 5385 - 323 X</mods:affiliation>
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<mods:namePart id="6DE3831CE2E6C464346A2EF8F5C4D65C">Zheng, Xinyi</mods:namePart>
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<mods:affiliation id="8C920A79AE8504B62F9E106584AB51B4">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China & zxy 780346931 @ 126. com; https: // orcid. org / 0000 - 0002 - 7396 - 7488</mods:affiliation>
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<mods:namePart id="183A493CA385C0753F1AFD37520E4FCF">Xing, Jichun</mods:namePart>
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<mods:affiliation id="69521DEF3CB528DCA95BFD2994253565">Institute of Entomology, Guizhou University; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources, Guizhou University; Guiyang, 550025 P. R. China</mods:affiliation>
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<mods:date id="08C323002CB68D2FD42D4F83A116101B">2021</mods:date>
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<treatment id="0390E60BFFCDFFE6FF09E595F43FFE39" ID-DOI="http://doi.org/10.5281/zenodo.4666519" ID-GBIF-Taxon="180877680" ID-Zenodo-Dep="4666519" LSID="urn:lsid:plazi:treatment:0390E60BFFCDFFE6FF09E595F43FFE39" httpUri="http://treatment.plazi.org/id/0390E60BFFCDFFE6FF09E595F43FFE39" lastPageId="2" lastPageNumber="364" pageId="1" pageNumber="364">
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Genus
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<taxonomicName id="4C392C9EFFCDFFE5FF70E595F44CFB09" ID-CoL="645J" ID-ENA="665292" authority="Kuwana, 1928" authorityName="Kuwana" authorityYear="1928" box="[238,559,1195,1222]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="B94D8B0FFFCDFFE5FF70E595F717FB0A" bold="true" box="[238,372,1195,1222]" italics="true" pageId="1" pageNumber="364">Nikkoaspis</emphasis>
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<bibRefCitation id="EFA82AECFFCDFFE5FEE5E595F44CFB09" author="Kuwana, S. I." box="[379,559,1195,1221]" pageId="1" pageNumber="364" pagination="1 - 39" refId="ref4357" refString="Kuwana, S. I. (1928) The diaspine Coccidae of Japan. V. Genera Chionaspis, Tsukushiaspis [n. gen.], Leucaspis, Nikkoaspis [n. gen.]. Scientific Bulletin. Japan, 1, 1 - 39." type="journal article" year="1928">Kuwana, 1928</bibRefCitation>
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</emphasis>
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</paragraph>
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<paragraph id="8B86571DFFCDFFE5FF09E5CEF339FACB" blockId="1.[151,1370,1263,1288]" box="[151,1370,1263,1288]" pageId="1" pageNumber="364">
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<treatmentCitation id="0A98710CFFCDFFE5FF09E5CEF7A9FAC4" author="Kuwana, S. I." box="[151,458,1264,1288]" page="37" pageId="1" pageNumber="364" year="1928">
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<taxonomicName id="4C392C9EFFCDFFE5FF09E5CEF7A9FAC4" ID-CoL="645J" authority="Kuwana, 1928: 37" authorityName="Kuwana" authorityPageNumber="37" authorityYear="1928" box="[151,458,1264,1288]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
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<emphasis id="B94D8B0FFFCDFFE5FF09E5CEF765FAC4" box="[151,262,1264,1288]" italics="true" pageId="1" pageNumber="364">Nikkoaspis</emphasis>
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<bibRefCitation id="EFA82AECFFCDFFE5FE93E5CEF7A9FAC4" author="Kuwana, S. I." box="[269,458,1264,1288]" pageId="1" pageNumber="364" pagination="1 - 39" refId="ref4357" refString="Kuwana, S. I. (1928) The diaspine Coccidae of Japan. V. Genera Chionaspis, Tsukushiaspis [n. gen.], Leucaspis, Nikkoaspis [n. gen.]. Scientific Bulletin. Japan, 1, 1 - 39." type="journal article" year="1928">Kuwana, 1928: 37</bibRefCitation>
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;
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<treatmentCitation id="0A98710CFFCDFFE5FE4BE5CEF4F8FACB" author="Chou, I." box="[469,667,1263,1288]" page="57" pageId="1" pageNumber="364" year="1982">
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<bibRefCitation id="EFA82AECFFCDFFE5FE4BE5CEF4F8FACB" author="Chou, I." box="[469,667,1263,1288]" pageId="1" pageNumber="364" refId="ref4108" refString="Chou, I. (1982) Monograph of the Diaspididae of China. Vol. 1. Shaanxi Publishing House of Science & Technology, Shaanxi, 58 pp." type="book" year="1982">Chou, 1982: 57–58</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A98710CFFCDFFE5FD3BE5CEF50FFAC4" author="Hu, J. L." box="[677,876,1264,1288]" page="138" pageId="1" pageNumber="364" year="1987">
