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<document id="DA618AE2418EC9110029C536E7712A4D" ID-DOI="10.11646/zootaxa.4000.1.1" ID-GBIF-Dataset="507a3784-35f5-46f5-9492-e0884a502f40" ID-ISSN="1175-5326" ID-Zenodo-Dep="289219" ID-ZooBank="00769C76-A2D4-4792-A5DD-5F2B7EA352CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461184674989" checkinUser="plazi" docAuthor="Allegro, Gianni &amp; Giachino, Pier Mauro" docDate="2015" docId="039087C7FFA3FF8BE28C4E8A315B6F82" docLanguage="en" docName="zt04000p048.pdf" docOrigin="Zootaxa 4000 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Blennidus (Agraphoderus) orbicollis Straneo 1993" docType="treatment" docVersion="6" lastPageNumber="14" masterDocId="FFA9FFBFFFAFFF86E21B4B7535296F7A" masterDocTitle="Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="48" masterPageNumber="1" pageNumber="13" updateTime="1698599231025" updateUser="plazi">
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<mods:title id="E70C9A177FBF881495E5DFA8F5A5E54E">Annotated checklist of the Blennidus subgenus Agraphoderus species from Peru with description of B. bombonensis n. sp. and synonymic notes (Coleoptera: Carabidae: Pterostichinae)</mods:title>
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<mods:namePart id="83B01AE40CBDE452C4E3AD9173B0ECA4">Allegro, Gianni</mods:namePart>
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<treatment id="039087C7FFA3FF8BE28C4E8A315B6F82" ID-DOI="http://doi.org/10.5281/zenodo.6096057" ID-GBIF-Taxon="127692760" ID-Zenodo-Dep="6096057" LSID="urn:lsid:plazi:treatment:039087C7FFA3FF8BE28C4E8A315B6F82" httpUri="http://treatment.plazi.org/id/039087C7FFA3FF8BE28C4E8A315B6F82" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="C323655AFFA3FF8AE28C4E8A344C6941" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B8636D1FFA3FF8AE28C4E8A363F6960" blockId="12.[151,790,1535,1595]" box="[151,790,1535,1562]" pageId="12" pageNumber="13">
<heading id="D0CE81BDFFA3FF8AE28C4E8A363F6960" bold="true" box="[151,790,1535,1562]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="4C394D52FFA3FF8AE28C4E8A363F6960" ID-CoL="M55X" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,790,1535,1562]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orbicollis" subGenus="Agraphoderus">
<emphasis id="B94DEAC3FFA3FF8AE28C4E8A363F6960" bold="true" box="[151,790,1535,1562]" pageId="12" pageNumber="13">
<emphasis id="B94DEAC3FFA3FF8AE28C4E8A343A6963" bold="true" box="[151,275,1535,1561]" italics="true" pageId="12" pageNumber="13">Blennidus</emphasis>
(
<emphasis id="B94DEAC3FFA3FF8AE33F4E8A34F96963" bold="true" box="[292,464,1535,1561]" italics="true" pageId="12" pageNumber="13">Agraphoderus</emphasis>
)
<emphasis id="B94DEAC3FFA3FF8AE3FA4E8A37786963" bold="true" box="[481,593,1535,1561]" italics="true" pageId="12" pageNumber="13">orbicollis</emphasis>
(
<bibRefCitation id="EFA84B20FFA3FF8AE07A4D7536276960" author="Straneo" box="[609,782,1536,1562]" pageId="12" pageNumber="13" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8636D1FFA3FF8AE28C4D57344C6941" blockId="12.[151,790,1535,1595]" box="[151,357,1570,1595]" pageId="12" pageNumber="13">
(
<figureCitation id="13022A54FFA3FF8AE2844D5735D56941" box="[159,252,1570,1595]" captionStart="FIGURES 32 35" captionStartId="32.[151,264,1953,1976]" captionTargetBox="[254,1347,215,1919]" captionTargetId="figure@32.[226,1361,193,1932]" captionTargetPageId="32" captionText="FIGURES 32 35. Habitus of Blennidus: nigritulus male HT (32); olivaceus male HT (33); orbicollis male PT (34); pachycerus male HT (35)." httpUri="https://zenodo.org/record/289228/files/figure.png" pageId="12" pageNumber="13">Figs. 34</figureCitation>
,
<figureCitation id="13022A54FFA3FF8AE3134D56340F6941" box="[264,294,1571,1595]" captionStart="FIGURES 66 73" captionStartId="39.[151,264,1916,1939]" captionTargetBox="[161,1422,202,1884]" captionTargetId="figure@39.[151,1436,193,1895]" captionTargetPageId="39" captionText="FIGURES 66 73. Pronotum of Blennidus: longiloba male HT (66); mateui male HT (67); mesotibialis male HT (68); negrei male HT (69); nigritulus male HT (70); olivaceus male HT (71); orbicollis male PT (72); pachycerus male HT (73)." httpUri="https://zenodo.org/record/289234/files/figure.png" pageId="12" pageNumber="13">72</figureCitation>
,
<figureCitation id="13022A54FFA3FF8AE3294D5634746941" box="[306,349,1571,1595]" captionStart="FIGURES 106 111" captionStartId="45.[151,264,1953,1976]" captionTargetBox="[267,1323,222,1909]" captionTargetId="figure@45.[234,1353,193,1932]" captionTargetPageId="45" captionText="FIGURES 106 111. Aedeagus (left lateral and dorsal aspects) of Blennidus: nigritulus HT (106); olivaceus HT (107); orbicollis PT (108); pachycerus HT (109); pascoensis HT (110); phaenogonus LT (111)." httpUri="https://zenodo.org/record/289240/files/figure.png" pageId="12" pageNumber="13">108</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA3FF8AE28C4D1F367469A6" pageId="12" pageNumber="13" type="reference_group">
<paragraph id="8B8636D1FFA3FF8AE28C4D1F376D69FA" blockId="12.[151,861,1642,1756]" box="[151,580,1642,1664]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB2911FFFFA3FF8AE28C4D1F376D69FA" box="[151,580,1642,1664]" pageId="12" pageNumber="13">
<taxonomicName id="4C394D52FFA3FF8AE28C4D1F376D69FA" ID-CoL="48Z76" authority="Straneo, 1993: 392" authorityName="Straneo" authorityPageNumber="392" authorityYear="1993" box="[151,580,1642,1664]" class="Insecta" family="Carabidae" genus="Ogmopleura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA3FF8AE28C4D1F345769FA" box="[151,382,1642,1664]" italics="true" pageId="12" pageNumber="13">Ogmopleura orbicollis</emphasis>
<treatmentCitation id="0A9810C0FFA3FF8AE39F4D1F376D69FA" author="Straneo" box="[388,580,1642,1664]" page="392" pageId="12" pageNumber="13" year="1993">
<bibRefCitation id="EFA84B20FFA3FF8AE39F4D1F373D69FA" author="Straneo" box="[388,532,1642,1664]" pageId="12" pageNumber="13" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
: 392
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</paragraph>
<paragraph id="8B8636D1FFA3FF8AE28C4DFC367469A6" blockId="12.[151,861,1642,1756]" pageId="12" pageNumber="13">
<treatmentCitationGroup id="AB2911FFFFA3FF8AE28C4DFC367469A6" pageId="12" pageNumber="13">
<taxonomicName id="4C394D52FFA3FF8AE28C4DFC372269E5" ID-CoL="M55X" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,523,1673,1695]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA3FF8AE28C4DFC344A69E5" box="[151,355,1673,1695]" italics="true" pageId="12" pageNumber="13">Blennidus orbicollis</emphasis>
(
<bibRefCitation id="EFA84B20FFA3FF8AE36A4DFC372B69E5" author="Straneo" box="[369,514,1673,1695]" pageId="12" pageNumber="13" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA3FF8AE00F4DFF37F069E5" author="Lorenz" box="[532,729,1673,1695]" page="263" pageId="12" pageNumber="13" year="2005">
<bibRefCitation id="EFA84B20FFA3FF8AE00F4DFF378269E5" author="Lorenz" box="[532,683,1673,1695]" pageId="12" pageNumber="13" refString="Lorenz, W. (2005 a) Systematic list of extant ground beetles of the world (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005" yearSuffix="a">Lorenz, 2005a</bibRefCitation>
: 263
</treatmentCitation>
<taxonomicName id="4C394D52FFA3FF8AE28C4DDD372269C4" ID-CoL="M55X" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,523,1704,1726]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA3FF8AE28C4DDD344A69C4" box="[151,355,1704,1726]" italics="true" pageId="12" pageNumber="13">Blennidus orbicollis</emphasis>
(
<bibRefCitation id="EFA84B20FFA3FF8AE36A4DDD372B69C4" author="Straneo" box="[369,514,1704,1726]" pageId="12" pageNumber="13" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA3FF8AE00F4DDC37F269C4" author="Lorenz" box="[532,731,1704,1726]" page="709" pageId="12" pageNumber="13" year="2005">
<bibRefCitation id="EFA84B20FFA3FF8AE00F4DDC378269C4" author="Lorenz" box="[532,683,1704,1726]" pageId="12" pageNumber="13" refString="Lorenz, W. (2005 b) Nomina Carabidarum. A directory of the scientific names of ground beetles (Insecta Coleoptera ' Geadephaga': Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 993 pp." type="book" year="2005" yearSuffix="b">Lorenz, 2005b</bibRefCitation>
: 709
</treatmentCitation>
<taxonomicName id="4C394D52FFA3FF8AE28C4DB3372269A6" ID-CoL="M55X" authority="Straneo, 1993" authorityName="Straneo" authorityYear="1993" box="[151,523,1734,1756]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA3FF8AE28C4DB3344A69A6" box="[151,355,1734,1756]" italics="true" pageId="12" pageNumber="13">Blennidus orbicollis</emphasis>
(
<bibRefCitation id="EFA84B20FFA3FF8AE36A4DB3372B69A6" author="Straneo" box="[369,514,1734,1756]" pageId="12" pageNumber="13" refString="Straneo, S. L. (1993) Nuove specie del genere Ogmopleura Tschitscherine (Coleoptera, Carabidae, Pterostichinae) del Peru e dell'Ecuador e chiave per la loro determinazione. Annali del Museo civico di Storia naturale &quot; G. Doria &quot;, 89, 351 - 399." type="journal article" year="1993">Straneo, 1993</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0A9810C0FFA3FF8AE00F4DB3367469A6" author="Allegro" box="[532,861,1734,1756]" page="170" pageId="12" pageNumber="13" year="2011">
<bibRefCitation id="EFA84B20FFA3FF8AE00F4DB3360469A6" author="Allegro" box="[532,813,1734,1756]" pageId="12" pageNumber="13" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino, 2011b</bibRefCitation>
: 170
</treatmentCitation>
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</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA3FF8AE28C4C7C31D568AC" pageId="12" pageNumber="13" type="materials_examined">
<paragraph id="8B8636D1FFA3FF8AE28C4C7C3467685B" blockId="12.[151,1437,1801,2006]" box="[151,334,1801,1826]" pageId="12" pageNumber="13">
<emphasis id="B94DEAC3FFA3FF8AE28C4C7C343D6858" bold="true" box="[151,276,1801,1826]" pageId="12" pageNumber="13">
<typeStatus id="54828873FFA3FF8AE28C4C7C35FB6858" box="[151,210,1801,1826]" pageId="12" pageNumber="13">Type</typeStatus>
area.
</emphasis>
<collectingCountry id="F32E7641FFA3FF8AE3004C7F3467685B" box="[283,334,1802,1825]" name="Peru" pageId="12" pageNumber="13">Peru</collectingCountry>
</paragraph>
<paragraph id="8B8636D1FFA3FF8AE2DC4C58367F683C" blockId="12.[151,1437,1801,2006]" box="[199,854,1837,1862]" pageId="12" pageNumber="13">
<emphasis id="B94DEAC3FFA3FF8AE2DC4C58349A683C" bold="true" box="[199,435,1837,1862]" pageId="12" pageNumber="13">Examined material.</emphasis>
1 PT ♂ and 1 PT ♀ (CSt at
<collectionCode id="ED28AE14FFA3FF8AE0EB4C5B3662683C" box="[752,843,1838,1862]" pageId="12" pageNumber="13">MSNM</collectionCode>
).
</paragraph>
<paragraph id="8B8636D1FFA3FF8AE2DC4C2431D568AC" blockId="12.[151,1437,1801,2006]" pageId="12" pageNumber="13">
<emphasis id="B94DEAC3FFA3FF8AE2DC4C24342F6810" bold="true" box="[199,262,1873,1898]" pageId="12" pageNumber="13">Note.</emphasis>
This species belongs to the
<taxonomicName id="4C394D52FFA3FF8AE09E4C2736306813" box="[645,793,1873,1897]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA3FF8AE09E4C2736306813" box="[645,793,1873,1897]" italics="true" pageId="12" pageNumber="13">B. orbicollis</emphasis>
</taxonomicName>
species group, which includes three species easily distinguished by the distinctive and very similar morphology of aedeagus (
<bibRefCitation id="EFA84B20FFA3FF8AE1C64C00302568F4" author="Allegro" box="[989,1292,1909,1934]" pageId="12" pageNumber="13" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino 2011b</bibRefCitation>
) as well as a further species (
<taxonomicName id="4C394D52FFA3FF8AE34E4CEF34E468CB" box="[341,461,1946,1969]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA3FF8AE34E4CEF34E468CB" box="[341,461,1946,1969]" italics="true" pageId="12" pageNumber="13">B. crassus</emphasis>
</taxonomicName>
) which is tentatively included in this group based on similar habitus, although its aedeagus is still unknown. The HT ♂ is deposited in Fundación Miguel Lillo, Tucumán (
<collectingCountry id="F32E7641FFA3FF8AE6614CC831D968AC" box="[1146,1264,1981,2006]" name="Argentina" pageId="12" pageNumber="13">Argentina</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C323655AFFA2FF8BE2DC4BE2315B6F82" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8B8636D1FFA2FF8BE2DC4BE2315B6F82" blockId="13.[151,1436,151,248]" pageId="13" pageNumber="14">
<emphasis id="B94DEAC3FFA2FF8BE2DC4BE234496FCA" bold="true" box="[199,352,151,176]" pageId="13" pageNumber="14">Distribution.</emphasis>
Even though the recorded specimens of
<taxonomicName id="4C394D52FFA2FF8BE13F4BEC36876FCA" box="[804,942,152,176]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="orbicollis">
<emphasis id="B94DEAC3FFA2FF8BE13F4BEC36876FCA" box="[804,942,152,176]" italics="true" pageId="13" pageNumber="14">B. orbicollis</emphasis>
</taxonomicName>
do not bear labels with precise locality data, it is very likely that this species, in analogy with the three others included in the same species group (
<taxonomicName id="4C394D52FFA2FF8BE7184BC8305F6FAE" box="[1283,1398,189,212]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="crassus">
<emphasis id="B94DEAC3FFA2FF8BE7184BC8305F6FAE" box="[1283,1398,189,212]" italics="true" pageId="13" pageNumber="14">B. crassus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C394D52FFA2FF8BE79F4BC835F66F8D" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pingui">
<emphasis id="B94DEAC3FFA2FF8BE79F4BC835F66F8D" italics="true" pageId="13" pageNumber="14">B. pingui</emphasis>
</taxonomicName>
s and
<taxonomicName id="4C394D52FFA2FF8BE33B4B9434F96F82" box="[288,464,224,248]" class="Insecta" family="Carabidae" genus="Blennidus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pseudolaevis">
<emphasis id="B94DEAC3FFA2FF8BE33B4B9434F96F82" box="[288,464,224,248]" italics="true" pageId="13" pageNumber="14">B. pseudolaevis</emphasis>
</taxonomicName>
), is restricted to northern
<collectingCountry id="F32E7641FFA2FF8BE0E94B95360C6F8D" box="[754,805,224,247]" name="Peru" pageId="13" pageNumber="14">Peru</collectingCountry>
(
<bibRefCitation id="EFA84B20FFA2FF8BE12F4BAA314E6F82" author="Allegro" box="[820,1127,223,248]" pageId="13" pageNumber="14" refString="Allegro, G. &amp; Giachino, P. M. (2011 b) Studies on the Blennidus Motschulsky subgen. Agraphoderus Bates from Peru: the orbicollis species group (Coleoptera: Carabidae: Pterostichini). Veroeffentlichungen des Naturkunde Museums Erfurt, 30, 167 - 173." type="journal article" year="2011" yearSuffix="b">Allegro &amp; Giachino 2011b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>