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<document id="011E14A1137ACB32A924652BA0FD974F" ID-DOI="10.12651/JSR.2014.3.2.127" ID-ISSN="2713-8615" ID-Zenodo-Dep="13143732" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722430635402" checkinUser="felipe" docAuthor="Blakemore, Robert J., Lee, Seunghan &amp; Seo, Hong-Yul" docDate="2014" docId="0390636CFFB4FFE0FF3DFD81FC328F4F" docLanguage="en" docName="JSpeciesRes.3.2.127-166.pdf" docOrigin="Journal of Species Research 3 (2)" docSource="http://dx.doi.org/10.12651/jsr.2014.3.2.127" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Drawida keikiensis Kobayashi 1938" docType="treatment" docVersion="2" lastPageNumber="147" masterDocId="FFA91B14FFA0FFF4FFA0FFE5FFA88C73" masterDocTitle="Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia" masterLastPageNumber="166" masterPageNumber="127" pageNumber="147" updateTime="1722431195132" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="824B5024B802B6F2E3C2D92F94341C72">Reports of Drawida (Oligochaeta: Moniligastridae) from far East Asia</mods:title>
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<mods:namePart id="2969C553BB914E2F9A8BB291AF72FF17">Blakemore, Robert J.</mods:namePart>
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<mods:namePart id="6D4ED782E57183EE67F46CFA7C5999CC">Lee, Seunghan</mods:namePart>
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<mods:namePart id="3495A8B2AEDBBABB78DE7F76AE7B9104">Seo, Hong-Yul</mods:namePart>
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<mods:title id="0E7940F96B331248AFAED5D507B0A4B3">Journal of Species Research</mods:title>
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<treatment id="0390636CFFB4FFE0FF3DFD81FC328F4F" LSID="urn:lsid:plazi:treatment:0390636CFFB4FFE0FF3DFD81FC328F4F" httpUri="http://treatment.plazi.org/id/0390636CFFB4FFE0FF3DFD81FC328F4F" lastPageNumber="147" pageId="20" pageNumber="147">
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<paragraph id="8B86D27AFFB4FFE0FF3DFD81FDF18E0F" blockId="20.[157,601,612,636]" box="[157,601,612,636]" pageId="20" pageNumber="147">
<heading id="D0CE6516FFB4FFE0FF3DFD81FDF18E0F" bold="true" box="[157,601,612,636]" fontSize="36" level="2" pageId="20" pageNumber="147" reason="3">
<emphasis id="B94D0E68FFB4FFE0FF3DFD81FDF18E0F" bold="true" box="[157,601,612,636]" pageId="20" pageNumber="147">
16.
<taxonomicName id="4C39A9F9FFB4FFE0FF65FD81FDF18E0F" authority="Kobayashi, 1938" authorityName="Kobayashi" authorityYear="1938" box="[197,601,612,636]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="20" pageNumber="147" phylum="Annelida" rank="species" species="keikiensis">
<emphasis id="B94D0E68FFB4FFE0FF65FD81FE3F8E0F" bold="true" box="[197,407,612,636]" italics="true" pageId="20" pageNumber="147">Drawida keikiensis</emphasis>
<bibRefCitation id="EFA8AF8BFFB4FFE0FE3DFD81FDF18E0F" author="Kobayashi, S." box="[413,601,612,636]" pageId="20" pageNumber="147" pagination="89 - 170" refId="ref22932" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi, 1938</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32381F1FFB4FFE0FF3DFD60FE458F4F" pageId="20" pageNumber="147" type="reference_group">
<paragraph id="8B86D27AFFB4FFE0FF3DFD60FE4D8EAF" blockId="20.[157,762,645,828]" pageId="20" pageNumber="147">
<treatmentCitationGroup id="AB29F554FFB4FFE0FF3DFD60FE4D8EAF" pageId="20" pageNumber="147">
<treatmentCitation id="0A98F46BFFB4FFE0FF3DFD60FDCC8EEF" author="Kobayashi, S." box="[157,612,645,668]" page="107" pageId="20" pageNumber="147" year="1938">
<taxonomicName id="4C39A9F9FFB4FFE0FF3DFD60FDCC8EEF" authority="Kobayashi, 1938: 107" authorityName="Kobayashi" authorityPageNumber="107" authorityYear="1938" box="[157,612,645,668]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="20" pageNumber="147" phylum="Annelida" rank="species" species="keikiensis">
<emphasis id="B94D0E68FFB4FFE0FF3DFD60FEC48EEF" box="[157,364,645,668]" italics="true" pageId="20" pageNumber="147">Drawida keikiensis</emphasis>
<materialsCitation id="3B51D827FFB4FFE0FED4FD60FDCC8EEF" box="[372,612,645,668]" county="From Suigen" location="Suwon" municipality="Keiki-do" pageId="20" pageNumber="147" specimenCount="1" stateProvince="Seoul">
<bibRefCitation id="EFA8AF8BFFB4FFE0FED4FD60FDCC8EEF" author="Kobayashi, S." box="[372,612,645,668]" pageId="20" pageNumber="147" pagination="89 - 170" refId="ref22932" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi, 1938: 107</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, fig. 4. [
<collectingCounty id="62E7AAF6FFB4FFE0FD1DFD60FF568ECF" pageId="20" pageNumber="147">From Suigen</collectingCounty>
,
<collectingMunicipality id="6BE24800FFB4FFE0FEA8FD40FECC8ECE" box="[264,356,677,701]" pageId="20" pageNumber="147">Keiki-do</collectingMunicipality>
(near
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,
<collectingRegion id="49FD1C98FFB4FFE0FE58FD40FD9F8ECF" box="[504,567,677,700]" country="South Korea" name="Seoul" pageId="20" pageNumber="147">Seoul</collectingRegion>
).
<typeStatus id="54826CD8FFB4FFE0FDE7FD40FD218ECF" box="[583,649,677,700]" pageId="20" pageNumber="147">Types</typeStatus>
, two semimature specimens now lost].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B86D27AFFB4FFE0FF3DFD00FE458F4F" blockId="20.[157,762,645,828]" pageId="20" pageNumber="147">
<treatmentCitationGroup id="AB29F554FFB4FFE0FF3DFD00FE458F4F" pageId="20" pageNumber="147">
<taxonomicName id="4C39A9F9FFB4FFE0FF3DFD00FEC68E8F" authorityName="Kobayashi" authorityYear="1938" box="[157,366,741,764]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="20" pageNumber="147" phylum="Annelida" rank="species" species="keikeiensis">
<emphasis id="B94D0E68FFB4FFE0FF3DFD00FEC68E8F" box="[157,366,741,764]" italics="true" pageId="20" pageNumber="147">Drawida keikeiensis</emphasis>
</taxonomicName>
: Kobayashi, 1941: 263;
<treatmentCitation id="0A98F46BFFB4FFE0FDCDFD00FF798F6F" author="Easton, E. G." page="39" pageId="20" pageNumber="147" year="1981">
<bibRefCitation id="EFA8AF8BFFB4FFE0FDCDFD00FF798F6F" author="Easton, E. G." pageId="20" pageNumber="147" pagination="33 - 65" refId="ref21957" refString="Easton, E. G. 1981. Japanese earthworms: a synopsis of the Megadrile species. ‾ Bulletin of the British Museum (Natural History) Zoology 40 (2): 33 - 65 [Available from: http: // www. archive. org / stream / bulletinofbritis 40 zoollond # pa ge / n 43 / mode / 2 up]." type="journal article" year="1981">Easton, 1981: 39</bibRefCitation>
</treatmentCitation>
, tab. 1;
<bibRefCitation id="EFA8AF8BFFB4FFE0FE8EFCE0FE438F6F" author="Blakemore, R. J." box="[302,491,773,796]" pageId="20" pageNumber="147" pagination="241 - 244" refId="ref20589" refString="Blakemore, R. J. 2003. Japanese earthworms (Annelida: Oligochaeta): a review and checklist of species. Organisms, Diversity and Evolution 3 (3): 241 - 244. [Available from: http: // www. urbanfischer. de / journals / ode /; www. sencken berg. de / odes / 03 - 11. htm]." type="journal article" year="2003">Blakemore, 2003</bibRefCitation>
, 2010, 2014; Blakemore &amp; Kupriyanova, 2010: tab. 3.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32381F1FFB4FFE0FF3DFC80FD1A8FCE" pageId="20" pageNumber="147" type="distribution">
<paragraph id="8B86D27AFFB4FFE0FF3DFC80FD1A8FCE" blockId="20.[157,762,869,1183]" pageId="20" pageNumber="147">
<emphasis id="B94D0E68FFB4FFE0FF3DFC80FE878F0E" bold="true" box="[157,303,869,893]" pageId="20" pageNumber="147">Distribution.</emphasis>
<collectingCountry id="F32E92EAFFB4FFE0FE98FC83FED18F0E" box="[312,377,870,893]" name="South Korea" pageId="20" pageNumber="147">Korea</collectingCountry>
and
<collectingCountry id="F32E92EAFFB4FFE0FE12FC83FE478F0E" box="[434,495,870,893]" name="Japan" pageId="20" pageNumber="147">Japan</collectingCountry>
(from Kobayashi, 1941: 263) Honshu in Chubu-chiho, Kinki-chiho, Gugoku-chiho and on Shikoku and Kyushu also, i.e. widespread.
</paragraph>
</subSubSection>
<subSubSection id="C32381F1FFB4FFE0FF3DFC23FC598E28" pageId="20" pageNumber="147" type="description">
<paragraph id="8B86D27AFFB4FFE0FF3DFC23FDA288ED" blockId="20.[157,762,869,1183]" pageId="20" pageNumber="147">
<emphasis id="B94D0E68FFB4FFE0FF3DFC23FE8E8FAD" bold="true" box="[157,294,966,990]" pageId="20" pageNumber="147">Description.</emphasis>
Grey. Size
<quantity id="4CC17F9FFFB4FFE0FE04FC23FDA78FAE" box="[420,527,966,989]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="5.4" metricValueMin="4.0" pageId="20" pageNumber="147" unit="mm" value="47.0" valueMax="54.0" valueMin="40.0">40-54 mm</quantity>
by up to
<quantity id="4CC17F9FFFB4FFE0FDD5FC23FD6B8FAE" box="[629,707,966,989]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="147" unit="mm" value="2.5">2.5 mm</quantity>
with 138-154 segments Prolobous. Setae small and closely paired (ab=cd). Dorsal pores absent. Clitellum begins to appear on 10-13. Nephropores in line with c. Spermathecal pores 7/8 nearer to c-lines. Male pore on blade-shap- ed penes in 10/11 a-c lines. Female pores paired in 11/
<quantity id="4CC17F9FFFB4FFE0FD7FFB82FF1A88ED" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="20" pageNumber="147" unit="in" value="12.0">12 in</quantity>
b-lines. Genital papillae absent.
</paragraph>
<paragraph id="8B86D27AFFB4FFE0FCEDFEE6FC598E28" blockId="20.[822,1428,259,828]" pageId="20" pageNumber="147">
Septa 5/6-8/9 thick; 10/11 and 11/12 combined in 11/ 12. Hearts last in 9. Spermathecal ampulla with convolut- ed duct to middle of atrium in 7/8 with accessory gland nearby. Testis sacs on 9/10 with very short vas deferens twisted and entering ental end of prostate. Prostate small, short but broad and warty on surface; medial to the prostate is large ovoid gland (not shown in Kobayashis fig. 4). Ovarian chamber in 10/11/12 with eggsacs extending to 16 or to 22. Gizzards three in
<specimenCount id="9D3F19F3FFB4FFE0FB3CFDE1FA988E68" box="[1180,1328,516,539]" count="1" pageId="20" pageNumber="147" type="generic">one specimen</specimenCount>
in 13-15 and four in the other in 12-15. Nephridia vesiculate, vestigial in 9 and 11.
</paragraph>
</subSubSection>
<subSubSection id="C32381F1FFB4FFE0FC96FD81FC328F4F" pageId="20" pageNumber="147" type="discussion">
<paragraph id="8B86D27AFFB4FFE0FC96FD81FC328F4F" blockId="20.[822,1428,259,828]" pageId="20" pageNumber="147">
<emphasis id="B94D0E68FFB4FFE0FC96FD81FC0D8E0F" bold="true" box="[822,933,612,636]" pageId="20" pageNumber="147">Remarks.</emphasis>
<bibRefCitation id="EFA8AF8BFFB4FFE0FC0FFD80FB018E0F" author="Kobayashi, S." box="[943,1193,613,636]" pageId="20" pageNumber="147" pagination="89 - 170" refId="ref22932" refString="Kobayashi, S. 1938. Earthworms of Korea I. Science Report of the Tohoku Imperial University 13 (2): 89 - 170." type="journal article" year="1938">Kobayashi (1938: 110)</bibRefCitation>
said his species was similar to
<taxonomicName id="4C39A9F9FFB4FFE0FC09FD60FB0E8EEF" authority="Chen, 1933" authorityName="Chen" authorityYear="1933" box="[937,1190,645,668]" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="20" pageNumber="147" phylum="Annelida" rank="species" species="syringa">
<emphasis id="B94D0E68FFB4FFE0FC09FD60FBB58EEF" box="[937,1053,645,668]" italics="true" pageId="20" pageNumber="147">D. syringa</emphasis>
<bibRefCitation id="EFA8AF8BFFB4FFE0FB88FD60FB0E8EEF" author="Chen, Y." box="[1064,1190,645,668]" pageId="20" pageNumber="147" pagination="177 - 296" refId="ref21788" refString="Chen, Y. 1933. Preliminary survey of the earthworms of the lower Yangtze Valley. Contributions from the Biological Laboratory of the Science Society of China (Zoology) 9: 177 - 296." type="journal article" year="1933">Chen, 1933</bibRefCitation>
</taxonomicName>
and also to
<taxonomicName id="4C39A9F9FFB4FFE0FA95FD60FC628ECF" authority="(Gates, 1935)" baseAuthorityName="Gates" baseAuthorityYear="1935" class="Clitellata" family="Moniligastridae" genus="Drawida" kingdom="Animalia" order="Moniligastrida" pageId="20" pageNumber="147" phylum="Annelida" rank="species" species="cheni">
<emphasis id="B94D0E68FFB4FFE0FA95FD60FA3B8EEF" box="[1333,1427,645,668]" italics="true" pageId="20" pageNumber="147">D. cheni</emphasis>
(
<bibRefCitation id="EFA8AF8BFFB4FFE0FC9EFD40FC6A8ECF" author="Gates, G. E." box="[830,962,677,700]" pageId="20" pageNumber="147" pagination="445 - 458" refId="ref22066" refString="Gates, G. E. 1935. On some Chinese earthworms. Lingnan Science Journal 14: 445 - 458." type="journal article" year="1935">Gates, 1935</bibRefCitation>
)
</taxonomicName>
but differs from the former mainly in the shape of the penis and from the latter by many characters. Its occurrence in
<collectingCountry id="F32E92EAFFB4FFE0FC5EFD00FB948E8F" box="[1022,1084,741,764]" name="Japan" pageId="20" pageNumber="147">Japan</collectingCountry>
has not since been confirmed although this may relate more to lack of study than true situation.
</paragraph>
</subSubSection>
</treatment>
</document>