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<bibRefCitation id="EFA82AECFFCDFFE5FD3BE5CEF50FFAC4" author="Hu, J. L." box="[677,876,1264,1288]" pageId="1" pageNumber="364" pagination="138 - 140" refId="ref4293" refString="Hu, J. L. (1987) A new species of Nikkoaspis (Coccoidea: Diaspididae). Contributions from Shanghai Institute of Entomology, 7, 138 - 140." type="journal article" year="1987">Hu, 1987: 138–140</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0A98710CFFCDFFE5FCE8E5CEF202FACB" author="Takagi, S." box="[886,1121,1263,1288]" page="114" pageId="1" pageNumber="364" year="1999">
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<bibRefCitation id="EFA82AECFFCDFFE5FCE8E5CEF202FACB" author="Takagi, S." box="[886,1121,1263,1288]" pageId="1" pageNumber="364" pagination="95 - 150" refId="ref4646" refString="Takagi, S. (1999) Notes on the scale insect subtribe Kuwanaspidina (Homoptera: Coccoidea: Diaspididae). Insecta Matsumurana New Series, 56, 95 - 150." type="journal article" year="1999">Takagi, 1999: 114–115</bibRefCitation>
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;
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<treatmentCitation id="0A98710CFFCDFFE5FBF5E5CEF353FACB" author="Normark, B. B. & Okusu, A. & Morse, G. E. & Peterson, D. A. & Itioka, T. & Schneider, S. A." box="[1131,1328,1263,1288]" page="52" pageId="1" pageNumber="364" year="2019">
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<bibRefCitation id="EFA82AECFFCDFFE5FBF5E5CEF353FACB" author="Normark, B. B. & Okusu, A. & Morse, G. E. & Peterson, D. A. & Itioka, T. & Schneider, S. A." box="[1131,1328,1263,1288]" pageId="1" pageNumber="364" pagination="1 - 98" refId="ref4519" refString="Normark, B. B., Okusu, A., Morse, G. E., Peterson, D. A., Itioka, T. & Schneider, S. A. (2019) Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae). Zootaxa, 4616 (1), 1 - 98. https: // doi. org / 10.11646 / zootaxa. 4616.1.1" type="journal article" year="2019">Normark, 2019: 52</bibRefCitation>
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, 67.
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</paragraph>
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<subSubSection id="C3230496FFCDFFE5FF09E413F33AF8D5" pageId="1" pageNumber="364" type="type_taxon">
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<paragraph id="8B86571DFFCDFFE5FF09E413F4D7FA89" blockId="1.[151,692,1325,1349]" box="[151,692,1325,1349]" pageId="1" pageNumber="364">
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<emphasis id="B94D8B0FFFCDFFE5FF09E413F740FA89" bold="true" box="[151,291,1325,1349]" pageId="1" pageNumber="364">
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<typeStatus id="5482E9BFFFCDFFE5FF09E413F6AFFA89" box="[151,204,1325,1349]" pageId="1" pageNumber="364">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C392C9EFFCDFFE5FEB4E413F4D3FA89" ID-CoL="6RZN4" authority="Kuwana, 1928" authorityName="Kuwana" authorityYear="1928" box="[298,688,1325,1349]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="shiranensis">
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<emphasis id="B94D8B0FFFCDFFE5FEB4E413F471FA89" box="[298,530,1325,1349]" italics="true" pageId="1" pageNumber="364">Nikkoaspis shiranensis</emphasis>
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<bibRefCitation id="EFA82AECFFCDFFE5FD87E413F4D3FA89" author="Kuwana, S. I." box="[537,688,1325,1349]" pageId="1" pageNumber="364" pagination="1 - 39" refId="ref4357" refString="Kuwana, S. I. (1928) The diaspine Coccidae of Japan. V. Genera Chionaspis, Tsukushiaspis [n. gen.], Leucaspis, Nikkoaspis [n. gen.]. Scientific Bulletin. Japan, 1, 1 - 39." type="journal article" year="1928">Kuwana, 1928</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8B86571DFFCDFFE5FF09E44DF75DFA7D" blockId="1.[151,1436,1395,2034]" pageId="1" pageNumber="364">
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<emphasis id="B94D8B0FFFCDFFE5FF09E44DF47DFA41" bold="true" box="[151,542,1395,1421]" pageId="1" pageNumber="364">Generic diagnosis (adult female).</emphasis>
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Scale cover of adult female white, elongate, convex; exuviae situated at anterior end, brownish.
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</paragraph>
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<paragraph id="8B86571DFFCDFFE5FF59E485F33AF8D5" blockId="1.[151,1436,1395,2034]" pageId="1" pageNumber="364">
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Slide-mounted adult female body elongate pyriform; cephalothorax tapering anteriorly; abdomen broad and expanded. Lateral margin of each abdominal segment with numerous plates and simple gland spines. Pygidium broad with posterior margin rounded. Pygidial lobes numbering 2 pairs; median lobes (L
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<subScript id="17BD5558FFCDFFE5FBB8E728F24DF9E8" attach="left" box="[1062,1070,1558,1572]" fontSize="6" pageId="1" pageNumber="364">1</subScript>
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) separated from each other, each L
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<subScript id="17BD5558FFCDFFE5FF36E704F6D3F984" attach="left" box="[168,176,1594,1608]" fontSize="6" pageId="1" pageNumber="364">1</subScript>
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split into 1 or 2 lobules; second lobes (L
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<subScript id="17BD5558FFCDFFE5FDE1E704F4E4F984" attach="left" box="[639,647,1594,1608]" fontSize="6" pageId="1" pageNumber="364">2</subScript>
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) each split into 2–4 lobules of equal size; each lobule slender, with or without apical incision. There are 2 plates between L
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<subScript id="17BD5558FFCDFFE5FD7BE760F48EF9A0" attach="left" box="[741,749,1630,1644]" fontSize="6" pageId="1" pageNumber="364">1</subScript>
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; space between L
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<subScript id="17BD5558FFCDFFE5FC2BE760F5DEF9A0" attach="left" box="[949,957,1630,1644]" fontSize="6" pageId="1" pageNumber="364">1</subScript>
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and L
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<subScript id="17BD5558FFCDFFE5FB9AE760F26FF9A0" attach="left" box="[1028,1036,1630,1644]" fontSize="6" pageId="1" pageNumber="364">2</subScript>
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with 1 or 2 plates. Margin lateral to L
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||
<subScript id="17BD5558FFCDFFE5FF36E7BCF6D3F95C" attach="left" box="[168,176,1666,1680]" fontSize="6" pageId="1" pageNumber="364">2</subScript>
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with some plates that form a continuous row with plates on margins of prepygidial abdominal segments. Gland spines absent from between L
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<subScript id="17BD5558FFCDFFE5FE7CE798F789F978" attach="left" box="[482,490,1702,1716]" fontSize="6" pageId="1" pageNumber="364">1</subScript>
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||
; 1 gland spine present lateral to L
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||
<subScript id="17BD5558FFCDFFE5FCFCE798F509F978" attach="left" box="[866,874,1702,1716]" fontSize="6" pageId="1" pageNumber="364">1</subScript>
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||
, and 1 or 2 gland spines situated lateral to L
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<subScript id="17BD5558FFCDFFE5FACCE798F339F978" attach="left" box="[1362,1370,1702,1716]" fontSize="6" pageId="1" pageNumber="364">2</subScript>
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. Anal opening situated slightly anterior to centre of pygidium. Perivulvar pores present in 5 groups. Dorsal macroducts small, densely covering the lateral margins of prepygidial abdominal segments, and extending along posterior margin of each pygidial segment to midline. Dorsal macroducts scattered and irregularly arranged (
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<bibRefCitation id="EFA82AECFFCDFFE5FB5AE7C1F32FF8D5" author="Chou, I." box="[1220,1356,1791,1817]" pageId="1" pageNumber="364" refId="ref4108" refString="Chou, I. (1982) Monograph of the Diaspididae of China. Vol. 1. Shaanxi Publishing House of Science & Technology, Shaanxi, 58 pp." type="book" year="1982">Chou, 1982</bibRefCitation>
|
||
).
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3230496FFCDFFE5FF59E61DF4AEF8F1" box="[199,717,1827,1854]" pageId="1" pageNumber="364" type="distribution">
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<paragraph id="8B86571DFFCDFFE5FF59E61DF4AEF8F1" blockId="1.[151,1436,1395,2034]" box="[199,717,1827,1854]" pageId="1" pageNumber="364">
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<emphasis id="B94D8B0FFFCDFFE5FF59E61DF702F8F1" bold="true" box="[199,353,1827,1853]" pageId="1" pageNumber="364">Distribution.</emphasis>
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<collectingCountry id="F32E178DFFCDFFE5FEF9E61DF7B3F8F1" box="[359,464,1827,1853]" name="Malaysia" pageId="1" pageNumber="364">Malaysia</collectingCountry>
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,
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||
<collectingCountry id="F32E178DFFCDFFE5FE42E61DF441F8F1" box="[476,546,1827,1853]" name="China" pageId="1" pageNumber="364">China</collectingCountry>
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,
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<collectingCountry id="F32E178DFFCDFFE5FDB3E61AF413F8F2" box="[557,624,1828,1854]" name="Japan" pageId="1" pageNumber="364">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32E178DFFCDFFE5FDE5E61DF4AAF8F1" box="[635,713,1827,1853]" name="Russia" pageId="1" pageNumber="364">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3230496FFCDFFE6FF59E679F43FFE39" lastPageId="2" lastPageNumber="365" pageId="1" pageNumber="364" type="discussion">
|
||
<paragraph id="8B86571DFFCDFFE6FF59E679F43FFE39" blockId="1.[151,1436,1395,2034]" lastBlockId="2.[151,1437,151,501]" lastPageId="2" lastPageNumber="365" pageId="1" pageNumber="364">
|
||
<emphasis id="B94D8B0FFFCDFFE5FF59E679F758F8AD" bold="true" box="[199,315,1863,1889]" pageId="1" pageNumber="364">Remarks.</emphasis>
|
||
<taxonomicName id="4C392C9EFFCDFFE5FED8E679F7A2F8AD" authorityName="Kuwana" authorityYear="1928" box="[326,449,1863,1889]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCDFFE5FED8E679F7A2F8AD" box="[326,449,1863,1889]" italics="true" pageId="1" pageNumber="364">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C392C9EFFCDFFE5FE60E676F4E4F8AD" authorityName="MacGillivray" authorityYear="1921" box="[510,647,1864,1889]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCDFFE5FE60E676F4E4F8AD" box="[510,647,1864,1889]" italics="true" pageId="1" pageNumber="364">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
are so similar that
|
||
<bibRefCitation id="EFA82AECFFCDFFE5FCF3E679F250F8AD" author="Takahashi, R." box="[877,1075,1863,1889]" pageId="1" pageNumber="364" pagination="1 - 45" refId="ref4714" refString="Takahashi, R. (1930) Observations on the Coccidae of Formosa, Part II. Report. Department of Agriculture Government Research Institute, Formosa, 43, 1 - 45." type="journal article" year="1930">Takahashi (1930)</bibRefCitation>
|
||
merged them into a single genus. However, according to
|
||
<bibRefCitation id="EFA82AECFFCDFFE5FE45E655F4C5F849" author="Takagi, S." box="[475,678,1899,1925]" pageId="1" pageNumber="364" pagination="1 - 146" refId="ref4597" refString="Takagi, S. (1970) Diaspididae of Taiwan based on material collected in connection with the Japan-U. S. Cooperative Science Programme, 1965 (Homoptera: Coccoidea). Pt. II. Insecta Matsumurana, New Series, 33, 1 - 146." type="journal article" year="1970">Takagi (1970:122</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA82AECFFCDFFE5FD2AE655F57DF84A" author="Takagi, S." box="[692,798,1899,1926]" pageId="1" pageNumber="364" pagination="95 - 150" refId="ref4646" refString="Takagi, S. (1999) Notes on the scale insect subtribe Kuwanaspidina (Homoptera: Coccoidea: Diaspididae). Insecta Matsumurana New Series, 56, 95 - 150." type="journal article" year="1999">1999:115</bibRefCitation>
|
||
)
|
||
<taxonomicName id="4C392C9EFFCDFFE5FCAEE655F5C8F849" authorityName="Kuwana" authorityYear="1928" box="[816,939,1899,1925]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCDFFE5FCAEE655F5C8F849" box="[816,939,1899,1925]" italics="true" pageId="1" pageNumber="364">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
species are found on high-altitude bamboo in boreo-montane forests, whereas
|
||
<taxonomicName id="4C392C9EFFCDFFE5FDBFE6AEF4C9F865" authorityName="MacGillivray" authorityYear="1921" box="[545,682,1936,1961]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCDFFE5FDBFE6AEF4C9F865" box="[545,682,1936,1961]" italics="true" pageId="1" pageNumber="364">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
species occur in a wide range of habitats from tropical lowlands to temperate regions. These differing distribution traits, together with the peculiar body shape and numerous dorsal macroducts, were used to justify acceptance of
|
||
<taxonomicName id="4C392C9EFFCDFFE5FD3AE6E9F57CF83D" authorityName="Kuwana" authorityYear="1928" box="[676,799,2007,2033]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCDFFE5FD3AE6E9F57CF83D" box="[676,799,2007,2033]" italics="true" pageId="1" pageNumber="364">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
as a valid genus by
|
||
<bibRefCitation id="EFA82AECFFCDFFE5FC63E6E9F2FEF83D" author="Takagi, S." box="[1021,1181,2007,2033]" pageId="1" pageNumber="364" pagination="1 - 146" refId="ref4597" refString="Takagi, S. (1970) Diaspididae of Taiwan based on material collected in connection with the Japan-U. S. Cooperative Science Programme, 1965 (Homoptera: Coccoidea). Pt. II. Insecta Matsumurana, New Series, 33, 1 - 146." type="journal article" year="1970">Takagi (1970)</bibRefCitation>
|
||
. However, in their molecular phylogenetic analysis,
|
||
<bibRefCitation id="EFA82AECFFCEFFE6FE72E1A9F480FF7D" author="Normark, B. B. & Okusu, A. & Morse, G. E. & Peterson, D. A. & Itioka, T. & Schneider, S. A." box="[492,739,151,177]" pageId="2" pageNumber="365" pagination="1 - 98" refId="ref4519" refString="Normark, B. B., Okusu, A., Morse, G. E., Peterson, D. A., Itioka, T. & Schneider, S. A. (2019) Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae). Zootaxa, 4616 (1), 1 - 98. https: // doi. org / 10.11646 / zootaxa. 4616.1.1" type="journal article" year="2019">
|
||
Normark
|
||
<emphasis id="B94D8B0FFFCEFFE6FDC6E1A9F4E9FF7D" box="[600,650,151,177]" italics="true" pageId="2" pageNumber="365">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
found that
|
||
<taxonomicName id="4C392C9EFFCEFFE6FCFBE1A9F583FF7D" authorityName="Kuwana" authorityYear="1928" box="[869,992,151,177]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FCFBE1A9F583FF7D" box="[869,992,151,177]" italics="true" pageId="2" pageNumber="365">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
is nested within
|
||
<taxonomicName id="4C392C9EFFCEFFE6FB3EE1A9F34AFF7C" authorityName="MacGillivray" authorityYear="1921" box="[1184,1321,151,176]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FB3EE1A9F34AFF7C" box="[1184,1321,151,176]" italics="true" pageId="2" pageNumber="365">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
, such that
|
||
<taxonomicName id="4C392C9EFFCEFFE6FF09E185F743FF18" authorityName="MacGillivray" authorityYear="1921" box="[151,288,187,212]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FF09E185F743FF18" box="[151,288,187,212]" italics="true" pageId="2" pageNumber="365">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
as presently constituted is paraphyletic, probably being comprised of multiple morphologically cryptic genera. Their analysis also showed
|
||
<taxonomicName id="4C392C9EFFCEFFE6FDB7E1E1F4C7FF35" authorityName="Kuwana" authorityYear="1928" box="[553,676,223,249]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FDB7E1E1F4C7FF35" box="[553,676,223,249]" italics="true" pageId="2" pageNumber="365">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
as being sister to a tropical clade of
|
||
<taxonomicName id="4C392C9EFFCEFFE6FBD8E1E1F2ACFF34" authorityName="MacGillivray" authorityYear="1921" box="[1094,1231,223,248]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FBD8E1E1F2ACFF34" box="[1094,1231,223,248]" italics="true" pageId="2" pageNumber="365">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
(all samples from lowland Borneo), whereas all the temperate samples of
|
||
<taxonomicName id="4C392C9EFFCEFFE6FC96E03DF5F2FED0" authorityName="MacGillivray" authorityYear="1921" box="[776,913,259,284]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FC96E03DF5F2FED0" box="[776,913,259,284]" italics="true" pageId="2" pageNumber="365">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
including the genotype,
|
||
<taxonomicName id="4C392C9EFFCEFFE6FB36E03DF6B8FE8D" authority="(Kuwana, 1902)" baseAuthorityName="Kuwana" baseAuthorityYear="1902" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="hikosani">
|
||
<emphasis id="B94D8B0FFFCEFFE6FB36E03DF344FED1" box="[1192,1319,259,285]" italics="true" pageId="2" pageNumber="365">K. hikosani</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA82AECFFCEFFE6FAA8E03DF6B0FE8D" author="Kuwana, S. I." pageId="2" pageNumber="365" pagination="43 - 98" refId="ref4326" refString="Kuwana, S. I. (1902) Coccidae (scale insects) of Japan. Proceedings of the California Academy of Sciences, 3, 43 - 98." type="journal article" year="1902">Kuwana, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, formed a separate clade. In principle, these clades might be accepted as separate genera, despite their close morphological similarity, but
|
||
<bibRefCitation id="EFA82AECFFCEFFE6FE7BE075F4BFFEA9" author="Normark, B. B. & Okusu, A. & Morse, G. E. & Peterson, D. A. & Itioka, T. & Schneider, S. A." box="[485,732,331,357]" pageId="2" pageNumber="365" pagination="1 - 98" refId="ref4519" refString="Normark, B. B., Okusu, A., Morse, G. E., Peterson, D. A., Itioka, T. & Schneider, S. A. (2019) Phylogeny and classification of armored scale insects (Hemiptera: Coccomorpha: Diaspididae). Zootaxa, 4616 (1), 1 - 98. https: // doi. org / 10.11646 / zootaxa. 4616.1.1" type="journal article" year="2019">
|
||
Normark
|
||
<emphasis id="B94D8B0FFFCEFFE6FDCFE075F4E0FEA9" box="[593,643,331,357]" italics="true" pageId="2" pageNumber="365">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
did not do so; nor did they synonymise
|
||
<taxonomicName id="4C392C9EFFCEFFE6FB37E075F347FEA9" authorityName="Kuwana" authorityYear="1928" box="[1193,1316,331,357]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FB37E075F347FEA9" box="[1193,1316,331,357]" italics="true" pageId="2" pageNumber="365">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
under
|
||
<taxonomicName id="4C392C9EFFCEFFE6FAECE075F69CFE44" authorityName="MacGillivray" authorityYear="1921" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FAECE075F69CFE44" italics="true" pageId="2" pageNumber="365">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
. Revision of the genus-level taxonomy is beyond the scope of the present article but future revision may result in splitting
|
||
<taxonomicName id="4C392C9EFFCEFFE6FEC3E0ADF785FE60" authorityName="MacGillivray" authorityYear="1921" box="[349,486,403,428]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FEC3E0ADF785FE60" box="[349,486,403,428]" italics="true" pageId="2" pageNumber="365">Kuwanaspis</emphasis>
|
||
</taxonomicName>
|
||
(as presently understood) into more than one genus. The temperate clade containing
|
||
<taxonomicName id="4C392C9EFFCEFFE6FF09E089F774FE1D" baseAuthorityName="Kuwana" baseAuthorityYear="1902" box="[151,279,439,465]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="hikosani">
|
||
<emphasis id="B94D8B0FFFCEFFE6FF09E089F774FE1D" box="[151,279,439,465]" italics="true" pageId="2" pageNumber="365">K. hikosani</emphasis>
|
||
</taxonomicName>
|
||
would retain the name
|
||
<emphasis id="B94D8B0FFFCEFFE6FDBFE089F4D3FE1C" box="[545,688,439,464]" italics="true" pageId="2" pageNumber="365">
|
||
<taxonomicName id="4C392C9EFFCEFFE6FDBFE089F4CEFE1C" authorityName="MacGillivray" authorityYear="1921" box="[545,685,439,464]" class="Insecta" family="Diaspididae" genus="Kuwanaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">Kuwanaspis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C392C9EFFCEFFE6FD76E089F500FE1D" authorityName="Kuwana" authorityYear="1928" box="[744,867,439,465]" class="Insecta" family="Diaspididae" genus="Nikkoaspis" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D8B0FFFCEFFE6FD76E089F500FE1D" box="[744,867,439,465]" italics="true" pageId="2" pageNumber="365">Nikkoaspis</emphasis>
|
||
</taxonomicName>
|
||
(as presently understood) might remain distinct or form part of the unnamed tropical clade.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |