348 lines
52 KiB
XML
348 lines
52 KiB
XML
<document ID-DOI="10.11646/zootaxa.5244.5.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7663766" ID-ZooBank="B55396DA-15DB-4CDF-81FD-78686A0EC3A2" approvalRequired="11" approvalRequired_for_textStreams="11" checkinTime="1677051835230" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Rahayu, Dwi Listyo" docDate="2023" docId="039087CD7E17733B47C6FB53F243FBA7" docLanguage="en" docName="zootaxa.5244.5.2.pdf" docOrigin="Zootaxa 5244 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.5.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocryptocoeloma Ward 1936" docType="treatment" docVersion="2" lastPageNumber="447" masterDocId="FFA9FFB57E0573284751FF9DF768FFDA" masterDocTitle="Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)" masterLastPageNumber="454" masterPageNumber="428" pageNumber="446" updateTime="1677052473057" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ng, Peter K. L.</mods:namePart>
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<mods:affiliation>Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<mods:nameIdentifier type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<mods:namePart>Rahayu, Dwi Listyo</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0003-4342-9045</mods:nameIdentifier>
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<mods:affiliation>Research Center for Marine and Land Bioindustry, National Research and Innovation Agency, Dusun Teluk Kodek, Pemenang, Lombok Utara 83756, NTB, Indonesia. dwilistyo @ yahoo. com; https: // orcid. org / 0000 - 0003 - 4342 - 9045</mods:affiliation>
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<mods:nameIdentifier type="email">dwilistyo@yahoo.com</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-22</mods:number>
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<mods:detail type="volume">
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<mods:number>5244</mods:number>
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<mods:number>5</mods:number>
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<mods:start>428</mods:start>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5244.5.2</mods:url>
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<mods:identifier type="DOI">10.11646/zootaxa.5244.5.2</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7663766</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7663762" ID-Zenodo-Dep="7663762" LSID="urn:lsid:plazi:treatment:039087CD7E17733B47C6FB53F243FBA7" httpUri="http://treatment.plazi.org/id/039087CD7E17733B47C6FB53F243FBA7" lastPageId="19" lastPageNumber="447" pageId="18" pageNumber="446">
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<paragraph blockId="18.[151,568,1230,1257]" box="[151,568,1230,1257]" pageId="18" pageNumber="446">
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<heading bold="true" box="[151,568,1230,1257]" fontSize="11" level="1" pageId="18" pageNumber="446" reason="1">
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<taxonomicName ID-CoL="7PWC6" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[151,568,1230,1257]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[151,568,1230,1257]" pageId="18" pageNumber="446">
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<emphasis bold="true" box="[151,415,1230,1257]" italics="true" pageId="18" pageNumber="446">Pseudocryptocoeloma</emphasis>
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<bibRefCitation author="Ward, M." box="[422,568,1230,1257]" pageId="18" pageNumber="446" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection box="[151,1065,1299,1324]" pageId="18" pageNumber="446" type="reference_group">
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<paragraph blockId="18.[151,1065,1299,1324]" box="[151,1065,1299,1324]" pageId="18" pageNumber="446">
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<treatmentCitationGroup box="[151,1065,1299,1324]" pageId="18" pageNumber="446">
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<treatmentCitation author="Ward, M." box="[151,529,1299,1323]" page="3" pageId="18" pageNumber="446" year="1936">
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<taxonomicName ID-CoL="7PWC6" authority="Ward, 1936: 3" authorityName="Ward" authorityPageNumber="3" authorityYear="1936" box="[151,529,1299,1323]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="genus">
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<emphasis box="[151,376,1299,1323]" italics="true" pageId="18" pageNumber="446">Pseudocryptocoeloma</emphasis>
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<bibRefCitation author="Ward, M." box="[383,529,1299,1323]" pageId="18" pageNumber="446" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936: 3</bibRefCitation>
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</taxonomicName>
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;
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<treatmentCitation author="Ng, P. K. L." box="[540,669,1299,1324]" page="97" pageId="18" pageNumber="446" year="1987">
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<bibRefCitation author="Ng, P. K. L." box="[540,669,1299,1324]" pageId="18" pageNumber="446" pagination="69 - 111" refId="ref16944" refString="Ng, P. K. L. (1987) The Indo-Pacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). Indo-Malayan Zoology, 1987, 4 (1), 69 - 111, pl. 1." type="journal article" year="1987">Ng 1987: 97</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Davie, P. J. F." box="[680,852,1299,1323]" page="426" pageId="18" pageNumber="446" year="2002">
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<bibRefCitation author="Davie, P. J. F." box="[680,852,1299,1323]" pageId="18" pageNumber="446" refId="ref14820" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. Zoological Catalogue of Australia. 19.3 B. CSIRO Publications, Melbourne, 641 pp." type="book" year="2002">Davie 2002: 426</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Ng, P. K. L. & D. Guinot & P. J. F. Davie" box="[862,1061,1299,1324]" page="144" pageId="18" pageNumber="446" year="2008">
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<bibRefCitation author="Ng, P. K. L. & D. Guinot & P. J. F. Davie" box="[862,1061,1299,1324]" pageId="18" pageNumber="446" pagination="1 - 286" refId="ref17397" refString="Ng, P. K. L., D. Guinot & P. J. F. Davie, 2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
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Ng
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<emphasis box="[899,950,1299,1323]" italics="true" pageId="18" pageNumber="446">et al.</emphasis>
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2008: 144
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</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<subSubSection box="[151,1228,1369,1396]" pageId="18" pageNumber="446" type="type_taxon">
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<paragraph blockId="18.[151,1437,1369,2008]" box="[151,1228,1369,1396]" pageId="18" pageNumber="446">
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<emphasis bold="true" box="[151,299,1369,1395]" pageId="18" pageNumber="446">
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<typeStatus box="[151,210,1369,1395]" pageId="18" pageNumber="446">Type</typeStatus>
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species
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</emphasis>
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.
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<taxonomicName authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[313,785,1369,1395]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="species" species="parvus">
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<emphasis box="[313,645,1369,1395]" italics="true" pageId="18" pageNumber="446">Pseudocryptocoeloma parvus</emphasis>
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<bibRefCitation author="Ward, M." box="[652,785,1369,1395]" pageId="18" pageNumber="446" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
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</taxonomicName>
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, by original designation; gender neuter.
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</paragraph>
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<subSubSection pageId="18" pageNumber="446" type="diagnosis">
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<paragraph blockId="18.[151,1437,1369,2008]" pageId="18" pageNumber="446">
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<emphasis bold="true" box="[199,314,1405,1431]" pageId="18" pageNumber="446">Diagnosis</emphasis>
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. Carapace subquadrate, broader than long; regions poorly defined; surface smooth, without granules or obvious rugosities, frontal and anterolateral margins covered with numerous soft setae that obscures margins, rest of carapace glabrous, appearing polished (
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<figureCitation box="[675,790,1477,1503]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="18" pageNumber="446">Figs. 13A</figureCitation>
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,
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<figureCitation box="[802,853,1477,1503]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">14A</figureCitation>
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,
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<figureCitation box="[865,917,1477,1504]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">15A</figureCitation>
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); frontal margin with 2 very low lobes, barely separated by faint concavity; lateral lobe not discernible (
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<figureCitation box="[775,886,1513,1539]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="18" pageNumber="446">Figs. 13A</figureCitation>
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,
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<figureCitation box="[895,946,1513,1539]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">14A</figureCitation>
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,
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<figureCitation box="[955,1007,1513,1540]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">15A</figureCitation>
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); anterolateral margin convex; external orbital tooth very low, wide; first and second anterolateral teeth low, wide (
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<figureCitation box="[991,1105,1549,1575]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="18" pageNumber="446">Figs. 13A</figureCitation>
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,
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<figureCitation box="[1117,1168,1549,1575]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">14A</figureCitation>
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,
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<figureCitation box="[1179,1231,1549,1576]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">15A</figureCitation>
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); posterior margin of epistome with wide median triangular lobe with median fissure, separated from lateral part by short fissure, lateral margin almost straight (
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||
<figureCitation box="[422,538,1621,1647]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">Figs. 14A</figureCitation>
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,
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<figureCitation box="[551,635,1621,1648]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">15B, C</figureCitation>
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). Merus of third maxilliped quadrate, with auriculiform anteroexternal angle, exopod tip almost reaching distal edge of merus (
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||
<figureCitation box="[783,888,1657,1684]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">Fig. 15D</figureCitation>
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). Chelipeds asymmetrical, relatively short, chela high, pubescence only on dorsal margin, outer surface smooth (
|
||
<figureCitation box="[877,1027,1693,1720]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="18" pageNumber="446">Figs. 13A, B</figureCitation>
|
||
,
|
||
<figureCitation box="[1040,1125,1693,1719]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">14A–D</figureCitation>
|
||
).P 2–5 short, stout; surface smooth; margins covered with pubescence, merus without cristae on dorsal margin (
|
||
<figureCitation box="[1088,1201,1729,1755]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="18" pageNumber="446">Figs. 13A</figureCitation>
|
||
,
|
||
<figureCitation box="[1213,1262,1729,1755]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">14E</figureCitation>
|
||
). Sternopleonal cavity reaching imaginary line connecting submedian part of coxae of chelipeds; sternite 8 exposed when pleon closed (
|
||
<figureCitation box="[237,348,1801,1827]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="18" pageNumber="446">Figs. 13B</figureCitation>
|
||
,
|
||
<figureCitation box="[359,403,1801,1827]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">14F</figureCitation>
|
||
,
|
||
<figureCitation box="[413,461,1801,1828]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">15E</figureCitation>
|
||
); pleon triangular, transversely narrow, somite 1 and 3 wide, reaching to P5 coxae (
|
||
<figureCitation captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="18" pageNumber="446">Figs. 14F</figureCitation>
|
||
,
|
||
<figureCitation box="[207,253,1837,1864]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">15F</figureCitation>
|
||
). G1 strongly sinuous, distal part strongly curved (
|
||
<figureCitation box="[821,958,1837,1864]" captionStart="FIGURE 15" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[166,1425,196,1762]" captionTargetId="figure-17@22.[151,1435,181,1776]" captionTargetPageId="22" captionText="FIGURE 15. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia. A, left side of carapace; B, frontal view of carapace showing antennae, antennules, orbit and epistome; C, posterior margin of epistome; D, left third maxilliped; E, anterior thoracic sternum; F, pleonal somites 1–4; G, left G1 (ventral view); H, distal part of left G1 (ventral view); I, distal part of left G1 (dorso-mesial view); K, left G2. Scales: A–F = 1.0 mm; G, K = 0.5 mm; H–J = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7663813" httpUri="https://zenodo.org/record/7663813/files/figure.png" pageId="18" pageNumber="446">Fig. 15G –J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="19" lastPageNumber="447" pageId="18" pageNumber="446" type="discussion">
|
||
<paragraph blockId="18.[151,1437,1369,2008]" lastBlockId="19.[151,1437,151,1149]" lastPageId="19" lastPageNumber="447" pageId="18" pageNumber="446">
|
||
<emphasis bold="true" box="[199,308,1873,1899]" pageId="18" pageNumber="446">Remarks</emphasis>
|
||
.
|
||
<bibRefCitation author="Ward, M." box="[325,501,1873,1899]" pageId="18" pageNumber="446" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward (1936: 3)</bibRefCitation>
|
||
established this genus for one small-sized species,
|
||
<taxonomicName authority="Ward, 1936" authorityName="Ward" authorityYear="1936" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="446" phylum="Arthropoda" rank="species" species="parvus">
|
||
<emphasis box="[1101,1436,1873,1899]" italics="true" pageId="18" pageNumber="446">Pseudocryptocoeloma parvus</emphasis>
|
||
<bibRefCitation author="Ward, M." box="[151,285,1909,1935]" pageId="18" pageNumber="446" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
, on the basis of the following characters: “Carapace three-quarters as long as it is broad. The posterior portions of the carapace are flat, smooth and shining. The epistome is twice as wide as it is long. The anterior margin of the buccal frame is raised and has an obsolete fissure on each side. The merus of the external maxilliped has the anterolateral angle sub-auriculate. The buccal frame is completely filled by the external maxillipeds.” He did not compare it with allied genera except
|
||
<taxonomicName authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[572,892,187,213]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[572,745,187,213]" italics="true" pageId="19" pageNumber="447">Cryptocoeloma</emphasis>
|
||
<bibRefCitation author="Miers, E. J." box="[753,892,187,213]" pageId="19" pageNumber="447" pagination="178 - 322" refId="ref16440" refString="Miers, E. J. (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. British Museum (Natural History), London, pp. 178 - 322, pls. 18 - 32" type="book chapter" year="1884">Miers, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="De Man, 1895" authorityName="De Man" authorityYear="1895" box="[953,1307,186,213]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[953,1132,187,213]" italics="true" pageId="19" pageNumber="447">Heteropilumnus</emphasis>
|
||
De
|
||
<bibRefCitation author="Man, J. G. de" box="[1182,1307,186,213]" pageId="19" pageNumber="447" pagination="485 - 609" refId="ref16137" refString="Man, J. G. de (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 485 - 609, pls. 12 - 14. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">Man, 1895</bibRefCitation>
|
||
</taxonomicName>
|
||
. Compared to
|
||
<taxonomicName authorityName="Miers" authorityYear="1884" box="[181,354,223,249]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[181,354,223,249]" italics="true" pageId="19" pageNumber="447">Cryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
, he noted that his new genus differed in that the eyes are visible in dorsal view (versus hidden below the frontal margins, the anterolateral margins are lined with setae of unequal lengths (versus lined with a fringe of setae with interspersed longer setae) and the exposed dorsal surface of the carapace is smooth and polished (versus surface duller, the regions more developed and sculptured on the branchial regions). From
|
||
<taxonomicName authorityName="De Man" authorityYear="1895" box="[1257,1436,331,357]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1257,1436,331,357]" italics="true" pageId="19" pageNumber="447">Heteropilumnus</emphasis>
|
||
</taxonomicName>
|
||
he commented that
|
||
<taxonomicName authorityName="Ward" authorityYear="1936" box="[383,632,367,393]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[383,632,367,393]" italics="true" pageId="19" pageNumber="447">Pseudocryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
differed in the anterolateral margin being entire, the wider posterior carapace margin and the structure of the chela.
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1437,151,1149]" pageId="19" pageNumber="447">
|
||
Despite
|
||
<bibRefCitation author="Ward, M." box="[296,461,439,465]" pageId="19" pageNumber="447" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward’s (1936)</bibRefCitation>
|
||
choice of the genus name,
|
||
<taxonomicName authorityName="Ward" authorityYear="1936" box="[797,1046,439,465]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[797,1046,439,465]" italics="true" pageId="19" pageNumber="447">Pseudocryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
is actually morphologically very different from
|
||
<taxonomicName authorityName="Miers" authorityYear="1884" box="[314,487,475,501]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[314,487,475,501]" italics="true" pageId="19" pageNumber="447">Cryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus box="[500,550,475,501]" pageId="19" pageNumber="447">type</typeStatus>
|
||
species
|
||
<taxonomicName authority="Rathbun, 1923" authorityName="Rathbun" authorityYear="1923" box="[643,1089,475,501]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="species" species="haswelli">
|
||
<emphasis box="[643,915,475,501]" italics="true" pageId="19" pageNumber="447">Cryptocoeloma haswelli</emphasis>
|
||
<bibRefCitation author="Rathbun, M. J." box="[922,1089,475,501]" pageId="19" pageNumber="447" pagination="95 - 156" refId="ref18036" refString="Rathbun, M. J. (1923) Report on the crabs obtained by the F. I. S. " Endeavour " on the coasts of Queensland, New South Wales, Victoria, South Australia and Tasmania. Report on the Brachyrhyncha, Oxystomata and Dromiacea. Biological Results of the Fishing Experiments carried on by the F. I. S. " Endeavour ", 1909 - 14, 5 (3), 95 - 156, figs. 1 - 3, pls. 16 - 42." type="journal article" year="1923">Rathbun, 1923</bibRefCitation>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation author="Ng, P. K. L. & Holthuis, L. B." box="[1135,1365,475,501]" pageId="19" pageNumber="447" pagination="203" refId="ref17449" refString="Ng, P. K. L. & Holthuis, L. B. (1989) Opinion 1554. Cryptocoeloma Miers, 1884 (Crustacea, Decapoda): Cryptocoeloma haswelli Rathbun, 1923 designated as the type species. Bulletin of Zoological Nomenclature, 46 (3), 203." type="journal article" year="1989">Ng & Holthuis 1989</bibRefCitation>
|
||
; Ng
|
||
<emphasis italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
2022). The differences highlighted by
|
||
<bibRefCitation author="Ward, M." box="[615,756,511,537]" pageId="19" pageNumber="447" pagination="1 - 13" refId="ref19005" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11 (1), 1 - 13, pls. 1 - 3." type="journal article" year="1936">Ward (1936)</bibRefCitation>
|
||
are valid at the genus level; the fringe of setae mentioned by Ward for
|
||
<taxonomicName authorityName="Miers" authorityYear="1884" box="[253,426,547,573]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[253,426,547,573]" italics="true" pageId="19" pageNumber="447">Cryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
is in fact very dense and formed by
|
||
<specimenCount box="[814,919,547,573]" pageId="19" pageNumber="447" type="generic" typeStatus="types">two types</specimenCount>
|
||
of long stiff setae which form a thick brush that completely obscures the frontal and anterolateral margins (cf. Ng 1989; Ng
|
||
<emphasis box="[1006,1062,583,609]" italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
2022). In
|
||
<taxonomicName authorityName="Ward" authorityYear="1936" box="[1180,1429,583,609]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1180,1429,583,609]" italics="true" pageId="19" pageNumber="447">Pseudocryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
, the frontal and anterolateral setae are composed of soft plumose setae, not forming a brush, and the rest of the dorsal surfaces are very smooth and glabrous (
|
||
<figureCitation box="[691,798,655,681]" captionStart="FIGURE 13" captionStartId="20.[151,250,1975,1999]" captionTargetBox="[151,1435,561,1952]" captionTargetId="figure-170@20.[151,1435,561,1952]" captionTargetPageId="20" captionText="FIGURE 13. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, overall dorsal view; B, ventral view of cephalothorax." figureDoi="http://doi.org/10.5281/zenodo.7663807" httpUri="https://zenodo.org/record/7663807/files/figure.png" pageId="19" pageNumber="447">Fig. 13A</figureCitation>
|
||
). The smoothness of the carapace is also unlike that of species of
|
||
<taxonomicName authorityName="De Man" authorityYear="1895" box="[272,451,691,717]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[272,451,691,717]" italics="true" pageId="19" pageNumber="447">Heteropilumnus</emphasis>
|
||
</taxonomicName>
|
||
, which have better defined carapace regions and are not as polished in appearance (cf.
|
||
<bibRefCitation author="Ng, P. K. L. & Tan, L. W. H." box="[151,328,727,753]" pageId="19" pageNumber="447" pagination="13 - 24" refId="ref17710" refString="Ng, P. K. L. & Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng & Tan 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ng, P. K. L. & Davie, P. J. F." box="[339,541,727,753]" pageId="19" pageNumber="447" pagination="517 - 524" refId="ref17332" refString="Ng, P. K. L. & Davie, P. J. F. (1991) The Indo-Pacific Pilumnidae VII. Notes on the taxonomy of Heteropilumnus sasekumari (Serene, 1971) and Cryptolutea Ward, 1936 (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 30 (3), 517 - 524." type="journal article" year="1991">Ng & Davie 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Yeo, D. C. J. & Rahayu, D. L. & Ng, P. K. L." box="[552,725,727,753]" pageId="19" pageNumber="447" pagination="79 - 88" refId="ref19082" refString="Yeo, D. C. J., Rahayu, D. L. & Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles Bulletin of Zoology, Supplement 11, 79 - 88." type="journal article" year="2004">
|
||
Yeo
|
||
<emphasis box="[603,659,727,753]" italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Maenosono, T." box="[736,933,727,753]" pageId="19" pageNumber="447" pagination="39 - 45" refId="ref16044" refString="Maenosono, T. (2016) First record of Pseudolitochira decharmoyi (Bouvier, 1915) (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Okinawa Island-group, Ryukyu Archipelago, Japan. Biological Magazine, Okinawa, 54, 39 - 45." type="journal article" year="2016">Maenosono 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ng, P. K. L. & Lin, C. - W. & Ho, P. - H." box="[944,1108,727,753]" pageId="19" pageNumber="447" pagination="12" refId="ref17562" refString="Ng, P. K. L., Lin, C. - W. & Ho, P. - H. (2018) On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on the genus Heteropilumnus De Man, 1895 (Crustacea: Brachyura). Zoological Studies, 57, 12." type="journal article" year="2018">
|
||
Ng
|
||
<emphasis box="[985,1041,727,753]" italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). Another allied genus whose identity was only recently clarified is
|
||
<taxonomicName authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[575,891,763,789]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[575,754,763,789]" italics="true" pageId="19" pageNumber="447">Pseudolitochira</emphasis>
|
||
<bibRefCitation author="Ward, M." box="[760,891,763,789]" pageId="19" pageNumber="447" pagination="49 - 113" refId="ref19040" refString="Ward, M. (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2), 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus box="[905,955,763,789]" pageId="19" pageNumber="447">type</typeStatus>
|
||
species
|
||
<taxonomicName authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[1050,1424,763,789]" class="Malacostraca" family="Goneplacidae" genus="Carcinoplax" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="species" species="integra">
|
||
<emphasis box="[1050,1277,763,789]" italics="true" pageId="19" pageNumber="447">Carcinoplax integra</emphasis>
|
||
<bibRefCitation author="Miers, E. J." box="[1284,1424,763,789]" pageId="19" pageNumber="447" pagination="178 - 322" refId="ref16440" refString="Miers, E. J. (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. British Museum (Natural History), London, pp. 178 - 322, pls. 18 - 32" type="book chapter" year="1884">Miers, 1884</bibRefCitation>
|
||
</taxonomicName>
|
||
), which superficially resembles
|
||
<taxonomicName authorityName="De Man" authorityYear="1895" box="[500,679,799,825]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[500,679,799,825]" italics="true" pageId="19" pageNumber="447">Heteropilumnus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Ng, P. K. L. & Clark, P. F. & Clark, B. & Kamanli, S. A." box="[696,882,799,825]" pageId="19" pageNumber="447" pagination="377 - 391" refId="ref17254" refString="Ng, P. K. L., Clark, P. F., Clark, B. & Kamanli, S. A. (2021) Pseudolitochira integra (Miers, 1884) (Crustacea: Brachyura: Pilumnidae): redescribed and illustrated from micro-CT scanning the type female. Zootaxa, 4969 (2), 377 - 391. https: // doi. org / 10.11646 / zootaxa. 4969.2.9" type="journal article" year="2021">
|
||
Ng
|
||
<emphasis box="[740,799,799,825]" italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
(2021)
|
||
</bibRefCitation>
|
||
showed that it can be easily distinguished from
|
||
<taxonomicName authorityName="De Man" authorityYear="1895" box="[151,330,835,861]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,330,835,861]" italics="true" pageId="19" pageNumber="447">Heteropilumnus</emphasis>
|
||
</taxonomicName>
|
||
as well as
|
||
<taxonomicName authorityName="Miers" authorityYear="1884" box="[463,636,835,861]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[463,636,835,861]" italics="true" pageId="19" pageNumber="447">Cryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Ward" authorityYear="1936" box="[695,944,835,861]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[695,944,835,861]" italics="true" pageId="19" pageNumber="447">Pseudocryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
by posterior margin of the epistome being entire, without any fissure or cleft separating the median triangular lobe from the rest of the structure (see also
|
||
<bibRefCitation author="Ng, P. K. L. & Clark, P. F." pageId="19" pageNumber="447" pagination="349 - 364" refId="ref17189" refString="Ng, P. K. L. & Clark, P. F. (2022) Descriptions of three new Pseudolitochira Ward, 1942 species (Crustacea: Brachyura: Pilumnidae) from the West Pacific. Journal of Natural History, 56 (5 - 8), 349 - 364. https: // doi. org / 10.1080 / 00222933.2022.2065221" type="journal article" year="2022">Ng & Clark, 2022</bibRefCitation>
|
||
). The chela of
|
||
<taxonomicName authorityName="Ward" authorityYear="1936" box="[477,726,907,933]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[477,726,907,933]" italics="true" pageId="19" pageNumber="447">Pseudocryptocoeloma</emphasis>
|
||
</taxonomicName>
|
||
is distinctive, with the chela high and the outer surfaces smooth even though the specimens are small (
|
||
<figureCitation box="[567,701,943,969]" captionStart="FIGURE 14" captionStartId="21.[151,250,1425,1449]" captionTargetBox="[151,1435,181,1401]" captionTargetId="figure-25@21.[151,1435,181,1401]" captionTargetPageId="21" captionText="FIGURE 14. Pseudocryptocoeloma parvum Ward, 1936, male (8.7 × 6.2 mm) (NHM 1937.7.15.20), Australia.A, frontal view of cephalothorax; B, dorsal view of right cheliped; C, right chela; D, left chela; E, right P5; F, male pleonal somites 1–4." figureDoi="http://doi.org/10.5281/zenodo.7663811" httpUri="https://zenodo.org/record/7663811/files/figure.png" pageId="19" pageNumber="447">Fig. 14C, D</figureCitation>
|
||
). The male chelae of species of
|
||
<taxonomicName authorityName="Tesch" authorityYear="1918" box="[1047,1167,943,969]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1047,1167,943,969]" italics="true" pageId="19" pageNumber="447">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Ward" authorityYear="1942" box="[1218,1397,943,969]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1218,1397,943,969]" italics="true" pageId="19" pageNumber="447">Pseudolitochira</emphasis>
|
||
</taxonomicName>
|
||
are not enlarged, relatively more slender and with the outer surface covered with small granules (cf. present study;
|
||
<bibRefCitation author="Ng, P. K. L. & Clark, P. F." pageId="19" pageNumber="447" pagination="349 - 364" refId="ref17189" refString="Ng, P. K. L. & Clark, P. F. (2022) Descriptions of three new Pseudolitochira Ward, 1942 species (Crustacea: Brachyura: Pilumnidae) from the West Pacific. Journal of Natural History, 56 (5 - 8), 349 - 364. https: // doi. org / 10.1080 / 00222933.2022.2065221" type="journal article" year="2022">Ng & Clark, 2022</bibRefCitation>
|
||
). In
|
||
<taxonomicName authorityName="De Man" authorityYear="1895" box="[366,545,1015,1041]" class="Malacostraca" family="Pilumnidae" genus="Heteropilumnus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[366,545,1015,1041]" italics="true" pageId="19" pageNumber="447">Heteropilumnus</emphasis>
|
||
</taxonomicName>
|
||
, the male chelae of most species are asymmetrical, with the larger chela longer and more slender (cf.
|
||
<bibRefCitation author="Ng, P. K. L. & Tan, L. W. H." box="[396,573,1051,1077]" pageId="19" pageNumber="447" pagination="13 - 24" refId="ref17710" refString="Ng, P. K. L. & Tan, L. W. H. (1988) The identities of Heteropilumnus subinteger (Lanchester, 1900) and Heteropilumnus hirsutior (Lanchester, 1900) stat. nov., with description of a new species, Heteropilumnus holthuisi sp. nov. (Crustacea, Decapoda, Brachyura, Pilumnidae). Crustaceana, 54 (1), 13 - 24. https: // doi. org / 10.1163 / 156854088 X 00582" type="journal article" year="1988">Ng & Tan 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ng, P. K. L. & Davie, P. J. F." box="[585,788,1051,1077]" pageId="19" pageNumber="447" pagination="517 - 524" refId="ref17332" refString="Ng, P. K. L. & Davie, P. J. F. (1991) The Indo-Pacific Pilumnidae VII. Notes on the taxonomy of Heteropilumnus sasekumari (Serene, 1971) and Cryptolutea Ward, 1936 (Crustacea: Decapoda: Brachyura). Memoirs of the Queensland Museum, 30 (3), 517 - 524." type="journal article" year="1991">Ng & Davie 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Yeo, D. C. J. & Rahayu, D. L. & Ng, P. K. L." box="[799,972,1051,1077]" pageId="19" pageNumber="447" pagination="79 - 88" refId="ref19082" refString="Yeo, D. C. J., Rahayu, D. L. & Ng, P. K. L. (2004) Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles Bulletin of Zoology, Supplement 11, 79 - 88." type="journal article" year="2004">
|
||
Yeo
|
||
<emphasis box="[849,906,1051,1077]" italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Maenosono, T." box="[983,1181,1051,1077]" pageId="19" pageNumber="447" pagination="39 - 45" refId="ref16044" refString="Maenosono, T. (2016) First record of Pseudolitochira decharmoyi (Bouvier, 1915) (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Okinawa Island-group, Ryukyu Archipelago, Japan. Biological Magazine, Okinawa, 54, 39 - 45." type="journal article" year="2016">Maenosono 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ng, P. K. L. & Lin, C. - W. & Ho, P. - H." box="[1192,1357,1051,1077]" pageId="19" pageNumber="447" pagination="12" refId="ref17562" refString="Ng, P. K. L., Lin, C. - W. & Ho, P. - H. (2018) On three species of reef-dwelling pilumnid crabs from Taiwan, with notes on the genus Heteropilumnus De Man, 1895 (Crustacea: Brachyura). Zoological Studies, 57, 12." type="journal article" year="2018">
|
||
Ng
|
||
<emphasis box="[1233,1289,1051,1077]" italics="true" pageId="19" pageNumber="447">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1437,151,1149]" pageId="19" pageNumber="447">
|
||
<bibRefCitation author="Edmondson, C. H." box="[199,418,1086,1113]" pageId="19" pageNumber="447" pagination="183 - 243" refId="ref15227" refString="Edmondson, C. H. (1951) Some Central Pacific Crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13), 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson (1951)</bibRefCitation>
|
||
described a second species,
|
||
<taxonomicName authorityName="Edmondson" authorityYear="1951" box="[758,1197,1087,1113]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="species" species="symmetrinudum">
|
||
<emphasis box="[758,1197,1087,1113]" italics="true" pageId="19" pageNumber="447">Pseudocryptocoeloma symmetrinudum</emphasis>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry box="[1280,1359,1087,1113]" name="Samoa" pageId="19" pageNumber="447">Samoa</collectingCountry>
|
||
, but it clearly does not belong to this genus and is here referred to
|
||
<taxonomicName authorityName="Tesch" authorityYear="1918" box="[820,940,1123,1149]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[820,940,1123,1149]" italics="true" pageId="19" pageNumber="447">Lophoplax</emphasis>
|
||
</taxonomicName>
|
||
(see Remarks under that species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |