treatments-xml/data/03/90/87/039087ED891588607AAAD421F6E9FC88.xml

<document id="110A78432CAC904EE4B149018C36A12E" ID-DOI="10.1590/S0031-10492010003700001" ID-ISSN="1807-0205" ID-Zenodo-Dep="13307773" IM.bibliography_requiresApprovalFor="GgImagineBatch" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1720049385726" checkinUser="felipe" docAuthor="De Armas, Luis F., Manzanilla, Osvaldo Villarreal &amp; Viquez, Carlos" docDate="2010" docId="039087ED891588607AAAD421F6E9FC88" docLanguage="es" docName="PapAvZool.50.37.579-586.pdf.imf" docOrigin="Papéis Avulsos de Zoologia 50 (37)" docStyle="DocumentStyle:342E0B2A517D47F211C16A9F07059896.3:PapAvZool.2002-2013.journal_article.type2" docStyleId="342E0B2A517D47F211C16A9F07059896" docStyleName="PapAvZool.2002-2013.journal_article.type2" docStyleVersion="3" docTitle="Surazomus brus Armas, Manzanilla &amp; Viquez, 2010, sp. nov." docType="treatment" docVersion="4" lastPageNumber="582" masterDocId="FFA9FF95891488637A47D759F407FF98" masterDocTitle="Nuevas Especies De Surazomus Reddell &amp; Cokendolpher, 1995 (Schizomida: Hubbardiidae) De Costa Rica" masterLastPageNumber="586" masterPageNumber="579" pageNumber="580" updateTime="1723490102617" updateUser="ExternalLinkService">
<mods:mods id="A5B2A606C471AC8F54394F61DC70CF2D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DF3DD87DF931874E2DCDB0898A5F063C">
<mods:title id="4F5E3D8A80D3269396716801A7859C22">Nuevas Especies De Surazomus Reddell &amp; Cokendolpher, 1995 (Schizomida: Hubbardiidae) De Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="E3F61069F350E90366E45AA9997B0D3F" type="personal">
<mods:role id="D39A35CB84028D2637077C404BBC7CD2">
<mods:roleTerm id="3B8EF31DEF8D8F15FE8408F3B3E569F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E05BFB7C9C894A23D12ACD8E0607161">De Armas, Luis F.</mods:namePart>
</mods:name>
<mods:name id="30CD77119A5FC260B239FF9100C4FAD0" type="personal">
<mods:role id="D5B6434CA200FE3529668E80B6843E56">
<mods:roleTerm id="27546FA1B54F564DC4AFCC4483A82508">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5DF2F28F560EFD937DA2A07A5C9B260D">Manzanilla, Osvaldo Villarreal</mods:namePart>
</mods:name>
<mods:name id="31D7E7CD1608911748EDFEDA396667A1" type="personal">
<mods:role id="692E625C5AD0A40166D4E13F01CA19C9">
<mods:roleTerm id="F130F748CC5BE1EAEFA43C36571F36C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B6D21EA13F771CF98B20A188DE99367">Viquez, Carlos</mods:namePart>
</mods:name>
<mods:typeOfResource id="99442ABE143BA5EA038C54D1A07F7012">text</mods:typeOfResource>
<mods:relatedItem id="F8EF8A1D6DE06F2CA6633ACE034288BA" type="host">
<mods:titleInfo id="38B9D86BF8AB7D2081E147C70A32F94B">
<mods:title id="9E323D96D2D56D6A65E41E4FFEF62989">Papéis Avulsos de Zoologia</mods:title>
</mods:titleInfo>
<mods:part id="A086F9E9CC99F0457BE65B621D8659E7">
<mods:date id="7E49128E40CE51962AF4D1626D225930">2010</mods:date>
<mods:detail id="88536450DEBD687D2E87B42FD24660F2" type="pubDate">
<mods:number id="14885F6C849C8987E6A41459C5B114EA">2010-12-31</mods:number>
</mods:detail>
<mods:detail id="CCAB778A0D59B9E57D0C138FD3CB2808" type="volume">
<mods:number id="B683E325DF910C3F786D90C866E837B5">50</mods:number>
</mods:detail>
<mods:detail id="E4099DFD1EFCC214BBD8A6D3663F046C" type="issue">
<mods:number id="335FC2609C9286BCBDE33C41E29C4CE4">37</mods:number>
</mods:detail>
<mods:extent id="E7B435650532DB78F84B0FDB610D374A" unit="page">
<mods:start id="2D7A525A59A596292DF8211453276250">579</mods:start>
<mods:end id="CD6DBA076E2926992AAB1033F3936E18">586</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6B66AA930AF330710CD016130D0C0B2F">journal article</mods:classification>
<mods:identifier id="AEF64D174A4BEA40FF267FC028FA6425" type="DOI">10.1590/S0031-10492010003700001</mods:identifier>
<mods:identifier id="9D38363CFC359C3A85362E57424D048B" type="ISSN">1807-0205</mods:identifier>
<mods:identifier id="3C69EF6A2AC88B8422E977E1604E363F" type="Zenodo-Dep">13307773</mods:identifier>
</mods:mods>
<treatment id="039087ED891588607AAAD421F6E9FC88" ID-DOI="http://doi.org/10.5281/zenodo.12685403" ID-Zenodo-Dep="12685403" LSID="urn:lsid:plazi:treatment:039087ED891588607AAAD421F6E9FC88" httpUri="http://treatment.plazi.org/id/039087ED891588607AAAD421F6E9FC88" lastPageId="3" lastPageNumber="582" pageId="1" pageNumber="580">
<subSubSection id="C3236570891588627AAAD421F5F3FC08" box="[237,500,887,912]" pageId="1" pageNumber="580" type="nomenclature">
<paragraph id="8B8636FB891588627AAAD421F5F3FC08" blockId="1.[200,537,887,944]" box="[237,500,887,912]" pageId="1" pageNumber="580">
<heading id="D0CE8197891588627AAAD421F5F3FC08" bold="true" box="[237,500,887,912]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="580" reason="2">
<emphasis id="B94DEAE9891588627AAAD421F5F3FC08" bold="true" box="[237,500,887,912]" pageId="1" pageNumber="580">
<taxonomicName id="4C394D78891588627AAAD421F59CFC17" ID-CoL="53H7S" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[237,411,887,912]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="1" pageNumber="580" phylum="Arthropoda" rank="species" species="brus" status="sp. nov.">
<emphasis id="B94DEAE9891588627AAAD421F59CFC17" bold="true" box="[237,411,887,912]" italics="true" pageId="1" pageNumber="580">Surazomus brus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E5792891588627BE6D421F5F3FC08" box="[417,500,888,912]" pageId="1" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3236570891588607A8FD4C1F6E9FC88" lastPageId="3" lastPageNumber="582" pageId="1" pageNumber="580" type="description">
<paragraph id="8B8636FB891588627A8FD4C1F61EFC28" blockId="1.[200,537,887,944]" box="[200,537,920,944]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE9891588627A8FD4C1F61EFC28" bold="true" box="[200,537,920,944]" pageId="1" pageNumber="580">
(
<figureCitation id="13022A7E891588627A88D4C1F513FC28" box="[207,276,920,944]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="1" pageNumber="580">Figs. 1</figureCitation>
A-E, 2A, 3, 4A, 
<tableCitation id="C6BB0340891588627B87D4C1F617FC28" box="[448,528,920,944]" captionStart="TABLA 1" captionStartId="1.[95,161,1366,1385]" captionText="TABLA 1: Dimensiones (mm) de tres nuevas especies costarricenses del género Surazomus. A. ancho; H, alto; L, longitud." pageId="1" pageNumber="580">Tabla 1</tableCitation>
)
</emphasis>
</paragraph>
<paragraph id="8B8636FB891588627A19D481F62AFBC8" blockId="1.[94,643,984,1104]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE9891588627A19D481F4EEFC68" box="[94,233,984,1008]" italics="true" pageId="1" pageNumber="580">Material tipo:</emphasis>
<materialsCitation id="3B513CA6891588627AB7D481F62AFBC8" collectionCode="INBio, R" collectorName="Estacion Biologica Las Cruces &amp; Punta Arenas &amp; L. F. Armas &amp; C. Viquez &amp; Gutierrez" pageId="1" pageNumber="580" specimenCount="1" typeStatus="holotype">
Macho 
<typeStatus id="54828859891588627B07D481F59FFC68" box="[320,408,984,1008]" pageId="1" pageNumber="580" type="holotype">holotipo</typeStatus>
(
<collectionCode id="ED28AE3E891588627BE0D481F5EBFC68" box="[423,492,984,1008]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="1" pageNumber="580">INBio</collectionCode>
, 
<collectorName id="26CC532D891588627BB0D481F515FB88" pageId="1" pageNumber="580">Estación Biológica Las Cruces</collectorName>
(1 100 msnm), 
<taxonomicName id="4C394D78891588627BF9D4A1F62BFB97" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[446,556,1016,1040]" class="Arachnida" family="Hubbardiidae" genus="Coto" kingdom="Animalia" order="Schizomida" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="brus">Coto Brus</taxonomicName>
, provincia 
<collectorName id="26CC532D891588627AC6D341F50BFBA8" box="[129,268,1048,1072]" pageId="1" pageNumber="580">Punta Arenas</collectorName>
, 3-4 de agosto, 2005, 
<collectorName id="26CC532D891588627BA9D341F667FBA8" box="[494,608,1048,1072]" pageId="1" pageNumber="580">L.F. Armas</collectorName>
, 
<collectorName id="26CC532D89158862782ED341F4A1FBC8" pageId="1" pageNumber="580">C. Víquez</collectorName>
&amp; 
<collectionCode id="ED28AE3E891588627A8DD361F4D9FBD7" box="[202,222,1080,1103]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="580">R</collectionCode>
. 
<collectorName id="26CC532D891588627AAFD361F54BFBC8" box="[232,332,1080,1104]" pageId="1" pageNumber="580">Gutiérrez</collectorName>
, bajo tronco podrido.
</materialsCitation>
</paragraph>
<paragraph id="8B8636FB891588627A19D320F4BAFB28" blockId="1.[94,642,1144,1200]" pageId="1" pageNumber="580">
<materialsCitation id="3B513CA6891588627A19D320F4BEFB28" collectionCode="INBio" pageId="1" pageNumber="580" specimenCount="2" typeStatus="Paratipo">
<emphasis id="B94DEAE9891588627A19D320F4BFFB08" box="[94,184,1145,1168]" italics="true" pageId="1" pageNumber="580">
<typeStatus id="54828859891588627A19D320F4B5FB08" box="[94,178,1145,1168]" pageId="1" pageNumber="580" type="paratype">Paratipo</typeStatus>
:
</emphasis>
Hembra (
<collectionCode id="ED28AE3E891588627B7FD321F579FB08" box="[312,382,1144,1168]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="1" pageNumber="580">INBio</collectionCode>
), iguales datos que el 
<typeStatus id="54828859891588627A19D3C1F4BEFB28" box="[94,185,1176,1200]" pageId="1" pageNumber="580" type="holotype">holotipo</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8636FB891588627A19D381F4B6FA88" blockId="1.[94,643,1240,1296]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE9891588627A19D381F4E5FB68" box="[94,226,1240,1264]" italics="true" pageId="1" pageNumber="580">Distribución:</emphasis>
Solo se conoce de la localidad tipo (
<figureCitation id="13022A7E891588627A21D3A1F4A3FA88" box="[102,164,1272,1296]" captionStart="FIGURA 3" captionStartId="3.[680,760,1826,1845]" captionTargetBox="[680,1228,1298,1789]" captionTargetId="figure-500@3.[680,1228,1298,1789]" captionTargetPageId="3" captionText="FIGURA 3: Distribución geográfica del género Surazomus en Costa Rica." figureDoi="http://doi.org/10.5281/zenodo.13307779" httpUri="https://zenodo.org/record/13307779/files/figure.png" pageId="1" pageNumber="580">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8B8636FB8915886278EFD7E1F702FF68" blockId="1.[680,1228,184,240]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE98915886278EFD7E1F71EFF48" box="[680,793,184,208]" italics="true" pageId="1" pageNumber="580">Etimología:</emphasis>
Nombre en aposición referido a la localidad tipo.
</paragraph>
<paragraph id="8B8636FB8915886278EFD640F7DEFCC8" blockId="1.[680,1229,280,848]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE98915886278EFD640F70AFEA8" box="[680,781,281,304]" italics="true" pageId="1" pageNumber="580">Diagnosis:</emphasis>
Macho 3,0 mm de longitud total; hembra 
<quantity id="4CC19B1E8915886278EFD661F6F8FED7" box="[680,767,312,336]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="580" unit="mm" value="3.5">3,5 mm</quantity>
; cuerpo de castaño verdoso oscuro casi uniforme; patela I con la mitad distal blanquecina. Propeltidio con cuatro pares de cerdas dorsales, manchas oculares prominentes, redondeadas. Metapeltidio dividido por una débil sutura longitudinal media. Esternón anterior con 10 + 2 cerdas. Fémur y patela del pedipalpo con sendos espolones bien desarrollados (
<figureCitation id="13022A7E89158862789FD541F71FFDA8" box="[728,792,536,560]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="1" pageNumber="580">Figs. 1</figureCitation>
A-B). Dedo móvil del quelícero sin dientes accesorios, con lamela (
<figureCitation id="13022A7E8915886279E5D561F7F1FDC8" box="[930,1014,568,592]" captionStart="FIGURA 2" captionStartId="3.[95,175,1826,1845]" captionTargetBox="[94,642,1110,1788]" captionTargetId="figure-472@3.[94,642,1109,1789]" captionTargetPageId="3" captionText="FIGURA 2: Detalle del dedo móvil del quelícero. A, Surazomus brus sp. nov., macho holotipo; B, S. selva, sp. nov., macho holotipo." figureDoi="http://doi.org/10.5281/zenodo.13307777" httpUri="https://zenodo.org/record/13307777/files/figure.png" pageId="1" pageNumber="580">Fig. 2A</figureCitation>
). Flagelo del macho trilobulado, con las cerdas 
<emphasis id="B94DEAE9891588627982D501F7EFFDED" box="[965,1000,600,629]" italics="true" pageId="1" pageNumber="580">
Dl 
<subScript id="17BD34BE8915886279A7D53EF7EFFDED" attach="left" box="[992,1000,615,629]" fontSize="6" pageId="1" pageNumber="580">1</subScript>
</emphasis>
y 
<emphasis id="B94DEAE9891588627E41D501F022FDED" box="[1030,1061,600,629]" italics="true" pageId="1" pageNumber="580">
Vl 
<subScript id="17BD34BE891588627E5AD53EF022FDED" attach="left" box="[1053,1061,615,629]" fontSize="6" pageId="1" pageNumber="580">1</subScript>
</emphasis>
modificadas en forma de alfanje; borde anterior del bulbo anguloso; áreas laterales y proximal de la depresión dorsomedial blanquecina (
<figureCitation id="13022A7E891588627975D5E1F784FD48" box="[818,899,696,720]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="1" pageNumber="580">Fig. 1D</figureCitation>
). Espermatecas sin arco quitinoso ni gonópodo; lóbulos medios que terminan en sendos bulbos bien desarrollados; lóbulos laterales más cortos que los medios y con el bulbo terminal mucho menos desarrollado (
<figureCitation id="13022A7E89158862793AD461F7C9FCC8" box="[893,974,824,848]" captionStart="FIGURA 4" captionStartId="4.[132,212,1851,1870]" captionTargetBox="[133,677,1608,1815]" captionTargetId="figure-694@4.[132,680,1607,1815]" captionTargetPageId="4" captionText="FIGURA 4: Surazomus brus sp. nov. Espermatecas." figureDoi="http://doi.org/10.5281/zenodo.13307781" httpUri="https://zenodo.org/record/13307781/files/figure.png" pageId="1" pageNumber="580">Fig. 4A</figureCitation>
).
</paragraph>
<paragraph id="8B8636FB8915886278EFD421F0C1FBE8" blockId="1.[680,1229,888,1296]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE98915886278EFD421F7E2FC08" box="[680,997,888,912]" italics="true" pageId="1" pageNumber="580">
Descripción del macho 
<typeStatus id="548288598915886279D7D421F7E7FC08" box="[912,992,888,912]" pageId="1" pageNumber="580" type="holotype">holotipo</typeStatus>
:
</emphasis>
Coloración. De color castaño verdoso oscuro, ligeramente más pálido en las patas y el metapeltidio; patela de la pata I con la mitad distal blanquecina. Propeltidio cuatro pares de cerdas dorsales; manchas oculares notables, redondeadas a ligeramente ovoides. Metapeltidio con una sutura longitudinal media poco notable. Esternón anterior con 10 + 2 cerdas; esternón posterior con seis cerdas.
</paragraph>
<paragraph id="8B8636FB8915886178A6D321F6AEFE88" blockId="1.[680,1229,888,1296]" lastBlockId="2.[132,681,184,784]" lastPageId="2" lastPageNumber="581" pageId="1" pageNumber="580">Abdomen: Terguitos I-VII con dos cerdas posteriores submedias; VIII con dos cerdas posteriores submedias y dos laterales; IX con cuatro cerdas posterolaterales. Segmentos X-XI con dos cerdas posterolaterales. Segmento XII con dos cerdas dorsoposteriores submedias, muy fuertes y cortas, espiniformes, más un par lateral; eminencia dorsoposterior media bien desarrollada, con un par de tubérculos cónicos submedios.</paragraph>
<caption id="DF466673891588627A18D20FF04DFAF1" box="[95,1098,1366,1385]" pageId="1" pageNumber="580" startId="1.[95,161,1366,1385]" targetType="table">
<paragraph id="8B8636FB891588627A18D20FF04DFAF1" blockId="1.[95,1098,1366,1385]" box="[95,1098,1366,1385]" pageId="1" pageNumber="580">
<emphasis id="B94DEAE9891588627A18D20FF4B0FAF1" bold="true" box="[95,183,1366,1385]" pageId="1" pageNumber="580">TABLA 1:</emphasis>
Dimensiones (mm) de tres nuevas especies costarricenses del género 
<emphasis id="B94DEAE98915886278AAD20FF74EFAF1" box="[749,841,1366,1385]" italics="true" pageId="1" pageNumber="580">
<taxonomicName id="4C394D788915886278AAD20FF742FAF1" authorityName="REDDELL &amp; COKENDOLPHER" authorityYear="1995" box="[749,837,1366,1385]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="1" pageNumber="580" phylum="Arthropoda" rank="genus">Surazomus</taxonomicName>
.
</emphasis>
A. ancho; H, alto; L, longitud.
</paragraph>
</caption>
<paragraph id="8B8636FB891588627AE2D2CEF083F8DF" pageId="1" pageNumber="580">
<table id="F939C45B8915779C7A19D2DEF09DF8DF" box="[94,1178,1415,1863]" gridcols="5" gridrows="14" pageId="1" pageNumber="580">
<tr id="350934B98915779C7A19D2DEF09DFA23" box="[94,1178,1415,1467]" gridrow="0" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D2DEF4FDFA23" box="[94,250,1415,1467]" gridcol="0" gridrow="0" pageId="1" pageNumber="580">Caracteres</th>
<th id="76D85DC58915779C7B33D2DEF5F4FA23" box="[372,499,1415,1467]" gridcol="1" gridrow="0" pageId="1" pageNumber="580">
<taxonomicName id="4C394D78891588627B33D2DEF5A9FA02" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[372,430,1415,1434]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="1" pageNumber="580" phylum="Arthropoda" rank="species" species="selva" status="sp. nov.">
<emphasis id="B94DEAE9891588627B33D2DEF5A9FA02" box="[372,430,1415,1434]" italics="true" pageId="1" pageNumber="580">S. selva</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E5792891588627BF4D2D1F5F4FA03" box="[435,499,1416,1435]" pageId="1" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
♂ Holotipo
</th>
<th id="76D85DC58915779C787DD2DEF6F5FA23" box="[570,754,1415,1467]" gridcol="2" gridrow="0" pageId="1" pageNumber="580">
<taxonomicName id="4C394D7889158862787DD2DEF6A9FA02" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[570,686,1415,1434]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="1" pageNumber="580" phylum="Arthropoda" rank="species" species="inexpectatus" status="sp. nov.">
<emphasis id="B94DEAE989158862787DD2DEF6A9FA02" box="[570,686,1415,1434]" italics="true" pageId="1" pageNumber="580">S. inexpectatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57928915886278F4D2D1F6F5FA03" box="[691,754,1416,1435]" pageId="1" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
♂ Holotipo
</th>
<th id="76D85DC58915779C797ED2DEF09DFA23" box="[825,1178,1415,1467]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="1" pageNumber="580">
<taxonomicName id="4C394D788915886279EAD2DEF7E4FA02" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[941,995,1415,1434]" class="Arachnida" family="Hubbardiidae" genus="Coto" kingdom="Animalia" order="Schizomida" pageId="1" pageNumber="580" phylum="Arthropoda" rank="species" species="brus" status="sp. nov.">
<emphasis id="B94DEAE98915886279EAD2DEF7E4FA02" box="[941,995,1415,1434]" italics="true" pageId="1" pageNumber="580">S. brus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E57928915886279AFD2D1F02FFA03" box="[1000,1064,1416,1435]" pageId="1" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
♂ Holotipo ♀ Paratipo
</th>
</tr>
<tr id="350934B98915779C7A19D29DF09DFA40" box="[94,1178,1476,1496]" gridrow="1" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D29DF4FDFA40" box="[94,250,1476,1496]" gridcol="0" gridrow="1" pageId="1" pageNumber="580">L total</th>
<td id="76D85DC58915779C7B33D29DF5F4FA40" box="[372,499,1476,1496]" gridcol="1" gridrow="1" pageId="1" pageNumber="580">2,75</td>
<td id="76D85DC58915779C787DD29DF6F5FA40" box="[570,754,1476,1496]" gridcol="2" gridrow="1" pageId="1" pageNumber="580">2,80</td>
<td id="76D85DC58915779C797ED29DF7B0FA40" box="[825,951,1476,1496]" gridcol="3" gridrow="1" pageId="1" pageNumber="580">3,00</td>
<td id="76D85DC58915779C7E5BD29DF09DFA40" box="[1052,1178,1476,1496]" gridcol="4" gridrow="1" pageId="1" pageNumber="580">3,50</td>
</tr>
<tr id="350934B98915779C7A19D2BAF09DFA6E" box="[94,1178,1507,1526]" gridrow="2" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D2BAF4FDFA6E" box="[94,250,1507,1526]" gridcol="0" gridrow="2" pageId="1" pageNumber="580">Propeltidio, L/A</th>
<td id="76D85DC58915779C7B33D2BAF5F4FA6E" box="[372,499,1507,1526]" gridcol="1" gridrow="2" pageId="1" pageNumber="580">0,94/0,49</td>
<td id="76D85DC58915779C787DD2BAF6F5FA6E" box="[570,754,1507,1526]" gridcol="2" gridrow="2" pageId="1" pageNumber="580">1,04/0,57</td>
<td id="76D85DC58915779C797ED2BAF7B0FA6E" box="[825,951,1507,1526]" gridcol="3" gridrow="2" pageId="1" pageNumber="580">1,04/0,55</td>
<td id="76D85DC58915779C7E5BD2BAF09DFA6E" box="[1052,1178,1507,1526]" gridcol="4" gridrow="2" pageId="1" pageNumber="580">1,09/0,65</td>
</tr>
<tr id="350934B98915779C7A19D158F09DF98D" box="[94,1178,1537,1557]" gridrow="3" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D158F4FDF98D" box="[94,250,1537,1557]" gridcol="0" gridrow="3" pageId="1" pageNumber="580">Flagelo, L/A/H</th>
<td id="76D85DC58915779C7B33D158F5F4F98D" box="[372,499,1537,1557]" gridcol="1" gridrow="3" pageId="1" pageNumber="580">0,31/0,34/0,18</td>
<td id="76D85DC58915779C787DD158F6F5F98D" box="[570,754,1537,1557]" gridcol="2" gridrow="3" pageId="1" pageNumber="580">0,44/0,49/0,23</td>
<td id="76D85DC58915779C797ED158F7B0F98D" box="[825,951,1537,1557]" gridcol="3" gridrow="3" pageId="1" pageNumber="580">0,42/0,42/0,25</td>
<td id="76D85DC58915779C7E5BD158F09DF98D" box="[1052,1178,1537,1557]" gridcol="4" gridrow="3" pageId="1" pageNumber="580">0,26/0,08/0,08</td>
</tr>
<tr id="350934B98915779C7A19D179F09DF9AC" box="[94,1178,1568,1588]" gridrow="4" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D179F4FDF9AC" box="[94,250,1568,1588]" gridcol="0" gridrow="4" pageId="1" pageNumber="580">Pedipalpo, L</th>
<td id="76D85DC58915779C7B33D179F5F4F9AC" box="[372,499,1568,1588]" gridcol="1" gridrow="4" pageId="1" pageNumber="580">1,85</td>
<td id="76D85DC58915779C787DD179F6F5F9AC" box="[570,754,1568,1588]" gridcol="2" gridrow="4" pageId="1" pageNumber="580">2,10</td>
<td id="76D85DC58915779C797ED179F7B0F9AC" box="[825,951,1568,1588]" gridcol="3" gridrow="4" pageId="1" pageNumber="580">2,05</td>
<td id="76D85DC58915779C7E5BD179F09DF9AC" box="[1052,1178,1568,1588]" gridcol="4" gridrow="4" pageId="1" pageNumber="580">1,79</td>
</tr>
<tr id="350934B98915779C7A19D166F09DF9CA" box="[94,1178,1599,1618]" gridrow="5" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D166F4FDF9CA" box="[94,250,1599,1618]" gridcol="0" gridrow="5" pageId="1" pageNumber="580">Trocánter, L/H</th>
<td id="76D85DC58915779C7B33D166F5F4F9CA" box="[372,499,1599,1618]" gridcol="1" gridrow="5" pageId="1" pageNumber="580">0,42/0,21</td>
<td id="76D85DC58915779C787DD166F6F5F9CA" box="[570,754,1599,1618]" gridcol="2" gridrow="5" pageId="1" pageNumber="580">0,47/0,23</td>
<td id="76D85DC58915779C797ED166F7B0F9CA" box="[825,951,1599,1618]" gridcol="3" gridrow="5" pageId="1" pageNumber="580">0,47/0,21</td>
<td id="76D85DC58915779C7E5BD166F09DF9CA" box="[1052,1178,1599,1618]" gridcol="4" gridrow="5" pageId="1" pageNumber="580">0,39/0,23</td>
</tr>
<tr id="350934B98915779C7A19D104F09DF9E8" box="[94,1178,1629,1648]" gridrow="6" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D104F4FDF9E8" box="[94,250,1629,1648]" gridcol="0" gridrow="6" pageId="1" pageNumber="580">Fémur, L/H</th>
<td id="76D85DC58915779C7B33D104F5F4F9E8" box="[372,499,1629,1648]" gridcol="1" gridrow="6" pageId="1" pageNumber="580">0,42/0,23</td>
<td id="76D85DC58915779C787DD104F6F5F9E8" box="[570,754,1629,1648]" gridcol="2" gridrow="6" pageId="1" pageNumber="580">0,49/0,29</td>
<td id="76D85DC58915779C797ED104F7B0F9E8" box="[825,951,1629,1648]" gridcol="3" gridrow="6" pageId="1" pageNumber="580">0,49/0,26</td>
<td id="76D85DC58915779C7E5BD104F09DF9E8" box="[1052,1178,1629,1648]" gridcol="4" gridrow="6" pageId="1" pageNumber="580">0,44/0,25</td>
</tr>
<tr id="350934B98915779C7A19D125F09DF917" box="[94,1178,1660,1679]" gridrow="7" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D125F4FDF917" box="[94,250,1660,1679]" gridcol="0" gridrow="7" pageId="1" pageNumber="580">Patela, L/H</th>
<td id="76D85DC58915779C7B33D125F5F4F917" box="[372,499,1660,1679]" gridcol="1" gridrow="7" pageId="1" pageNumber="580">0,52/0,16</td>
<td id="76D85DC58915779C787DD125F6F5F917" box="[570,754,1660,1679]" gridcol="2" gridrow="7" pageId="1" pageNumber="580">0,52/0,21</td>
<td id="76D85DC58915779C797ED125F7B0F917" box="[825,951,1660,1679]" gridcol="3" gridrow="7" pageId="1" pageNumber="580">0,52/0,18</td>
<td id="76D85DC58915779C7E5BD125F09DF917" box="[1052,1178,1660,1679]" gridcol="4" gridrow="7" pageId="1" pageNumber="580">0,44/0,18</td>
</tr>
<tr id="350934B98915779C7A19D1C3F09DF936" box="[94,1178,1690,1710]" gridrow="8" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D1C3F4FDF936" box="[94,250,1690,1710]" gridcol="0" gridrow="8" pageId="1" pageNumber="580">Tibia, L</th>
<td id="76D85DC58915779C7B33D1C3F5F4F936" box="[372,499,1690,1710]" gridcol="1" gridrow="8" pageId="1" pageNumber="580">0,31</td>
<td id="76D85DC58915779C787DD1C3F6F5F936" box="[570,754,1690,1710]" gridcol="2" gridrow="8" pageId="1" pageNumber="580">0,39</td>
<td id="76D85DC58915779C797ED1C3F7B0F936" box="[825,951,1690,1710]" gridcol="3" gridrow="8" pageId="1" pageNumber="580">0,39</td>
<td id="76D85DC58915779C7E5BD1C3F09DF936" box="[1052,1178,1690,1710]" gridcol="4" gridrow="8" pageId="1" pageNumber="580">0,34</td>
</tr>
<tr id="350934B98915779C7A19D1E0F09DF955" box="[94,1178,1721,1741]" gridrow="9" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D1E0F4FDF955" box="[94,250,1721,1741]" gridcol="0" gridrow="9" pageId="1" pageNumber="580">Tarso, L</th>
<td id="76D85DC58915779C7B33D1E0F5F4F955" box="[372,499,1721,1741]" gridcol="1" gridrow="9" pageId="1" pageNumber="580">0,18</td>
<td id="76D85DC58915779C787DD1E0F6F5F955" box="[570,754,1721,1741]" gridcol="2" gridrow="9" pageId="1" pageNumber="580">0,23</td>
<td id="76D85DC58915779C797ED1E0F7B0F955" box="[825,951,1721,1741]" gridcol="3" gridrow="9" pageId="1" pageNumber="580">0,18</td>
<td id="76D85DC58915779C7E5BD1E0F09DF955" box="[1052,1178,1721,1741]" gridcol="4" gridrow="9" pageId="1" pageNumber="580">0,18</td>
</tr>
<tr id="350934B98915779C7A19D181F09DF973" box="[94,1178,1752,1771]" gridrow="10" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D181F4FDF973" box="[94,250,1752,1771]" gridcol="0" gridrow="10" pageId="1" pageNumber="580">Pata I, L</th>
<td id="76D85DC58915779C7B33D181F5F4F973" box="[372,499,1752,1771]" gridcol="1" gridrow="10" pageId="1" pageNumber="580">4,00</td>
<td id="76D85DC58915779C787DD181F6F5F973" box="[570,754,1752,1771]" gridcol="2" gridrow="10" pageId="1" pageNumber="580">4,53</td>
<td id="76D85DC58915779C797ED181F7B0F973" box="[825,951,1752,1771]" gridcol="3" gridrow="10" pageId="1" pageNumber="580">4,03</td>
<td id="76D85DC58915779C7E5BD181F09DF973" box="[1052,1178,1752,1771]" gridcol="4" gridrow="10" pageId="1" pageNumber="580">3,42</td>
</tr>
<tr id="350934B98915779C7A19D1AFF09DF892" box="[94,1178,1782,1802]" gridrow="11" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D1AFF4FDF892" box="[94,250,1782,1802]" gridcol="0" gridrow="11" pageId="1" pageNumber="580">Fémur, L</th>
<td id="76D85DC58915779C7B33D1AFF5F4F892" box="[372,499,1782,1802]" gridcol="1" gridrow="11" pageId="1" pageNumber="580">1,01</td>
<td id="76D85DC58915779C787DD1AFF6F5F892" box="[570,754,1782,1802]" gridcol="2" gridrow="11" pageId="1" pageNumber="580">1,09</td>
<td id="76D85DC58915779C797ED1AFF7B0F892" box="[825,951,1782,1802]" gridcol="3" gridrow="11" pageId="1" pageNumber="580">0,99</td>
<td id="76D85DC58915779C7E5BD1AFF09DF892" box="[1052,1178,1782,1802]" gridcol="4" gridrow="11" pageId="1" pageNumber="580">0,91</td>
</tr>
<tr id="350934B98915779C7A19D04CF09DF8B0" box="[94,1178,1813,1832]" gridrow="12" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D04CF4FDF8B0" box="[94,250,1813,1832]" gridcol="0" gridrow="12" pageId="1" pageNumber="580">Pata IV, L</th>
<td id="76D85DC58915779C7B33D04CF5F4F8B0" box="[372,499,1813,1832]" gridcol="1" gridrow="12" pageId="1" pageNumber="580">3,25</td>
<td id="76D85DC58915779C787DD04CF6F5F8B0" box="[570,754,1813,1832]" gridcol="2" gridrow="12" pageId="1" pageNumber="580">3,64</td>
<td id="76D85DC58915779C797ED04CF7B0F8B0" box="[825,951,1813,1832]" gridcol="3" gridrow="12" pageId="1" pageNumber="580">3,30</td>
<td id="76D85DC58915779C7E5BD04CF09DF8B0" box="[1052,1178,1813,1832]" gridcol="4" gridrow="12" pageId="1" pageNumber="580">3,27</td>
</tr>
<tr id="350934B98915779C7A19D06DF09DF8DF" box="[94,1178,1844,1863]" gridrow="13" pageId="1" pageNumber="580">
<th id="76D85DC58915779C7A19D06DF4FDF8DF" box="[94,250,1844,1863]" gridcol="0" gridrow="13" pageId="1" pageNumber="580">Fémur, L/H</th>
<td id="76D85DC58915779C7B33D06DF5F4F8DF" box="[372,499,1844,1863]" gridcol="1" gridrow="13" pageId="1" pageNumber="580">1,04/0,49</td>
<td id="76D85DC58915779C787DD06DF6F5F8DF" box="[570,754,1844,1863]" gridcol="2" gridrow="13" pageId="1" pageNumber="580">1,12/0,49</td>
<td id="76D85DC58915779C797ED06DF7B0F8DF" box="[825,951,1844,1863]" gridcol="3" gridrow="13" pageId="1" pageNumber="580">1,01/0,47</td>
<td id="76D85DC58915779C7E5BD06DF09DF8DF" box="[1052,1178,1844,1863]" gridcol="4" gridrow="13" pageId="1" pageNumber="580">0,99/0,44</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8636FB891688617AFAD641F639FE48" blockId="2.[132,681,184,784]" pageId="2" pageNumber="581">
Flagelo: Trilobulado (
<figureCitation id="13022A7E891688617BE5D641F5E0FEA8" box="[418,487,280,304]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="2" pageNumber="581">Figs. 1</figureCitation>
C-E), ligeramente más largo que ancho y con el pedúnculo muy corto; el borde anterior del bulbo forma un ángulo obtuso de aproximadamente 140°; depresión dorsal media flanqueada por un par de eminencias blanquecinas; cerdas 
<emphasis id="B94DEAE9891688617AC3D6E1F4A0FE4D" box="[132,167,440,469]" italics="true" pageId="2" pageNumber="581">
Dl 
<subScript id="17BD34BE891688617AD8D69EF4A0FE4D" attach="left" box="[159,167,455,469]" fontSize="6" pageId="2" pageNumber="581">1</subScript>
</emphasis>
y 
<emphasis id="B94DEAE9891688617A87D6E1F4E5FE4D" box="[192,226,440,469]" italics="true" pageId="2" pageNumber="581">
Dl 
<subScript id="17BD34BE891688617A9DD69EF4E5FE4D" attach="left" box="[218,226,455,469]" fontSize="6" pageId="2" pageNumber="581">2</subScript>
</emphasis>
modificadas, en forma de alfanje.
</paragraph>
<paragraph id="8B8636FB891688617AFAD681F63FFD28" blockId="2.[132,681,184,784]" pageId="2" pageNumber="581">
Pedipalpo (
<figureCitation id="13022A7E891688617B75D681F589FE68" box="[306,398,472,496]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="2" pageNumber="581">Figs. 1D</figureCitation>
, 
<figureCitation id="13022A7E891688617BDCD681F5B9FE68" box="[411,446,472,496]" captionStart="FIGURA 2" captionStartId="3.[95,175,1826,1845]" captionTargetBox="[94,642,1110,1788]" captionTargetId="figure-472@3.[94,642,1109,1789]" captionTargetPageId="3" captionText="FIGURA 2: Detalle del dedo móvil del quelícero. A, Surazomus brus sp. nov., macho holotipo; B, S. selva, sp. nov., macho holotipo." figureDoi="http://doi.org/10.5281/zenodo.13307777" httpUri="https://zenodo.org/record/13307777/files/figure.png" pageId="2" pageNumber="581">2A</figureCitation>
): Trocánter con un diminuto espolón interno y el ápice prominente; fémur corto y robusto, con un fuerte espolón anteroventral, ligeramente dirigido hacia arriba (
<figureCitation id="13022A7E891688617BAAD561F638FDC8" box="[493,575,568,592]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="2" pageNumber="581">Fig. 1A</figureCitation>
), con dos cerdas espiniformes en la superficie interna; patela con un espolón distal interno que es aproximadamente 0,3 veces tan largo como la tibia (
<figureCitation id="13022A7E891688617B9BD5C1F62DFD28" box="[476,554,664,688]" captionStart="FIGURA 1" captionStartId="2.[132,212,1826,1845]" captionTargetBox="[137,1262,828,1786]" captionTargetId="figure-277@2.[132,1266,826,1790]" captionTargetPageId="2" captionText="FIGURA 1: Surazomus brus sp. nov. A-E, Macho holotipo. A-B, pedipalpo: A, vista externa; B, espolón patelar; C-E, flagelo, vistas lateral (C), dorsal (D) y ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="2" pageNumber="581">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8B8636FB891688617AFAD5E1F75FFEA8" blockId="2.[132,681,184,784]" lastBlockId="2.[718,1266,184,336]" pageId="2" pageNumber="581">
Quelícero: Dedo fijo con seis dientes accesorios entre los dos mayores; dedo móvil sin dientes accesorios, con una lamela (
<figureCitation id="13022A7E891688617BC4D5A1F5D1FC88" box="[387,470,760,784]" captionStart="FIGURA 2" captionStartId="3.[95,175,1826,1845]" captionTargetBox="[94,642,1110,1788]" captionTargetId="figure-472@3.[94,642,1109,1789]" captionTargetPageId="3" captionText="FIGURA 2: Detalle del dedo móvil del quelícero. A, Surazomus brus sp. nov., macho holotipo; B, S. selva, sp. nov., macho holotipo." figureDoi="http://doi.org/10.5281/zenodo.13307777" httpUri="https://zenodo.org/record/13307777/files/figure.png" pageId="2" pageNumber="581">Fig. 2A</figureCitation>
), sérrula compuesta por 15 dientes que terminan en un diente guardián grande; cerdas de la serie 1 con el tallo casi desnudo, solo con tubérculos dispersos muy pequeños, no espiniformes.
</paragraph>
<paragraph id="8B8636FB891688617940D661F7FFFEC8" blockId="2.[718,1266,184,336]" box="[775,1016,312,336]" pageId="2" pageNumber="581">
Dimensiones (
<tableCitation id="C6BB03408916886179DCD661F7EFFEC8" box="[923,1000,312,336]" captionStart="TABLA 1" captionStartId="1.[95,161,1366,1385]" captionText="TABLA 1: Dimensiones (mm) de tres nuevas especies costarricenses del género Surazomus. A. ancho; H, alto; L, longitud." pageId="2" pageNumber="581">Tabla 1</tableCitation>
).
</paragraph>
<paragraph id="8B8636FB891688617889D621F7B7FE48" blockId="2.[718,1266,376,656]" pageId="2" pageNumber="581">
<emphasis id="B94DEAE9891688617889D621F722FE08" box="[718,805,376,400]" italics="true" pageId="2" pageNumber="581">Hembra:</emphasis>
Similar al macho, pero con el flagelo trisegmentado y los pedipalpos no modificados, sin espolones en fémur y patela.
</paragraph>
<paragraph id="8B8636FB891688617940D681F757FDE8" blockId="2.[718,1266,376,656]" pageId="2" pageNumber="581">
Espermatecas: Sin arco quitinoso ni gonópodo; lóbulos medios más largos que los laterales, con el bulbo terminal algo mayor que el de estos (
<figureCitation id="13022A7E891688617ED2D541F0E1FDA8" box="[1173,1254,536,560]" captionStart="FIGURA 4" captionStartId="4.[132,212,1851,1870]" captionTargetBox="[133,677,1608,1815]" captionTargetId="figure-694@4.[132,680,1607,1815]" captionTargetPageId="4" captionText="FIGURA 4: Surazomus brus sp. nov. Espermatecas." figureDoi="http://doi.org/10.5281/zenodo.13307781" httpUri="https://zenodo.org/record/13307781/files/figure.png" pageId="2" pageNumber="581">Fig. 4A</figureCitation>
); todos los lóbulos con escasos poros irregularmente distribuidos.
</paragraph>
<paragraph id="8B8636FB891688617940D521F7FFFD08" blockId="2.[718,1266,376,656]" box="[775,1016,632,656]" pageId="2" pageNumber="581">
Dimensiones (
<tableCitation id="C6BB03408916886179DCD521F7EFFD08" box="[923,1000,632,656]" captionStart="TABLA 1" captionStartId="1.[95,161,1366,1385]" captionText="TABLA 1: Dimensiones (mm) de tres nuevas especies costarricenses del género Surazomus. A. ancho; H, alto; L, longitud." pageId="2" pageNumber="581">Tabla 1</tableCitation>
).
</paragraph>
<paragraph id="8B8636FB891688607889D5E0F583FE68" blockId="2.[718,1266,696,784]" lastBlockId="3.[94,643,184,496]" lastPageId="3" lastPageNumber="582" pageId="2" pageNumber="581">
<emphasis id="B94DEAE9891688617889D5E0F772FD48" box="[718,885,696,720]" italics="true" pageId="2" pageNumber="581">Historia natural:</emphasis>
El área de la Estación Biológica “Las Cruces” (1 100 msnm) está mayormente cubierta por pluvisilva. La precipitación promedio anual es de 4 000 mm y la temperatura varía entre 21 y 26°C durante el día y entre 15 y 21°C en las noches. La serie tipo de 
<taxonomicName id="4C394D78891788607AF4D7A1F4FDFE88" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[179,250,248,272]" class="Arachnida" family="Hubbardiidae" genus="Coto" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="brus" status="sp. nov.">
<emphasis id="B94DEAE9891788607AF4D7A1F4FDFE88" box="[179,250,248,272]" italics="true" pageId="3" pageNumber="582">S. brus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E5792891788607B44D7A1F553FE97" box="[259,340,248,271]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
fue recolectada bajo troncos podridos, a los lados del recorrido turístico que lleva del Jardín Botánico “Robert y Catherine Wilson” al río Java. A pesar del intenso esfuerzo de recolecta manual durante 21 horas/hombre, únicamente se vieron los dos especímenes que conforman la serie tipo y el macho de otra especie nueva, descrita más adelante, con la que vive en simpatría.
</paragraph>
<caption id="DF466673891688617AC3D07BF5ABF8D6" ID-DOI="http://doi.org/10.5281/zenodo.13307775" ID-Zenodo-Dep="13307775" httpUri="https://zenodo.org/record/13307775/files/figure.png" pageId="2" pageNumber="581" startId="2.[132,212,1826,1845]" targetBox="[137,1262,828,1786]" targetPageId="2" targetType="figure">
<paragraph id="8B8636FB891688617AC3D07BF5ABF8D6" blockId="2.[132,1267,1826,1871]" pageId="2" pageNumber="581">
<emphasis id="B94DEAE9891688617AC3D07BF4EAF8AD" bold="true" box="[132,237,1826,1845]" pageId="2" pageNumber="581">FIGURA 1:</emphasis>
<taxonomicName id="4C394D78891688617AB2D07BF572F8AD" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[245,373,1826,1845]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="2" pageNumber="581" phylum="Arthropoda" rank="species" species="brus" status="sp. nov.">
<emphasis id="B94DEAE9891688617AB2D07BF572F8AD" box="[245,373,1826,1845]" italics="true" pageId="2" pageNumber="581">Surazomus brus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E5792891688617B3AD07BF5C7F8AD" box="[381,448,1826,1845]" pageId="2" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B94DEAE9891688617B81D07BF5EAF8AD" bold="true" box="[454,493,1826,1845]" pageId="2" pageNumber="581">A-E,</emphasis>
Macho holotipo. 
<emphasis id="B94DEAE98916886178CCD07BF6B4F8AD" bold="true" box="[651,691,1826,1845]" pageId="2" pageNumber="581">A-B,</emphasis>
pedipalpo: 
<emphasis id="B94DEAE989168861795DD07BF72BF8AD" bold="true" box="[794,812,1826,1845]" pageId="2" pageNumber="581">A,</emphasis>
vista externa; 
<emphasis id="B94DEAE98916886179ECD07BF7B9F8AD" bold="true" box="[939,958,1826,1845]" pageId="2" pageNumber="581">B,</emphasis>
espolón patelar; 
<emphasis id="B94DEAE9891688617E14D07BF07CF8AD" bold="true" box="[1107,1147,1826,1845]" pageId="2" pageNumber="581">C-E,</emphasis>
flagelo, vistas lateral 
<emphasis id="B94DEAE9891688617AFCD062F4D9F8D6" bold="true" box="[187,222,1851,1870]" pageId="2" pageNumber="581">(C),</emphasis>
dorsal 
<emphasis id="B94DEAE9891688617B5DD062F53EF8D6" bold="true" box="[282,313,1851,1871]" pageId="2" pageNumber="581">(D)</emphasis>
y ventral (E).
</paragraph>
</caption>
<paragraph id="8B8636FB891788607A19D541F56AFB88" blockId="3.[94,643,536,1040]" pageId="3" pageNumber="582">
<emphasis id="B94DEAE9891788607A19D541F4FEFDB7" box="[94,249,536,559]" italics="true" pageId="3" pageNumber="582">Comparaciones:</emphasis>
Esta especie se parece a 
<emphasis id="B94DEAE989178860784FD541F49DFDC8" italics="true" pageId="3" pageNumber="582">
<taxonomicName id="4C394D7889178860784FD541F491FDC8" baseAuthorityName="Rowland &amp; Reddell" baseAuthorityYear="1979" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="pallipatellatus">S. pallipatellatus</taxonomicName>
,
</emphasis>
sin embargo las hembras de ambos taxones se pueden separar por la forma y longitud relativa de los lóbulos medios de las espermatecas, los cuales tienen aspecto de maza en 
<emphasis id="B94DEAE9891788607B69D5C1F5DDFD28" box="[302,474,664,688]" italics="true" pageId="3" pageNumber="582">
<taxonomicName id="4C394D78891788607B69D5C1F5D1FD28" baseAuthorityName="Rowland &amp; Reddell" baseAuthorityYear="1979" box="[302,470,664,688]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="pallipatellatus">S. pallipatellatus</taxonomicName>
,
</emphasis>
mucho más engrosados distalmente que en 
<taxonomicName id="4C394D78891788607BCFD5E1F5CAFD48" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[392,461,696,720]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="brus" status="sp. nov.">
<emphasis id="B94DEAE9891788607BCFD5E1F5CAFD48" box="[392,461,696,720]" italics="true" pageId="3" pageNumber="582">S. brus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E5792891788607B93D5E1F624FD57" box="[468,547,696,719]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
y en esta última especie de largo relativo mucho mayor que los lóbulos laterales (1,42 en 
<taxonomicName id="4C394D78891788607B2CD5A1F5B6FC88" authorityName="De Armas &amp; Manzanilla &amp; Viquez" authorityYear="2010" box="[363,433,760,784]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="brus" status="sp. nov.">
<emphasis id="B94DEAE9891788607B2CD5A1F5B6FC88" box="[363,433,760,784]" italics="true" pageId="3" pageNumber="582">S. brus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27E5792891788607BFED5A1F60EFC97" box="[441,521,760,783]" pageId="3" pageNumber="582" rank="species">sp. nov.</taxonomicNameLabel>
vs. 1,12 en 
<taxonomicName id="4C394D78891788607A19D441F504FCA8" baseAuthorityName="Rowland &amp; Reddell" baseAuthorityYear="1979" box="[94,259,792,816]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="pallipatellatus">
<emphasis id="B94DEAE9891788607A19D441F504FCA8" box="[94,259,792,816]" italics="true" pageId="3" pageNumber="582">S. pallipatellatus</emphasis>
</taxonomicName>
). El macho de 
<taxonomicName id="4C394D78891788607BE7D441F643FCA8" baseAuthorityName="Rowland &amp; Reddell" baseAuthorityYear="1979" box="[416,580,792,816]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="pallipatellatus">
<emphasis id="B94DEAE9891788607BE7D441F643FCA8" box="[416,580,792,816]" italics="true" pageId="3" pageNumber="582">S. pallipatellatus</emphasis>
</taxonomicName>
posee el flagelo más globoso, con el borde anterior del bulbo arqueado (no anguloso) y presenta las cerdas 
<emphasis id="B94DEAE9891788607827D401F684FCED" box="[608,643,856,885]" italics="true" pageId="3" pageNumber="582">
Dl 
<subScript id="17BD34BE89178860783CD43EF684FCED" attach="right" box="[635,643,871,885]" fontSize="6" pageId="3" pageNumber="582">1</subScript>
</emphasis>
y 
<emphasis id="B94DEAE9891788607A35D421F496FC0D" box="[114,145,888,917]" italics="true" pageId="3" pageNumber="582">
Vl 
<subScript id="17BD34BE891788607ACED4DEF496FC0D" attach="left" box="[137,145,903,917]" fontSize="6" pageId="3" pageNumber="582">1</subScript>
</emphasis>
no modificadas; adicionalmente, la forma de la proyección anterior del trocánter en ambas especies es característica (más aguda en 
<taxonomicName id="4C394D78891788607BC6D4E1F623FC48" baseAuthorityName="Rowland &amp; Reddell" baseAuthorityYear="1979" box="[385,548,952,976]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="pallipatellatus">
<emphasis id="B94DEAE9891788607BC6D4E1F623FC48" box="[385,548,952,976]" italics="true" pageId="3" pageNumber="582">S. pallipatellatus</emphasis>
</taxonomicName>
), al igual que el espolón del fémur, que es mucho más largo y agudo en la nueva especie.
</paragraph>
<paragraph id="8B8636FB8917886078A6D7E1F6E9FC88" blockId="3.[680,1229,184,784]" pageId="3" pageNumber="582">
Según 
<bibRefCitation id="EFA84B0A89178860796DD7E1F054FF48" author="ROWLAND, J. M. &amp; REDDELL, J. R." box="[810,1107,184,208]" pageId="3" pageNumber="582" pagination="89 - 119" refId="ref3808" refString="ROWLAND, J. M. &amp; REDDELL, J. R. 1979. The order Schizomida (Arachnida) in the New World. II. simonis and brasiliensis group (Schizomidae: Schizomus). Journal Arachnology, 7: 89 - 119." type="journal article" year="1979">Rowland &amp; Reddell (1979)</bibRefCitation>
, 
<emphasis id="B94DEAE9891788607E25D7E1F6F2FF68" italics="true" pageId="3" pageNumber="582">
<taxonomicName id="4C394D78891788607E25D7E1F6F7FF68" baseAuthorityName="Rowland &amp; Reddell" baseAuthorityYear="1979" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="pallipatellatus">S. pallipatellatus</taxonomicName>
,
</emphasis>
<taxonomicName id="4C394D78891788607947D781F02DFF68" authority="(Kraus, 1957)" baseAuthorityName="Kraus" baseAuthorityYear="1957" box="[768,1066,216,240]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="macarensis">
<emphasis id="B94DEAE9891788607947D781F78FFF68" box="[768,904,216,240]" italics="true" pageId="3" pageNumber="582">S. macarensis</emphasis>
(Kraus, 1957)
</taxonomicName>
, 
<taxonomicName id="4C394D78891788607E7ED781F73FFE88" authority="(Kraus, 1957)" baseAuthorityName="Kraus" baseAuthorityYear="1957" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="species" species="cumbalensis">
<emphasis id="B94DEAE9891788607E7ED781F0CBFF68" box="[1081,1228,216,240]" italics="true" pageId="3" pageNumber="582">S. cumbalensis</emphasis>
(Kraus, 1957)
</taxonomicName>
(estas dos últimas, de 
<collectingCountry id="F32E766B891788607E59D7A1F080FE88" box="[1054,1159,248,272]" name="Colombia" pageId="3" pageNumber="582">Colombia</collectingCountry>
) y una especie innominada de 
<collectingCountry id="F32E766B8917886079EDD641F007FEA8" box="[938,1024,280,304]" name="Ecuador" pageId="3" pageNumber="582">Ecuador</collectingCountry>
presentan la patela del primer par de patas de color blanquecino, un carácter que también lo comparten las tres especies costarricenses tratadas en este trabajo. Este patrón de colorido es único entre los 
<taxonomicName id="4C394D788917886079FBD6C1F04DFE28" authorityName="Cook" authorityYear="1899" box="[956,1098,408,432]" class="Arachnida" family="Hubbardiidae" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="family">Hubbardiidae</taxonomicName>
de América, por lo que tal vez pudiera representar una sinapomorfía de dicho grupo de especies, así mismo la presencia de un espolón interno en la patela pedipalpar de los machos de las tres especies descritas aquí, y la presencia de áreas blanquecinas en los bordes de la depresión dorsomedial del flagelo masculino pareciera soportar una relación mas estrecha entre estas especies. Un análisis filogenético para hipotetizar las relaciones de grupos dentro de 
<taxonomicName id="4C394D78891788607927D5E1F7CCFD57" authorityName="REDDELL &amp; COKENDOLPHER" authorityYear="1995" box="[864,971,696,719]" class="Arachnida" family="Hubbardiidae" genus="Surazomus" kingdom="Animalia" order="Schizomida" pageId="3" pageNumber="582" phylum="Arthropoda" rank="genus">
<emphasis id="B94DEAE9891788607927D5E1F7CCFD57" box="[864,971,696,719]" italics="true" pageId="3" pageNumber="582">Surazomus</emphasis>
</taxonomicName>
es objeto de un presente estudio no publicado aun (Villarreal 
<emphasis id="B94DEAE9891788607E74D580F06FFD68" box="[1075,1128,728,752]" italics="true" pageId="3" pageNumber="582">et al.</emphasis>
datos no publ.).
</paragraph>
</subSubSection>
</treatment>
</document>