335 lines
40 KiB
XML
335 lines
40 KiB
XML
<document id="0FA096CBC05B4F542347D906A85926F3" ID-CLB-Dataset="298355" ID-DOI="10.5194/fr-24-19-2021" ID-GBIF-Dataset="cc5ca7c4-8577-4583-b28a-d9b10e9c5c36" ID-ISSN="2193-0074" ID-Zenodo-Dep="10965766" IM.bibliography_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1712928138748" checkinUser="felipe" docAuthor="Alekseev, Vitalii I., Mitchell, Jerit, McKellar, Ryan C., Barbi, Mauricio, Larsson, Hans C. E. & Bukejs, Andris" docDate="2021" docId="03901871FFDEFFC365AEF9EA8B4BBD8F" docLanguage="en" docName="FossRec.24.1.19-32.pdf" docOrigin="Fossil Record 24 (1)" docSource="http://dx.doi.org/10.5194/fr-24-19-2021" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Chelonarium dingansich Alekseev and Bukejs 2021, sp. nov." docType="treatment" docVersion="6" lastPageNumber="29" masterDocId="FFA96009FFDBFFC9651FFFD48A14BB23" masterDocTitle="The first described turtle beetles from Eocene Baltic amber, with notes on fossil Chelonariidae (Coleoptera: Byrrhoidea)" masterLastPageNumber="32" masterPageNumber="19" pageNumber="24" updateTime="1718077761551" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="26B38A513D268258E3A2536FF10570FF">
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<mods:title id="E80CE52A20DB7316166262FA92969968">The first described turtle beetles from Eocene Baltic amber, with notes on fossil Chelonariidae (Coleoptera: Byrrhoidea)</mods:title>
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<mods:name id="1995B1BAEC8ACB7075A767F0C2D0BD49" type="personal">
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<mods:namePart id="76A2A35A95F867C87DB0E64956832575">Alekseev, Vitalii I.</mods:namePart>
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<mods:namePart id="EF874AF7C3EDB27193457FC7ADBA3F0D">Mitchell, Jerit</mods:namePart>
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<mods:roleTerm id="5C9AE182BCFE3F19010ACEC07E98E5E3">Author</mods:roleTerm>
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<mods:namePart id="34B0D602E9073C7B2288324B4A4027CC">McKellar, Ryan C.</mods:namePart>
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<mods:roleTerm id="F86E8F3BF6CCBF5FB1CFD5662B541EA4">Author</mods:roleTerm>
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<mods:namePart id="3ABC56DA065B0878287A51F89F1901E6">Barbi, Mauricio</mods:namePart>
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<mods:namePart id="E85220F552F82A6807333E9A149DC316">Larsson, Hans C. E.</mods:namePart>
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<mods:namePart id="4E89D6973018AD07A26752FFCDC6E483">Bukejs, Andris</mods:namePart>
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<mods:title id="BA46DB911E53E416E4E9894F7E874FAA">Fossil Record</mods:title>
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<mods:part id="D3D705DA31DB256BD9025E3F8788E17A">
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<mods:date id="4CE4C140B1FFFE512F95CD46B5429F34">2021</mods:date>
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<mods:number id="1E9FACF633D1ABA22F285468D317084D">2021-02-10</mods:number>
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<mods:number id="1280B45F041C693C490E0E6AEAB62B8A">24</mods:number>
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<mods:url id="C9BF9FE14578FB08105B8F7BFEC9912C">http://dx.doi.org/10.5194/fr-24-19-2021</mods:url>
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<treatment id="03901871FFDEFFC365AEF9EA8B4BBD8F" ID-DOI="http://doi.org/10.5281/zenodo.11558751" ID-GBIF-Taxon="232728040" ID-Zenodo-Dep="11558751" LSID="urn:lsid:plazi:treatment:03901871FFDEFFC365AEF9EA8B4BBD8F" httpUri="http://treatment.plazi.org/id/03901871FFDEFFC365AEF9EA8B4BBD8F" lastPageId="10" lastPageNumber="29" pageId="5" pageNumber="24">
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<subSubSection id="C323FAECFFDEFFCC65AEF9EA88AEBDA2" pageId="5" pageNumber="24" type="nomenclature">
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<paragraph id="8B86A967FFDEFFCC65AEF9EA88E3BD76" blockId="5.[177,759,1598,1697]" box="[177,759,1598,1622]" pageId="5" pageNumber="24">
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<heading id="D0CE1E0BFFDEFFCC65AEF9EA88E3BD76" box="[177,759,1598,1622]" fontSize="10" level="2" pageId="5" pageNumber="24" reason="5">
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<taxonomicName id="4C39D2E4FFDEFFCC65AEF9EA888BBD76" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[177,671,1598,1622]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="dingansich" status="sp. nov.">
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<emphasis id="B94D7575FFDEFFCC65AEF9EA8BACBD75" box="[177,440,1598,1622]" italics="true" pageId="5" pageNumber="24">Chelonarium dingansich</emphasis>
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Alekseev and Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="A27EC80EFFDEFFCC67BAF9EA88E3BD76" box="[677,759,1598,1621]" pageId="5" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8B86A967FFDEFFCC65AEF9BE88AEBDA2" blockId="5.[177,759,1598,1697]" box="[177,698,1642,1665]" pageId="5" pageNumber="24">urn:lsid:zoobank.org:act:CE0A8F66-563F-4995-</paragraph>
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</subSubSection>
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<subSubSection id="C323FAECFFDEFFCC65AEF95E8B2FBDEE" pageId="5" pageNumber="24" type="description">
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<paragraph id="8B86A967FFDEFFCC65AEF95E8BBDBD82" blockId="5.[177,759,1598,1697]" box="[177,425,1674,1697]" pageId="5" pageNumber="24">8AD4-F7846E1FE27D</paragraph>
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<paragraph id="8B86A967FFDEFFCC65AEF9628B2FBDEE" blockId="5.[177,315,1718,1741]" box="[177,315,1718,1741]" pageId="5" pageNumber="24">
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<figureCitation id="1302B5E2FFDEFFCC65AEF9628B02BDEE" box="[177,278,1718,1741]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStartId-0="7.[124,191,1532,1554]" captionStartId-1="8.[124,191,1527,1549]" captionTargetBox-0="[294,1292,177,1501]" captionTargetBox-1="[294,1292,177,1496]" captionTargetId-0="figure-15@7.[294,1292,177,1501]" captionTargetId-1="figure-15@8.[294,1292,177,1496]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figure 6. Photomicrographs of Chelonarium dingansich Alekseev and Bukejs sp. nov., holotype, 6696 [MAIG], habitus: (a) dorsal view, (b) ventral view and (c) left lateral view. Scale bar = 1.0 mm." captionText-1="Figure 7. X-ray micro-CT renderings of Chelonarium dingansich Alekseev and Bukejs sp. nov., holotype, 6696 [MAIG], habitus: (a) dorsal view, (b) ventral view and (c) right lateral view. Scale bar = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10965778" figureDoi-1="http://doi.org/10.5281/zenodo.10965782" httpUri-0="https://zenodo.org/record/10965778/files/figure.png" httpUri-1="https://zenodo.org/record/10965782/files/figure.png" pageId="5" pageNumber="24">Figs. 6–7</figureCitation>
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,
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<figureCitation id="1302B5E2FFDEFFCC643EF9628B2FBDEE" box="[289,315,1718,1741]" captionStart="Figure 8" captionStartId="9.[124,191,1243,1265]" captionTargetBox="[302,1285,177,1212]" captionTargetId="figure-15@9.[302,1285,177,1212]" captionTargetPageId="9" captionText="Figure 8. Elytral vestiture and setation shape in fossil Chelonarium: (a) Ch. dingansich Alekseev and Bukejs sp. nov., right elytron, and (b) Ch. andabata Alekseev and Bukejs sp. nov., left elytron. Scale bars = 0.5 mm. Abbreviation: pa – patches of paler and denser setae." figureDoi="http://doi.org/10.5281/zenodo.10965784" httpUri="https://zenodo.org/record/10965784/files/figure.png" pageId="5" pageNumber="24">8a</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C323FAECFFDEFFCC659CF8DA8F92BDF6" pageId="5" pageNumber="24" type="etymology">
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<paragraph id="8B86A967FFDEFFCC659CF8DA8B5CBC05" blockId="5.[131,328,1806,1830]" box="[131,328,1806,1830]" pageId="5" pageNumber="24">
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<emphasis id="B94D7575FFDEFFCC659CF8DA8B5CBC05" bold="true" box="[131,328,1806,1830]" italics="true" pageId="5" pageNumber="24">Derivatio nominis</emphasis>
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</paragraph>
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<paragraph id="8B86A967FFDEFFCC6563F89E8F92BDF6" blockId="5.[124,767,1866,1921]" lastBlockId="5.[819,1462,1598,1749]" pageId="5" pageNumber="24">
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The specific epithet
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<taxonomicName id="4C39D2E4FFDEFFCC6442F89E8BC8BC41" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[349,476,1866,1890]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="dingansich">
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“
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<emphasis id="B94D7575FFDEFFCC6477F89E8BC8BC41" box="[360,476,1866,1890]" italics="true" pageId="5" pageNumber="24">dingansich</emphasis>
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</taxonomicName>
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” is derived from the German word combination “
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<emphasis id="B94D7575FFDEFFCC649BF8BD881FBCA2" box="[388,523,1897,1921]" italics="true" pageId="5" pageNumber="24">Ding an sich</emphasis>
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” (in English: “thing in itself”), a well-known philosophic concept introduced by Immanuel Kant. The species name is dedicated to this famous Königsberg (now
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<collectingRegion id="49FD6785FFDEFFCC66E6F9AA8E6EBDB6" box="[1017,1146,1662,1685]" country="Russia" name="Kaliningrad" pageId="5" pageNumber="24">Kaliningrad</collectingRegion>
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)-native Prussian philosopher and thinker, in the run-up to the 300-year anniversary of his birth in 2024. The name is used as a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="C323FAECFFDEFFC1662CF8D0884FBDC0" lastPageId="8" lastPageNumber="27" pageId="5" pageNumber="24" type="materials_examined">
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<paragraph id="8B86A967FFDEFFCC662CF8D089C6BC3F" blockId="5.[819,978,1796,1820]" box="[819,978,1796,1820]" pageId="5" pageNumber="24">
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<emphasis id="B94D7575FFDEFFCC662CF8D089C6BC3F" bold="true" box="[819,978,1796,1820]" pageId="5" pageNumber="24">Type material</emphasis>
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</paragraph>
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<paragraph id="8B86A967FFDEFFCE662CF89E8AC3BC4B" blockId="5.[819,1462,1866,1921]" lastBlockId="7.[124,767,1650,1896]" lastPageId="7" lastPageNumber="26" pageId="5" pageNumber="24">
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<materialsCitation id="3B51A33AFFDEFFCC662CF89E8E1DBCA2" collectionCode="MAIG" collectorName="Jonas Damzen" pageId="5" pageNumber="24" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548217C5FFDEFFCC662CF89E898EBC42" box="[819,922,1866,1889]" pageId="5" pageNumber="24" type="holotype">Holotype</typeStatus>
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: collection number “6696” [
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<collectionCode id="ED2831A2FFDEFFCC61DEF89E8F19BC42" box="[1217,1293,1866,1889]" pageId="5" pageNumber="24">MAIG</collectionCode>
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] (ex. coll.
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<collectorName id="26CCCCB1FFDEFFCC6064F89E8998BCA2" pageId="5" pageNumber="24">Jonas Damzen</collectorName>
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JDC 7210)
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</materialsCitation>
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, “
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<typeStatus id="548217C5FFDEFFCC6102F8BE8E95BCA2" box="[1053,1153,1898,1921]" pageId="5" pageNumber="24" type="holotype">Holotype</typeStatus>
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/
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<taxonomicName id="4C39D2E4FFDEFFCC6195F8BD8F85BCA2" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[1162,1425,1897,1921]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="dingansich">
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<emphasis id="B94D7575FFDEFFCC6195F8BD8F85BCA2" box="[1162,1425,1897,1921]" italics="true" pageId="5" pageNumber="24">Chelonarium dingansich</emphasis>
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</taxonomicName>
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sp nov./Alekseev and Bukejs des. 2021” [red printed label] [
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<collectionCode id="ED2831A2FFDCFFCE659DF9468AC5BD8A" box="[130,209,1682,1705]" pageId="7" pageNumber="26">MAIG</collectionCode>
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]; adult, sex apparently female. Almost complete beetle (antennomeres 4–11 of both antennae lacking) included in a transparent, yellow amber piece with approximate dimensions of
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<quantity id="4CC10482FFDCFFCE6415F9258B46BC2B" box="[266,338,1777,1800]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="26" unit="mm" value="40.0">40 mm</quantity>
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×
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<quantity id="4CC10482FFDCFFCE646FF9258BACBC2B" box="[368,440,1777,1800]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="26" unit="mm" value="20.0">20 mm</quantity>
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and a maximum thickness of
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<quantity id="4CC10482FFDCFFCE6563F8C58ADCBC0B" box="[124,200,1809,1832]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="26" unit="mm" value="10.0">10 mm</quantity>
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; preserved without supplementary fixation. Syninclusions: four very small stellate trichomes and a few small gas vesicles.
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</paragraph>
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<paragraph id="8B86A967FFDEFFCC60ADF8BE8FA2BCA2" blockId="5.[819,1462,1866,1921]" box="[1458,1462,1898,1921]" pageId="5" pageNumber="24">.</paragraph>
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<caption id="DF46F9EFFFDDFFCF6563FCC789F3B865" ID-DOI="http://doi.org/10.5281/zenodo.10965774" ID-Zenodo-Dep="10965774" httpUri="https://zenodo.org/record/10965774/files/figure.png" pageId="6" pageNumber="25" startId="6.[124,191,787,809]" targetBox="[339,1203,177,756]" targetPageId="6" targetType="figure">
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<paragraph id="8B86A967FFDDFFCF6563FCC789F3B865" blockId="6.[124,1462,787,838]" pageId="6" pageNumber="25">
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<emphasis id="B94D7575FFDDFFCF6563FCC78ACDB80A" bold="true" box="[124,217,787,809]" pageId="6" pageNumber="25">Figure 4.</emphasis>
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Antennae of
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<taxonomicName id="4C39D2E4FFDDFFCF6440FCC08907B80A" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[351,787,788,809]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
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<emphasis id="B94D7575FFDDFFCF6440FCC0882BB80A" box="[351,575,788,809]" italics="true" pageId="6" pageNumber="25">Chelonarium andabata</emphasis>
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Alekseev and Bukejs
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</taxonomicName>
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<taxonomicNameLabel id="A27EC80EFFDDFFCF6604FCC0897DB80A" box="[795,873,788,809]" pageId="6" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype, RSKM_P3000.141 [RSKM]:
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<emphasis id="B94D7575FFDDFFCF61EAFCC78F04B80A" bold="true" box="[1269,1296,787,809]" pageId="6" pageNumber="25">(a)</emphasis>
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X-ray micro-CT rendering and
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<emphasis id="B94D7575FFDDFFCF6419FCE48B37B865" bold="true" box="[262,291,816,838]" pageId="6" pageNumber="25">(b)</emphasis>
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reconstruction. Abbreviations: a1–a11 – antennomeres 1–11 respectively.
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</paragraph>
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</caption>
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<caption id="DF46F9EFFFDDFFCF6563F933898ABC39" ID-DOI="http://doi.org/10.5281/zenodo.10965776" ID-Zenodo-Dep="10965776" httpUri="https://zenodo.org/record/10965776/files/figure.png" pageId="6" pageNumber="25" startId="6.[124,191,1767,1789]" targetBox="[226,1360,885,1736]" targetPageId="6" targetType="figure">
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<paragraph id="8B86A967FFDDFFCF6563F933898ABC39" blockId="6.[124,1462,1767,1818]" pageId="6" pageNumber="25">
|
||
<emphasis id="B94D7575FFDDFFCF6563F9338ACFBDDE" bold="true" box="[124,219,1767,1789]" pageId="6" pageNumber="25">Figure 5.</emphasis>
|
||
X-ray micro-CT renderings of
|
||
<taxonomicName id="4C39D2E4FFDDFFCF6706F93C89C0BDDE" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[537,980,1768,1789]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFDDFFCF6706F93C88EEBDDE" box="[537,762,1768,1789]" italics="true" pageId="6" pageNumber="25">Chelonarium andabata</emphasis>
|
||
Alekseev and Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFDDFFCF66C2F93C8E39BDDE" box="[989,1069,1768,1789]" pageId="6" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, RSKM_P3000.141 [RSKM], aedeagus:
|
||
<emphasis id="B94D7575FFDDFFCF65FFF8D08AE8BC39" bold="true" box="[224,252,1796,1818]" pageId="6" pageNumber="25">(a)</emphasis>
|
||
dorsal view,
|
||
<emphasis id="B94D7575FFDDFFCF6464F8D08B8CBC39" bold="true" box="[379,408,1796,1818]" pageId="6" pageNumber="25">(b)</emphasis>
|
||
lateral view and
|
||
<emphasis id="B94D7575FFDDFFCF6723F8D08842BC39" bold="true" box="[572,598,1796,1818]" pageId="6" pageNumber="25">(c)</emphasis>
|
||
ventral view. Scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF46F9EFFFDCFFCE6563FA2888D1BD0C" ID-DOI="http://doi.org/10.5281/zenodo.10965778" ID-Zenodo-Dep="10965778" httpUri="https://zenodo.org/record/10965778/files/figure.png" pageId="7" pageNumber="26" startId="7.[124,191,1532,1554]" targetBox="[294,1292,177,1501]" targetPageId="7" targetType="figure">
|
||
<paragraph id="8B86A967FFDCFFCE6563FA2888D1BD0C" blockId="7.[124,1462,1532,1584]" pageId="7" pageNumber="26">
|
||
<emphasis id="B94D7575FFDCFFCE6563FA288ACDBD31" bold="true" box="[124,217,1532,1554]" pageId="7" pageNumber="26">Figure 6.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C39D2E4FFDCFFCE64ACFA298961BD31" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[435,885,1533,1554]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="dingansich" status="sp. nov.">
|
||
<emphasis id="B94D7575FFDCFFCE64ACFA2988B6BD31" box="[435,674,1533,1554]" italics="true" pageId="7" pageNumber="26">Chelonarium dingansich</emphasis>
|
||
Alekseev and Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFDCFFCE6662FA2989DFBD31" box="[893,971,1533,1554]" pageId="7" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, 6696 [MAIG], habitus:
|
||
<emphasis id="B94D7575FFDCFFCE6001FA288F2EBD31" bold="true" box="[1310,1338,1532,1554]" pageId="7" pageNumber="26">(a)</emphasis>
|
||
dorsal view,
|
||
<emphasis id="B94D7575FFDCFFCE6563F9CE8A8DBD13" bold="true" box="[124,153,1562,1584]" pageId="7" pageNumber="26">(b)</emphasis>
|
||
ventral view and
|
||
<emphasis id="B94D7575FFDCFFCE645BF9CE8B4ABD13" bold="true" box="[324,350,1562,1584]" pageId="7" pageNumber="26">(c)</emphasis>
|
||
left lateral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B86A967FFDCFFCE662CF9A589D8BDAA" blockId="7.[819,972,1649,1673]" box="[819,972,1649,1673]" pageId="7" pageNumber="26">
|
||
<emphasis id="B94D7575FFDCFFCE662CF9A589D8BDAA" bold="true" box="[819,972,1649,1673]" pageId="7" pageNumber="26">Type stratum</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFDCFFCE662CF97889B4BC00" blockId="7.[819,1462,1708,1827]" pageId="7" pageNumber="26">
|
||
Baltic amber from Eocene amber-bearing Blue Earth layers; a predominantly Bartonian age has been interpreted for the extinct central European resin-producing forests (
|
||
<bibRefCitation id="EFA8D496FFDCFFCE6051F9388981BC00" author="Bukejs, A. & Alekseev, V. I. & Pollock, D. A." pageId="7" pageNumber="26" pagination="261 - 273" refId="ref7344" refString="Bukejs, A., Alekseev, V. I., and Pollock, D. A.: Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location, Zootaxa, 4664, 261 - 273, https: // doi. org / 10.11646 / zootaxa. 4664.2.8, 2019." type="journal article" year="2019">Bukejs et al., 2019</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF46F9EFFFD3FFC16563FA23891FBD09" ID-DOI="http://doi.org/10.5281/zenodo.10965782" ID-Zenodo-Dep="10965782" httpUri="https://zenodo.org/record/10965782/files/figure.png" pageId="8" pageNumber="27" startId="8.[124,191,1527,1549]" targetBox="[294,1292,177,1496]" targetPageId="8" targetType="figure">
|
||
<paragraph id="8B86A967FFD3FFC16563FA23891FBD09" blockId="8.[124,1461,1527,1578]" pageId="8" pageNumber="27">
|
||
<emphasis id="B94D7575FFD3FFC16563FA238AC3BD2E" bold="true" box="[124,215,1527,1549]" pageId="8" pageNumber="27">Figure 7.</emphasis>
|
||
X-ray micro-CT renderings of
|
||
<taxonomicName id="4C39D2E4FFD3FFC1671CFA2389A9BD2E" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[515,957,1527,1549]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="dingansich" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD3FFC1671CFA2388E4BD2F" box="[515,752,1527,1548]" italics="true" pageId="8" pageNumber="27">Chelonarium dingansich</emphasis>
|
||
Alekseev and Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD3FFC166DCFA2C8E1BBD2E" box="[963,1039,1528,1549]" pageId="8" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, 6696 [MAIG], habitus:
|
||
<emphasis id="B94D7575FFD3FFC16045FA238F61BD2E" bold="true" box="[1370,1397,1527,1549]" pageId="8" pageNumber="27">(a)</emphasis>
|
||
dorsal view,
|
||
<emphasis id="B94D7575FFD3FFC165ABF9C08AC5BD09" bold="true" box="[180,209,1556,1578]" pageId="8" pageNumber="27">(b)</emphasis>
|
||
ventral view and
|
||
<emphasis id="B94D7575FFD3FFC16463F9C08B82BD09" bold="true" box="[380,406,1556,1578]" pageId="8" pageNumber="27">(c)</emphasis>
|
||
right lateral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B86A967FFD3FFC16563F9B88B19BDA7" blockId="8.[124,269,1644,1668]" box="[124,269,1644,1668]" pageId="8" pageNumber="27">
|
||
<emphasis id="B94D7575FFD3FFC16563F9B88B19BDA7" bold="true" box="[124,269,1644,1668]" pageId="8" pageNumber="27">Type locality</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD3FFC16563F978884FBDC0" blockId="8.[124,767,1708,1763]" pageId="8" pageNumber="27">
|
||
Yantarny settlement (formerly Palmnicken), Sambian (Samland) Peninsula,
|
||
<collectingRegion id="49FD6785FFD3FFC16431F9188816BDC0" box="[302,514,1740,1763]" country="Russia" name="Kaliningrad" pageId="8" pageNumber="27">Kaliningrad Region</collectingRegion>
|
||
,
|
||
<collectingCountry id="F32EE9F7FFD3FFC16712F9188843BDC0" box="[525,599,1740,1763]" name="Russia" pageId="8" pageNumber="27">Russia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323FAECFFD3FFC36563F8DE8868BF27" lastPageId="10" lastPageNumber="29" pageId="8" pageNumber="27" type="description">
|
||
<paragraph id="8B86A967FFD3FFC16563F8DE8AEBBC01" blockId="8.[124,255,1802,1826]" box="[124,255,1802,1826]" pageId="8" pageNumber="27">
|
||
<emphasis id="B94D7575FFD3FFC16563F8DE8AEBBC01" bold="true" box="[124,255,1802,1826]" pageId="8" pageNumber="27">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD3FFC16563F89E8E04BD87" blockId="8.[124,767,1866,1921]" lastBlockId="8.[819,1463,1645,1892]" pageId="8" pageNumber="27">
|
||
Measurements: body length
|
||
<quantity id="4CC10482FFD3FFC164AEF89E8816BC42" box="[433,514,1866,1889]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="27" unit="mm" value="6.2">6.2 mm</quantity>
|
||
, maximum body width
|
||
<quantity id="4CC10482FFD3FFC16563F8BE8ADBBCA2" box="[124,207,1898,1921]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="27" unit="mm" value="2.7">2.7 mm</quantity>
|
||
; pronotum length
|
||
<quantity id="4CC10482FFD3FFC164ADF8BE8810BCA2" box="[434,516,1898,1921]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="27" unit="mm" value="1.4">1.4 mm</quantity>
|
||
, maximum pronotum width
|
||
<quantity id="4CC10482FFD3FFC16663F9B989DABDA7" box="[892,974,1645,1668]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="27" unit="mm" value="2.3">2.3 mm</quantity>
|
||
; elytra length
|
||
<quantity id="4CC10482FFD3FFC1616BF9B98ED2BDA7" box="[1140,1222,1645,1668]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="8" pageNumber="27" unit="mm" value="4.9">4.9 mm</quantity>
|
||
, maximum combined elytra width
|
||
<quantity id="4CC10482FFD3FFC166A5F9598E18BD87" box="[954,1036,1677,1700]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="27" unit="mm" value="2.7">2.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD3FFC06651F9798B38BEEB" blockId="8.[819,1463,1645,1892]" lastBlockId="9.[124,767,1362,1895]" lastPageId="9" lastPageNumber="28" pageId="8" pageNumber="27">
|
||
Body elongate oval, biconvex; total body length
|
||
<emphasis id="B94D7575FFD3FFC16665F9198992BDC6" bold="true" box="[890,902,1741,1765]" italics="true" pageId="8" pageNumber="27">/</emphasis>
|
||
maximum body width equal to 2.3; integument unicolorous dark brown (as preserved). Pubescence dark, with small patches of denser and paler setae on elytral disc: dorsum covered with short, semierect setae in moderate density; lateral sides of elytra and pronotum with slightly longer, distinctly curved setae; elytral vestiture simple, without any trichome-like structures; venter and legs with dense, recumbent, almost straight setation, setae shorter than setae of dorsum.
|
||
</paragraph>
|
||
<caption id="DF46F9EFFFD2FFC06563FB0F8F9EBE2D" ID-DOI="http://doi.org/10.5281/zenodo.10965784" ID-Zenodo-Dep="10965784" httpUri="https://zenodo.org/record/10965784/files/figure.png" pageId="9" pageNumber="28" startId="9.[124,191,1243,1265]" targetBox="[302,1285,177,1212]" targetPageId="9" targetType="figure">
|
||
<paragraph id="8B86A967FFD2FFC06563FB0F8F9EBE2D" blockId="9.[124,1462,1243,1294]" pageId="9" pageNumber="28">
|
||
<emphasis id="B94D7575FFD2FFC06563FB0F8ACDBFD2" bold="true" box="[124,217,1243,1265]" pageId="9" pageNumber="28">Figure 8.</emphasis>
|
||
Elytral vestiture and setation shape in fossil
|
||
<taxonomicName id="4C39D2E4FFD2FFC0678AFB088907BFD2" authorityName="Fabricius" authorityYear="1801" box="[661,787,1244,1265]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94D7575FFD2FFC0678AFB088907BFD2" box="[661,787,1244,1265]" italics="true" pageId="9" pageNumber="28">Chelonarium</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="B94D7575FFD2FFC0663EFB0F8929BFD2" bold="true" box="[801,829,1243,1265]" pageId="9" pageNumber="28">(a)</emphasis>
|
||
<taxonomicName id="4C39D2E4FFD2FFC0665AFB088EB8BFD2" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[837,1196,1244,1265]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="dingansich" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD2FFC0665AFB0889CCBFD2" box="[837,984,1244,1265]" italics="true" pageId="9" pageNumber="28">Ch. dingansich</emphasis>
|
||
Alekseev and Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD2FFC061ABFB088F16BFD2" box="[1204,1282,1244,1265]" pageId="9" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, right elytron, and
|
||
<emphasis id="B94D7575FFD2FFC06563FB2C8A8DBE2D" bold="true" box="[124,153,1272,1294]" pageId="9" pageNumber="28">(b)</emphasis>
|
||
<taxonomicName id="4C39D2E4FFD2FFC06580FB2D8BE4BE2D" authority="Alekseev and Bukejs" authorityName="Alekseev and Bukejs" authorityYear="2021" box="[159,496,1273,1294]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="28" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD2FFC06580FB2D8B35BE2D" box="[159,289,1273,1294]" italics="true" pageId="9" pageNumber="28">Ch. andabata</emphasis>
|
||
Alekseev and Bukejs
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD2FFC064E9FB2D8857BE2D" box="[502,579,1273,1294]" pageId="9" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, left elytron. Scale bars = 0.5 mm. Abbreviation: pa – patches of paler and denser setae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B86A967FFD2FFC06588FA0588CFBDC4" blockId="9.[124,767,1362,1895]" pageId="9" pageNumber="28">Head declined, not visible from above, slightly convex; densely covered with small punctation; distance between punctures smaller than diameter of one puncture. Clypeus not distinct; frontoclypeal suture absent. Eyes small, entire, almost round, slightly convex, finely facetted, without interfacetal setation; interocular frontal distance equal to about 2× diameter of one eye. Antennal insertions hidden. Antennae rather long; scape hidden; pedicel short, about 0.4× as long as antennomere 3; antennomere 3 longest, flattened.</paragraph>
|
||
<paragraph id="8B86A967FFD2FFC06588F9248E3CBD8B" blockId="9.[124,767,1362,1895]" lastBlockId="9.[819,1462,1362,1895]" pageId="9" pageNumber="28">Pronotum transverse, 1.6× as wide as long, widest at base; densely covered with small punctuation; distance between punctures less than diameter of one puncture. Posterior angles of pronotum acute. Anterior edge of pronotum almost semicircular in dorsal view; posterior edge bisinuate and crenulate. Pronotum margined anteriorly and laterally, with sharp, raised carina dividing pronotum into upper (moderately convex dorsally) and lower (inclined posteriad) parts. Lower pronotal area, between prominent anterior carina and posterior edge of concealed head, wider than protibial width. Pro- and mesosternum with deep, elongate, intercoxal median excavation for reception of basal antennomeres. Hypomeron excavated to receive profemora; mesoand metaventrite with excavations for receiving meso- and metafemora and tibiae.</paragraph>
|
||
<paragraph id="8B86A967FFD2FFC36651F9648B95BA84" blockId="9.[819,1462,1362,1895]" lastBlockId="10.[124,768,176,1028]" lastPageId="10" lastPageNumber="29" pageId="9" pageNumber="28">Scutellum subpentagonal, almost as long as wide, densely covered with semierect, pale setae. Elytra moderately convex, forming elongate oval about 1.9× as long as combined width; slightly wider than pronotal posterior margin, about 3.6× as long as pronotum. Elytral punctation irregular, small, dense and round, with distance between punctures less than diameter of one puncture; interspaces almost flat. Epipleura narrow, reaching abdominal ventrite 1. Metepisternum wide, about 2× as wide as epipleural maximum width; with dense, small punctation. Metaventrite slightly convex laterally, with almost flat disc; densely covered with small punctation; distance between punctures smaller than diameter of one puncture; interspaces slightly convex; discrimen distinct in anterior half of metaventrite.</paragraph>
|
||
<paragraph id="8B86A967FFD1FFC36588FE7B8B4AB866" blockId="10.[124,768,176,1028]" pageId="10" pageNumber="29">Legs rather short, flattened; densely covered with small punctures; distance between punctures smaller than diameter of one puncture. All coxae distinctly separated; procoxa nearly round; mesocoxa oval; metacoxa narrow, strongly transverse, excavate, with short metacoxal plates and with triangular outer edge. Femora and tibiae subequal in length; pro- and mesofemora comparatively wider than metafemur. Tibiae narrower than femora, protibia about 0.6× as wide as profemur, mesotibia about 0.3× as wide mesofemur, metatibia about 0.6× as wide as metafemur; protibia spinose along inner margin. Tarsi 5 segmented, tarsomere 3 apparently with membranous lobe ventrally, tarsomere 4 minute. Tarsal claws with denticle basally.</paragraph>
|
||
<paragraph id="8B86A967FFD1FFC36588FC9A8868BF27" blockId="10.[124,768,176,1028]" pageId="10" pageNumber="29">Abdomen with five visible ventrites; abdominal sutures entire, slightly concave to almost straight; ventrite 5 simple, with widely triangular apical margin; finely and densely punctate; distance between punctures equal to 0.5–1.0× diameter of one puncture. Relative length ratios of abdominal ventrites 1–5 equal to 6: 4: 4: 3.5: 7 (medially).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323FAECFFD1FFC36563FBF388A6BEFB" pageId="10" pageNumber="29" type="diagnosis">
|
||
<paragraph id="8B86A967FFD1FFC36563FBF38B7ABF1C" blockId="10.[124,366,1063,1087]" box="[124,366,1063,1087]" pageId="10" pageNumber="29">
|
||
<emphasis id="B94D7575FFD1FFC36563FBF38B7ABF1C" bold="true" box="[124,366,1063,1087]" pageId="10" pageNumber="29">Differential diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD1FFC36563FBB688A6BEFB" blockId="10.[124,767,1122,1496]" pageId="10" pageNumber="29">
|
||
<taxonomicName id="4C39D2E4FFD1FFC36563FBB68B90BF59" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[124,388,1122,1146]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="dingansich" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC36563FBB68B90BF59" box="[124,388,1122,1146]" italics="true" pageId="10" pageNumber="29">Chelonarium dingansich</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC36493FBB68BCBBF5A" box="[396,479,1122,1145]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C39D2E4FFD1FFC36772FBB688EABF59" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[621,766,1122,1146]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC36772FBB688EABF59" box="[621,766,1122,1146]" italics="true" pageId="10" pageNumber="29">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC36563FB568AC4BFBA" box="[124,208,1154,1177]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in possessing longer and distinctly curved setae on elytral and pronotal lateral sides (setae comparatively shorter and almost straight in
|
||
<taxonomicName id="4C39D2E4FFD1FFC36476FB158BEDBFFA" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[361,505,1217,1241]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC36476FB158BEDBFFA" box="[361,505,1217,1241]" italics="true" pageId="10" pageNumber="29">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC3671EFB168842BFFA" box="[513,598,1218,1241]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); an elytral disc with small patches of paler setae (without small patches of paler setae in
|
||
<taxonomicName id="4C39D2E4FFD1FFC36400FAD58BA1BE3A" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[287,437,1281,1305]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC36400FAD58BA1BE3A" box="[287,437,1281,1305]" italics="true" pageId="10" pageNumber="29">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC364DEFAD68809BE3A" box="[449,541,1282,1305]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); a widely rounded, semicircular apical margin of abdominal ventrite 5 (widely triangular in
|
||
<taxonomicName id="4C39D2E4FFD1FFC36414FA958B8ABE7A" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[267,414,1345,1369]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC36414FA958B8ABE7A" box="[267,414,1345,1369]" italics="true" pageId="10" pageNumber="29">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC364B7FA958815BE7B" box="[424,513,1345,1368]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
); and a distinctly more elongate oval body: less transverse pronotum, 1.6× as wide as long (2.1× as wide as long in
|
||
<taxonomicName id="4C39D2E4FFD1FFC364C5FA55887EBEBA" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[474,618,1409,1433]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC364C5FA55887EBEBA" box="[474,618,1409,1433]" italics="true" pageId="10" pageNumber="29">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC3676EFA5588D2BEBB" box="[625,710,1409,1432]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and more elongate elytra, 1.9× as long as combined width (1.4× as long as combined width in
|
||
<taxonomicName id="4C39D2E4FFD1FFC364A6FA14885EBEFB" authority="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs, 2021" authorityName="Alekseev & Mitchell & McKellar & Barbi & Larsson & Bukejs" authorityYear="2021" box="[441,586,1472,1496]" class="Insecta" family="Chelonariidae" genus="Chelonarium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="29" phylum="Arthropoda" rank="species" species="andabata" status="sp. nov.">
|
||
<emphasis id="B94D7575FFD1FFC364A6FA14885EBEFB" box="[441,586,1472,1496]" italics="true" pageId="10" pageNumber="29">Ch. andabata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27EC80EFFD1FFC3674EFA1588B2BEFB" box="[593,678,1473,1496]" pageId="10" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C323FAECFFD1FFC36563FA2E8B4BBD8F" pageId="10" pageNumber="29" type="discussion">
|
||
<paragraph id="8B86A967FFD1FFC36563FA2E8AF7BD31" blockId="10.[124,227,1530,1554]" box="[124,227,1530,1554]" pageId="10" pageNumber="29">
|
||
<emphasis id="B94D7575FFD1FFC36563FA2E8AF7BD31" bold="true" box="[124,227,1530,1554]" pageId="10" pageNumber="29">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B86A967FFD1FFC36563F9E28B4BBD8F" blockId="10.[124,767,1590,1708]" pageId="10" pageNumber="29">The sex of the examined specimen is determined based on micro-CT results. There is no sclerotized aedeagus-like structure present inside the abdomen; therefore, the specimen appears to be female.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |