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<document id="418B50A5810D2EAE6BA57858B4FA5B15" ID-DOI="10.5281/zenodo.5312492" ID-ISSN="0374-1036" ID-Zenodo-Dep="5312492" ID-ZooBank="456DA4F5-6AB8-4BE5-BBFC-EDCC3A3A692DP" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630119533070" checkinUser="marcus" docAuthor="Hernando, Carles, Ribera, Ignacio &amp; Csic" docDate="2014" docId="03900A19502BFF99FE72FE7B69F58E9A" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.173-189.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Byrrhinus helicophallus Hernando &amp; Ribera &amp; Csic 2014, sp. nov." docType="treatment" docUuid="BD692DE0-A352-41FE-AE28-54EA1460E104" docUuidSource="ZooBank" docVersion="5" lastPageNumber="182" masterDocId="FFA97261502CFF90FFD7FF806B4F8A69" masterDocTitle="The Limnichidae (Coleoptera) of the Arabian Peninsula and the island of Socotra" masterLastPageNumber="189" masterPageNumber="173" pageNumber="180" updateTime="1699159315458" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="EA62E400F2EFAFEDF196D9F8B46BA84A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8BB6ABE031DE21A94CA404A8BA8673D7">
<mods:title id="2747EDE770D65FFEEEB0C22B65DBE89A">The Limnichidae (Coleoptera) of the Arabian Peninsula and the island of Socotra</mods:title>
</mods:titleInfo>
<mods:name id="1ADD16DB4857F478293A47A3EC3E6CEE" type="personal">
<mods:role id="8AA2E678660FB9073C7565B73CF2A953">
<mods:roleTerm id="243EBF46E794AD8C5A415AAF345359C2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F563968DFD484959B8C3B181973B009D">Hernando, Carles</mods:namePart>
<mods:affiliation id="3F00B5B185DC7EC53322C29E2D2BD7A2">P. O. box 118, E- 08911 Badalona, Catalonia, Spain; e-mail: leptotyphlus @ gmail. com</mods:affiliation>
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</mods:role>
<mods:namePart id="F3D55529AD65F69163D8CE9455555AC7">Ribera, Ignacio</mods:namePart>
<mods:affiliation id="204BCC8816937699EA0AF56D683C46ED">Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, E- 08003 Barcelona, Spain; e-mail: Ignacio. Ribera @ ibe. upf-csic. es</mods:affiliation>
</mods:name>
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<mods:namePart id="F73E4032E460BE2BF99CEF25DC38E05A">Csic</mods:namePart>
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<mods:typeOfResource id="4BC5DF77ECB4DDD301AFDC879CD0E985">text</mods:typeOfResource>
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<mods:title id="0FB25088918D7B754BEF62F4C429BBB7">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
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<mods:date id="4CC0DA6122EDD24C36B21374F368869C">2014</mods:date>
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<mods:number id="1545D09CD2D004B32D192BCE60936835">2014-12-30</mods:number>
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<mods:number id="704253B69E950EFFA313EBDEF214C6AB">54</mods:number>
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<treatment id="03900A19502BFF99FE72FE7B69F58E9A" ID-DOI="http://doi.org/10.5281/zenodo.5449316" ID-Zenodo-Dep="5449316" LSID="urn:lsid:zoobank.org:act:BD692DE0-A352-41FE-AE28-54EA1460E104" httpUri="http://treatment.plazi.org/id/03900A19502BFF99FE72FE7B69F58E9A" lastPageId="9" lastPageNumber="182" pageId="7" pageNumber="180">
<subSubSection id="C323E884502BFF97FE72FE7B6842887A" box="[421,781,507,531]" pageId="7" pageNumber="180" type="nomenclature">
<paragraph id="8B86BB0F502BFF97FE72FE7B6842887A" blockId="7.[421,781,507,531]" box="[421,781,507,531]" pageId="7" pageNumber="180">
<heading id="D0CE0C63502BFF97FE72FE7B6842887A" bold="true" box="[421,781,507,531]" centered="true" fontSize="8" level="2" pageId="7" pageNumber="180" reason="3">
<emphasis id="B94D671D502BFF97FE72FE7B6842887A" bold="true" box="[421,781,507,531]" pageId="7" pageNumber="180">
<taxonomicName id="4C39C08C502BFF97FE72FE7B69FE887A" authority="Hernando &amp; Ribera &amp; Csic, 2014" authorityName="Hernando &amp; Ribera &amp; Csic" authorityYear="2014" box="[421,689,507,531]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="180" phylum="Arthropoda" rank="species" species="helicophallus" status="sp. nov.">
<emphasis id="B94D671D502BFF97FE72FE7B69FE887A" bold="true" box="[421,689,507,531]" italics="true" pageId="7" pageNumber="180">Byrrhinus helicophallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EDA66502BFF97FD60FE7C6842887A" box="[695,781,508,531]" pageId="7" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C323E884502BFF97FDCFFD9A69D58844" box="[536,666,538,557]" pageId="7" pageNumber="180" type="description">
<paragraph id="8B86BB0F502BFF97FDCFFD9A69D58844" blockId="7.[536,666,538,557]" box="[536,666,538,557]" pageId="7" pageNumber="180">
(
<figureCitation id="1302A78A502BFF97FDC9FD9A691A8844" box="[542,597,538,557]" captionStart="Figs 15" captionStartId="2.[151,188,1545,1564]" captionTargetBox="[171,1118,231,1467]" captionTargetId="figure-11@2.[145,1126,219,1508]" captionTargetPageId="2" captionText="Figs 15. Habitus of the Limnichidae of Arabian Peninsula and Socotra Islands. 1 Pelochares sinbad sp. nov.; 2 P. sabaeanus sp. nov.; 3 Limnichus arabicus sp. nov.; 4 Byrrhinus socotrensis sp. nov.; 5 B. helicophallus sp. nov. Upper scale bar applied to Figs 13, bottom scale bar to Figs 45." figureDoi="http://doi.org/10.5281/zenodo.5312494" httpUri="https://zenodo.org/record/5312494/files/figure.png" pageId="7" pageNumber="180">Figs 5</figureCitation>
, 1011)
</paragraph>
</subSubSection>
<subSubSection id="C323E884502BFF97FFA6FDC46A1C89A5" pageId="7" pageNumber="180" type="materials_examined">
<paragraph id="8B86BB0F502BFF97FFA6FDC469AF883E" blockId="7.[113,1090,580,972]" box="[113,736,580,599]" pageId="7" pageNumber="180">
<emphasis id="B94D671D502BFF97FFA6FDC46BA5883E" bold="true" box="[113,234,580,599]" pageId="7" pageNumber="180">
<typeStatus id="548205AD502BFF97FFA6FDC46BD1883E" box="[113,158,580,599]" pageId="7" pageNumber="180">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32EFB9F502BFF97FF38FDC46A62883E" box="[239,301,580,599]" name="Yemen" pageId="7" pageNumber="180">Yemen</collectingCountry>
,
<collectingRegion id="49FD75ED502BFF97FEE3FDC46956883E" box="[308,537,580,599]" country="Yemen" name="Al Hudaydah" pageId="7" pageNumber="180">Al Hudaydah Governorate</collectingRegion>
, Jabal Bura (
<figureCitation id="1302A78A502BFF97FD50FDC4699A883E" box="[647,725,580,599]" captionStart="Fig" captionStartId="14.[151,182,932,951]" captionTargetBox="[152,1126,219,913]" captionTargetId="figure-188@14.[152,1127,219,913]" captionTargetPageId="14" captionText="Fig. 15. Representative habitats where specimens of Limnichidae were found. A Oman, wadi 15 km southwest of Sur, type locality of Pelochares sinbad sp. nov.; B Oman, wadi west of Qalhat, also with P. sinbad sp. nov.; C Western Yemen, Jabal Bura, type locality of Pelochares sabaeanus sp. nov. and Byrrhinus helicophallus sp. nov.; D Socotra, wadi Dirhor (Esgego), type locality of B. socotrensis sp. nov.(photos:AB A. Cieslak; CD J. Hájek)" figureDoi="http://doi.org/10.5281/zenodo.5312508" httpUri="https://zenodo.org/record/5312508/files/figure.png" pageId="7" pageNumber="180">Fig. 15C</figureCitation>
).
</paragraph>
<paragraph id="8B86BB0F502BFF97FFA6FDDE687D8915" blockId="7.[113,1090,580,972]" pageId="7" pageNumber="180">
<emphasis id="B94D671D502BFF97FFA6FDDE6BB98818" bold="true" box="[113,246,606,625]" pageId="7" pageNumber="180">Type material.</emphasis>
<materialsCitation id="3B51B152502BFF97FF2DFDDE699488CE" collectingDate="2010-11-04" collectionCode="NMPC" collectorName="J. Bezdek" country="Yemen" elevation="295" location="Jabal Bura valley" longitude="25.2" pageId="7" pageNumber="180" specimenCount="1" stateProvince="Al Hudaydah" typeStatus="holotype">
<typeStatus id="548205AD502BFF97FF2DFDDE6A018819" box="[250,334,606,625]" pageId="7" pageNumber="180" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED2823CA502BFF97FEA4FDDE6AFB8818" box="[371,436,606,625]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="180">NMPC</collectionCode>
),
<collectingCountry id="F32EFB9F502BFF97FE12FDDE69588818" box="[453,535,606,625]" name="Yemen" pageId="7" pageNumber="180">YEMEN</collectingCountry>
,
<collectingRegion id="49FD75ED502BFF97FDC9FDDE69DE8818" box="[542,657,606,625]" country="Yemen" name="Al Hudaydah" pageId="7" pageNumber="180">Al Hudaydah</collectingRegion>
gov. /
<location id="8EE6EDD4502BFF97FD1EFDDE68138818" LSID="urn:lsid:plazi:treatment:03900A19502BFF99FE72FE7B69F58E9A:8EE6EDD4502BFF97FD1EFDDE68138818" box="[713,860,606,625]" country="Yemen" longitude="25.2" name="Jabal Bura valley" pageId="7" pageNumber="180" stateProvince="Al Hudaydah">Jabal Bura valley</location>
forest NP / (stream valley; at light),
<quantity id="4CC116EA502BFF97FF14FDF96A5488E5" box="[195,283,633,652]" metricMagnitude="2" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="7" pageNumber="180" unit="m" value="295.0" valueMax="350.0" valueMin="240.0">
<elevation id="00145C3C502BFF97FF14FDF96A5488E5" box="[195,283,633,652]" metricMagnitude="2" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="7" pageNumber="180" unit="m" value="295.0" valueMax="350.0" valueMin="240.0">240-350m</elevation>
</quantity>
/ 14º52.4-5ʹN, 43º24.6-
<geoCoordinate id="EE0DDDC8502BFF97FE3DFDF9696D88E5" box="[490,546,633,652]" degrees="25.2" direction="east" orientation="longitude" pageId="7" pageNumber="180" precision="5555" value="25.2">25.2ʹE</geoCoordinate>
/
<collectorName id="26CCDED9502BFF97FDE0FDF969C388E5" box="[567,652,633,652]" pageId="7" pageNumber="180">J. Bezděk</collectorName>
leg.
<date id="FF879DCF502BFF97FD6DFDF9684288E5" box="[698,781,633,652]" pageId="7" pageNumber="180" value="2010-11-04">
<collectingDate id="EFC36427502BFF97FD6DFDF9684288E5" box="[698,781,633,652]" pageId="7" pageNumber="180" value="2010-11-04">4.xi.2010</collectingDate>
</date>
[genitalia and abdominal ventrites mounted in DMHF on two transparent cards pinned with the specimen]
</materialsCitation>
.
<materialsCitation id="3B51B152502BFF97FD33FD14695D88A8" collectingDate="2010-11-04" collectionCode="CHBS, IBEB, NHMW, NMPC" collectorName="J. Bezdek" country="Yemen" elevation="295" location="Jabal Bura valley" longitude="25.2" pageId="7" pageNumber="180" specimenCount="32" stateProvince="Al Hudaydah" typeStatus="paratype">
<typeStatus id="548205AD502BFF97FD33FD14687388CC" box="[740,828,660,679]" pageId="7" pageNumber="180" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="9D3F7086502BFF97FC90FD1468F188CE" box="[839,958,660,679]" pageId="7" pageNumber="180" type="generic">26 specimens</specimenCount>
,
<collectionCode id="ED2823CA502BFF97FC1FFD146F4D88CE" box="[968,1026,660,679]" pageId="7" pageNumber="180">CHBS</collectionCode>
,
<collectionCode id="ED2823CA502BFF97FBDCFD146F7188CE" box="[1035,1086,660,679]" pageId="7" pageNumber="180">IBEB</collectionCode>
,
<collectionCode id="ED2823CA502BFF97FFA6FD2E6BF888A8" box="[113,183,686,705]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="180" type="Museum">NHMW</collectionCode>
,
<collectionCode id="ED2823CA502BFF97FF17FD2E6A4E88A8" box="[192,257,686,705]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="7" pageNumber="180">NMPC</collectionCode>
):
<specimenCount id="9D3F7086502BFF97FED9FD2E6A0288A8" box="[270,333,686,705]" pageId="7" pageNumber="180" type="generic">6 spec.</specimenCount>
: same data as holotype
</materialsCitation>
;
<materialsCitation id="3B51B152502BFF97FDCDFD2E688288A8" box="[538,973,686,705]" collectingDate="2010-11-04" collectionCode="NMPC" collectorName="L. Purchart" country="Yemen" elevation="295" location="Jabal Bura valley" longitude="25.2" pageId="7" pageNumber="180" specimenCount="4" stateProvince="Al Hudaydah" typeStatus="paratype">
<specimenCount id="9D3F7086502BFF97FDCDFD2E691788A8" box="[538,600,686,705]" pageId="7" pageNumber="180" type="generic">4 spec.</specimenCount>
, same data as holotype, but
<collectorName id="26CCDED9502BFF97FC91FD2E68EA88A8" box="[838,933,686,705]" pageId="7" pageNumber="180">L. Purchart</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B51B152502BFF97FC02FD2E6AD188B5" collectingDate="2010-11-04" collectionCode="NMPC" collectorName="Jiri Hajek" country="Yemen" elevation="295" location="Jabal Bura valley" longitude="25.2" pageId="7" pageNumber="180" specimenCount="6" stateProvince="Al Hudaydah" typeStatus="paratype">
<specimenCount id="9D3F7086502BFF97FC02FD2E6F5E88A8" box="[981,1041,686,705]" pageId="7" pageNumber="180" type="generic">6 spec.</specimenCount>
same data as holotype, but
<collectorName id="26CCDED9502BFF97FEF3FD496A3888B5" box="[292,375,713,732]" pageId="7" pageNumber="180">Jiří Hájek</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B51B152502BFF97FE71FD496A8F889E" collectingDate="2007-04-09" collectingDateMax="2007-04-11" collectingDateMin="2007-04-09" collectorName="P. Kabatek'" country="Yemen" elevation="431" latitude="14.866667" location="Jabal Bura'" longLatPrecision="1286" longitude="43.4" pageId="7" pageNumber="180" specimenCount="1" stateProvince="Al Hudaydah" typeStatus="paratype">
<specimenCount id="9D3F7086502BFF97FE71FD496AAB88B5" box="[422,484,713,732]" pageId="7" pageNumber="180" type="generic">1 spec.</specimenCount>
: W
<collectingCountry id="F32EFB9F502BFF97FDDDFD49691788B5" box="[522,600,713,732]" name="Yemen" pageId="7" pageNumber="180">YEMEN</collectingCountry>
,
<location id="8EE6EDD4502BFF97FD88FD49698E88B5" LSID="urn:lsid:plazi:treatment:03900A19502BFF99FE72FE7B69F58E9A:8EE6EDD4502BFF97FD88FD49698E88B5" box="[607,705,713,732]" country="Yemen" latitude="14.866667" longLatPrecision="1286" longitude="43.4" name="Jabal Bura'" pageId="7" pageNumber="180" stateProvince="Al Hudaydah">Jabal Bura´</location>
NEE /
<collectingRegion id="49FD75ED502BFF97FD2BFD49683F88B5" box="[764,880,713,732]" country="Yemen" name="Al Hudaydah" pageId="7" pageNumber="180">Al Hudaydah</collectingRegion>
,
<geoCoordinate id="EE0DDDC8502BFF97FCAFFD4968F188B5" box="[888,958,713,732]" degrees="14" direction="north" minutes="52" orientation="latitude" pageId="7" pageNumber="180" precision="925" value="14.866667">N14º52ʹ</geoCoordinate>
/
<geoCoordinate id="EE0DDDC8502BFF97FC1BFD496F5F88B5" box="[972,1040,713,732]" degrees="43" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="180" precision="925" value="43.4">E43º24ʹ</geoCoordinate>
,
<quantity id="4CC116EA502BFF97FBCEFD496BE6889E" metricMagnitude="2" metricUnit="m" metricValue="4.305" metricValueMax="6.0" metricValueMin="2.61" pageId="7" pageNumber="180" unit="m" value="430.5" valueMax="600.0" valueMin="261.0">
<elevation id="00145C3C502BFF97FBCEFD496BE6889E" metricMagnitude="2" metricUnit="m" metricValue="4.305" metricValueMax="6.0" metricValueMin="2.61" pageId="7" pageNumber="180" unit="m" value="430.5" valueMax="600.0" valueMin="261.0">261- 600 m</elevation>
</quantity>
, /
<date id="FF879DCF502BFF97FF6BFD646A7D889E" box="[188,306,740,759]" pageId="7" pageNumber="180" value="2007-04-09" valueMax="2007-04-11" valueMin="2007-04-09">
<collectingDate id="EFC36427502BFF97FF6BFD646A7D889E" box="[188,306,740,759]" pageId="7" pageNumber="180" value="2007-04-09" valueMax="2007-04-11" valueMin="2007-04-09">9.-11.IV.2007</collectingDate>
</date>
, lgt.
<collectorName id="26CCDED9502BFF97FE8BFD646A8F889E" box="[348,448,740,759]" pageId="7" pageNumber="180">P. Kabátek</collectorName>
</materialsCitation>
;
<materialsCitation id="3B51B152502BFF97FE1EFD6468B58978" collectorName="S. Kadlec' &amp; S. Kadlec &amp; National Museum &amp; Prague &amp; Czech Republic'" country="Yemen" elevation="413" latitude="14.866667" location="Jabal Bura'" longLatPrecision="1286" longitude="43.4" pageId="7" pageNumber="180" specimenCount="3" stateProvince="Al Hudaydah" typeStatus="paratype">
<specimenCount id="9D3F7086502BFF97FE1EFD646947889E" box="[457,520,740,759]" pageId="7" pageNumber="180" type="generic">3 spec.</specimenCount>
: W
<collectingCountry id="F32EFB9F502BFF97FDFAFD646934889E" box="[557,635,740,759]" name="Yemen" pageId="7" pageNumber="180">YEMEN</collectingCountry>
,
<location id="8EE6EDD4502BFF97FD54FD6469A8889E" LSID="urn:lsid:plazi:treatment:03900A19502BFF99FE72FE7B69F58E9A:8EE6EDD4502BFF97FD54FD6469A8889E" box="[643,743,740,759]" country="Yemen" latitude="14.866667" longLatPrecision="1286" longitude="43.4" name="Jabal Bura'" pageId="7" pageNumber="180" stateProvince="Al Hudaydah">Jabal Bura</location>
, / NEE
<collectingRegion id="49FD75ED502BFF97FCF0FD6468D4889E" box="[807,923,740,759]" country="Yemen" name="Al Hudaydah" pageId="7" pageNumber="180">Al Hudaydah</collectingRegion>
,
<geoCoordinate id="EE0DDDC8502BFF97FC74FD6468A6889E" box="[931,1001,740,759]" degrees="14" direction="north" minutes="52" orientation="latitude" pageId="7" pageNumber="180" precision="925" value="14.866667">N14º52ʹ</geoCoordinate>
/
<geoCoordinate id="EE0DDDC8502BFF97FC2FFD646F73889E" box="[1016,1084,740,759]" degrees="43" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="180" precision="925" value="43.4">E43º24ʹ</geoCoordinate>
,
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/ 30.X.-1.XI.05, lgt.
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, ex coll.
<collectorName id="26CCDED9502BFF97FDF6FD7E693D8978" box="[545,626,766,785]" pageId="7" pageNumber="180">S. Kadlec</collectorName>
/
<collectorName id="26CCDED9502BFF97FD57FD7E68578978" box="[640,792,766,785]" pageId="7" pageNumber="180">National Museum</collectorName>
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<collectorName id="26CCDED9502BFF97FCF1FD7E682C8978" box="[806,867,766,785]" pageId="7" pageNumber="180">Prague</collectorName>
,
<collectorName id="26CCDED9502BFF97FCBDFD7E68B58978" box="[874,1018,766,785]" pageId="7" pageNumber="180">Czech Republic</collectorName>
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;
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: W
<collectingCountry id="F32EFB9F502BFF97FF45FC996BAF8945" box="[146,224,793,812]" name="Yemen" pageId="7" pageNumber="180">YEMEN</collectingCountry>
,
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NEE /
<collectingRegion id="49FD75ED502BFF97FE5AFC99694C8945" box="[397,515,793,812]" country="Yemen" name="Al Hudaydah" pageId="7" pageNumber="180">Al Hudaydah</collectingRegion>
,
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/
<geoCoordinate id="EE0DDDC8502BFF97FDB2FC9969E78945" box="[613,680,793,812]" degrees="43" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="180" precision="925" value="43.4">E43º24ʹ</geoCoordinate>
,
<quantity id="4CC116EA502BFF97FD63FC99685B8945" box="[692,788,793,812]" metricMagnitude="2" metricUnit="m" metricValue="4.125" metricValueMax="6.0" metricValueMin="2.25" pageId="7" pageNumber="180" unit="m" value="412.5" valueMax="600.0" valueMin="225.0">
<elevation id="00145C3C502BFF97FD63FC99685B8945" box="[692,788,793,812]" metricMagnitude="2" metricUnit="m" metricValue="4.125" metricValueMax="6.0" metricValueMin="2.25" pageId="7" pageNumber="180" unit="m" value="412.5" valueMax="600.0" valueMin="225.0">225-600 m</elevation>
</quantity>
, 30.X.- / 1.XI.05, lgt.
<collectorName id="26CCDED9502BFF97FC00FC996F728945" box="[983,1085,793,812]" pageId="7" pageNumber="180">P. Kabátek</collectorName>
</materialsCitation>
;
<materialsCitation id="3B51B152502BFF97FFA6FCB468618915" collectingDate="2005-10-29" collectorName="S. Kadlec' &amp; S. Kadlec &amp; National Museum &amp; Prague &amp; Czech Republic'" country="Yemen" elevation="1794" latitude="14.666667" location="Dhawran" longLatPrecision="1287" longitude="44.216667" pageId="7" pageNumber="180" specimenCount="1" typeStatus="paratype">
1 J: W
<collectingCountry id="F32EFB9F502BFF97FF6AFCB46A44892E" box="[189,267,820,839]" name="Yemen" pageId="7" pageNumber="180">YEMEN</collectingCountry>
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NW /
<location id="8EE6EDD4502BFF97FE52FCB46A9A892E" LSID="urn:lsid:plazi:treatment:03900A19502BFF99FE72FE7B69F58E9A:8EE6EDD4502BFF97FE52FCB46A9A892E" box="[389,469,820,839]" country="Yemen" latitude="14.666667" longLatPrecision="1287" longitude="44.216667" name="Dhawran" pageId="7" pageNumber="180">Dhawran</location>
,
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<geoCoordinate id="EE0DDDC8502BFF97FDFEFCB46922892E" box="[553,621,820,839]" degrees="44" direction="east" minutes="13" orientation="longitude" pageId="7" pageNumber="180" precision="925" value="44.216667">E44º13ʹ</geoCoordinate>
/
<quantity id="4CC116EA502BFF97FDA9FCB4698E892E" box="[638,705,820,839]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="7" pageNumber="180" unit="m" value="1794.0">
<elevation id="00145C3C502BFF97FDA9FCB4698E892E" box="[638,705,820,839]" metricMagnitude="3" metricUnit="m" metricValue="1.794" pageId="7" pageNumber="180" unit="m" value="1794.0">1794 m</elevation>
</quantity>
,
<date id="FF879DCF502BFF97FD1DFCB46869892E" box="[714,806,820,839]" pageId="7" pageNumber="180" value="2005-10-29">
<collectingDate id="EFC36427502BFF97FD1DFCB46869892E" box="[714,806,820,839]" pageId="7" pageNumber="180" value="2005-10-29">29.X.2005</collectingDate>
</date>
, [
<emphasis id="B94D671D502BFF97FCEAFCB468CC892F" box="[829,899,819,838]" italics="true" pageId="7" pageNumber="180">ex larva</emphasis>
crossed with pencil], / lgt.
<collectorName id="26CCDED9502BFF97FF48FCCE6BB38908" box="[159,252,846,865]" pageId="7" pageNumber="180">S. Kadlec</collectorName>
, ex coll.
<collectorName id="26CCDED9502BFF97FE86FCCE6AEB8908" box="[337,420,846,865]" pageId="7" pageNumber="180">S. Kadlec</collectorName>
/
<collectorName id="26CCDED9502BFF97FE60FCCE691E8908" box="[439,593,846,865]" pageId="7" pageNumber="180">National Museum</collectorName>
/
<collectorName id="26CCDED9502BFF97FDB5FCCE69D08908" box="[610,671,846,865]" pageId="7" pageNumber="180">Prague</collectorName>
,
<collectorName id="26CCDED9502BFF97FD7EFCCE68788908" box="[681,823,846,865]" pageId="7" pageNumber="180">Czech Republic</collectorName>
[immature specimen, genitalia dissected and mounted in DMHF on a transparent card pinned with the specimen]
</materialsCitation>
.
</paragraph>
<paragraph id="8B86BB0F502BFF97FFA6FC046A1C89A5" blockId="7.[113,1090,580,972]" pageId="7" pageNumber="180">
<emphasis id="B94D671D502BFF97FFA6FC046A6A89FE" bold="true" box="[113,293,900,919]" pageId="7" pageNumber="180">Dubious specimens.</emphasis>
<specimenCount id="9D3F7086502BFF97FEFDFC046A1589FF" box="[298,346,893,919]" pageId="7" pageNumber="180" type="female">2 ♀♀</specimenCount>
(
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):
<collectingCountry id="F32EFB9F502BFF97FE6EFC04696E89FE" box="[441,545,900,919]" name="Ethiopia" pageId="7" pageNumber="180">ETHIOPIA</collectingCountry>
,
<collectingRegion id="49FD75ED502BFF97FDFDFC0469C589FE" box="[554,650,900,919]" country="Ethiopia" name="Dire Dawa" pageId="7" pageNumber="180">Dire Dawa</collectingRegion>
, /
<geoCoordinate id="EE0DDDC8502BFF97FD49FC0469BA89FE" box="[670,757,900,919]" degrees="9" direction="north" minutes="39,27" orientation="latitude" pageId="7" pageNumber="180" precision="9" value="9.6545">9º39,27ʹN</geoCoordinate>
,
<geoCoordinate id="EE0DDDC8502BFF97FD28FC04682689FE" box="[767,873,900,919]" degrees="41" direction="east" minutes="55,813" orientation="longitude" pageId="7" pageNumber="180" precision="1" value="41.930218">41º55,813ʹE</geoCoordinate>
, /
<quantity id="4CC116EA502BFF97FCAAFC04688F89FE" box="[893,960,900,919]" metricMagnitude="3" metricUnit="m" metricValue="1.133" pageId="7" pageNumber="180" unit="m" value="1133.0">1133 m</quantity>
,
<date id="FF879DCF502BFF97FC1EFC046F5589FE" box="[969,1050,900,919]" pageId="7" pageNumber="180" value="2011-06-05">5.vi.2011</date>
, / V. Hula &amp; J. Niedobová leg. [genitalia dissected and mounted in DMHF on separate transparent cards pinned with the respective specimens].
</paragraph>
</subSubSection>
<subSubSection id="C323E884502BFF99FFA6FC65691F88FA" lastPageId="9" lastPageNumber="182" pageId="7" pageNumber="180" type="description">
<paragraph id="8B86BB0F502BFF97FFA6FC6569018E55" blockId="7.[113,1090,997,1628]" pageId="7" pageNumber="180">
<emphasis id="B94D671D502BFF97FFA6FC656BB38995" bold="true" box="[113,252,997,1020]" pageId="7" pageNumber="180">Description.</emphasis>
Length
<quantity id="4CC116EA502BFF97FE85FC656A9A8995" box="[338,469,997,1020]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="7" pageNumber="180" unit="mm" value="3.2" valueMax="3.3" valueMin="3.1">3.13.3 mm</quantity>
; maximum width
<quantity id="4CC116EA502BFF97FD45FC65685A8995" box="[658,789,997,1020]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="7" pageNumber="180" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.81.9 mm</quantity>
. Body oval, strongly convex (
<figureCitation id="1302A78A502BFF97FFAEFB856BF58E75" box="[121,186,1029,1052]" captionStart="Figs 15" captionStartId="2.[151,188,1545,1564]" captionTargetBox="[171,1118,231,1467]" captionTargetId="figure-11@2.[145,1126,219,1508]" captionTargetPageId="2" captionText="Figs 15. Habitus of the Limnichidae of Arabian Peninsula and Socotra Islands. 1 Pelochares sinbad sp. nov.; 2 P. sabaeanus sp. nov.; 3 Limnichus arabicus sp. nov.; 4 Byrrhinus socotrensis sp. nov.; 5 B. helicophallus sp. nov. Upper scale bar applied to Figs 13, bottom scale bar to Figs 45." figureDoi="http://doi.org/10.5281/zenodo.5312494" httpUri="https://zenodo.org/record/5312494/files/figure.png" pageId="7" pageNumber="180">Fig. 5</figureCitation>
). Dorsal surface brown to dark brown, covered with yellowish, long, dense erect pubescence. Body appendages paler than body.
</paragraph>
<paragraph id="8B86BB0F502BFF97FF47FBC56F0E8E95" blockId="7.[113,1090,997,1628]" pageId="7" pageNumber="180">Head. Eyes convex, visible from above; upper margin of eyes bordered, border reaching anteriorly insertion of antennae; extended posteriorly to eyes, weakening when reaching posterior margin of head. Surface of head posterior to eyes flat, without depressions or fossae, with erect pubescence and fine and sparse puncturation, denser and coarser on clypeus; surface between punctures smooth and shiny. Antennae strongly pubescent, antennomeres IX symmetrical, antennomere XI asymmetrical, apical four antennomeres forming a loose club.</paragraph>
<paragraph id="8B86BB0F502BFF97FF47FA856F738F15" blockId="7.[113,1090,997,1628]" pageId="7" pageNumber="180">Pronotum transverse (ratio median length / width at base = 0.450.47); anterior margin straight, without crenulations, very finely bordered; posterior margin with a strong double sinuation; lateral margins arched, with prominent borders. Puncturation similar to that on head; pubescence slightly decumbent posteriorly. Hypomeron flat, without depressions or fossae.</paragraph>
<paragraph id="8B86BB0F502BFF97FF47FA056F478FB5" blockId="7.[113,1090,997,1628]" pageId="7" pageNumber="180">Elytra with a weak humeral callus; lateral margins bordered from humerus to apex, finer towards apex; apex acuminate, with a locking device. Membranous wings well developed. Surface of elytra with strong irregular punctures; pubescence slightly bent backwards.</paragraph>
<paragraph id="8B86BB0F502BFF99FF47FA65686F8BFA" blockId="7.[113,1090,997,1628]" lastBlockId="9.[113,1095,220,1267]" lastPageId="9" lastPageNumber="182" pageId="7" pageNumber="180">Ventral surface with strong and dense puncturation, with dense, long recumbent pubescence except on metaventrite, which has only isolated setae. Surface of abdominal ventrites with uniform, fine and very dense punctures. Three first ventrites connate, surface between punctures with polygonal reticle, with median pore; from ventrite IV without polygonal reticle. Inter-metacoxal plate on ventrite I triangular, strongly acuminate; abdominal ventrite I with depressions for reception of metafemora and metatibiae. Ventral surface with pubescence finer than on dorsal surface, recumbent.Apical abdominal ventrite strongly emarginate, with blunt median protuberance; pubescence denser in medial area (in males denser than in females). When seen with transmitted light, posterior margin of apical abdominal ventrite of male with a series of likely glandular tubules, not visible in dry specimens.</paragraph>
<paragraph id="8B86BB0F5025FF99FF47FE1C6AE58B9A" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">Protibia without preapical comb of setae or denticles; apex of protibia with sulcus for reception of tarsi on external surface; apex of meso- and metatibiae with anterior surface flattened, not forming sulcus.</paragraph>
<paragraph id="8B86BB0F5025FF99FF47FE7C6895881A" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">Male genitalia (Fig. 10). Parameres, median lobe of aedeagus and phallobase twisted, strongly asymmetrical and strongly sclerotized. Genital segment asymmetrical, strongly sclerotized; struts asymmetrical, with different length; short strut fused with lamina. Ventrite VIII U-shaped, with narrow apical membranous lamina. Spiculum as in Fig. 10D.</paragraph>
<paragraph id="8B86BB0F5025FF99FF47FDFC691F88FA" blockId="9.[113,1095,220,1267]" box="[144,592,636,659]" pageId="9" pageNumber="182">Female genitalia. Ovipositor as in Fig. 11.</paragraph>
</subSubSection>
<subSubSection id="C323E8845025FF99FFA6FD1C682D893A" pageId="9" pageNumber="182" type="diagnosis">
<paragraph id="8B86BB0F5025FF99FFA6FD1C682D893A" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">
<emphasis id="B94D671D5025FF99FFA6FD1C6A2488DA" bold="true" box="[113,363,668,691]" pageId="9" pageNumber="182">Differential diagnosis.</emphasis>
<taxonomicName id="4C39C08C5025FF99FEA6FD1B693E88DA" authority="Hernando &amp; Ribera &amp; Csic, 2014" authorityName="Hernando &amp; Ribera &amp; Csic" authorityYear="2014" box="[369,625,667,691]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="182" phylum="Arthropoda" rank="species" species="helicophallus" status="sp. nov.">
<emphasis id="B94D671D5025FF99FEA6FD1B693E88DA" box="[369,625,667,691]" italics="true" pageId="9" pageNumber="182">Byrrhinus helicophallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EDA665025FF99FDA0FD1C698588DA" box="[631,714,668,691]" pageId="9" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
does not seem to be closely related to any of the Afrotropical species figured by
<bibRefCitation id="EFA8C6FE5025FF99FD98FD3C69A088BA" author="DELEVE J." box="[591,751,700,723]" pageId="9" pageNumber="182" pagination="212 - 274" refId="ref7744" refString="DELEVE J. 1968: Contribution a l'etude des &quot; Dryopoidea &quot;. Les Limnichinae d'Afrique (Coleoptera Limnichidae). Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 104: 212 - 274." type="journal article" year="1968">DELÈVE (1968)</bibRefCitation>
: the whole aedeagus is strongly asymmetrical, heavily sclerotized, with a very peculiar spiculum. Only one of the known African species has an asymmetrical aedeagus (
<taxonomicName id="4C39C08C5025FF99FDB3FD7C68CF897A" authority="(Deleve, 1968)" baseAuthorityName="Deleve" baseAuthorityYear="1968" box="[612,896,763,787]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="182" phylum="Arthropoda" rank="species" species="bomansi">
<emphasis id="B94D671D5025FF99FDB3FD7C6995897A" box="[612,730,763,787]" italics="true" pageId="9" pageNumber="182">B. bomansi</emphasis>
(
<bibRefCitation id="EFA8C6FE5025FF99FD30FD7C6837897A" author="DELEVE J." box="[743,888,764,787]" pageId="9" pageNumber="182" pagination="212 - 274" refId="ref7744" refString="DELEVE J. 1968: Contribution a l'etude des &quot; Dryopoidea &quot;. Les Limnichinae d'Afrique (Coleoptera Limnichidae). Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 104: 212 - 274." type="journal article" year="1968">Delève, 1968</bibRefCitation>
)
</taxonomicName>
; see
<bibRefCitation id="EFA8C6FE5025FF99FC66FD7C6F72897A" author="DELEVE J." box="[945,1085,764,787]" pageId="9" pageNumber="182" pagination="212 - 274" refId="ref7744" refString="DELEVE J. 1968: Contribution a l'etude des &quot; Dryopoidea &quot;. Les Limnichinae d'Afrique (Coleoptera Limnichidae). Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 104: 212 - 274." type="journal article" year="1968">DELÈVE 1968</bibRefCitation>
: Fig. 43), but nothing comparable to that of
<taxonomicName id="4C39C08C5025FF99FDE9FC9C69A3895A" authority="Hernando &amp; Ribera &amp; Csic, 2014" authorityName="Hernando &amp; Ribera &amp; Csic" authorityYear="2014" box="[574,748,795,819]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="182" phylum="Arthropoda" rank="species" species="helicophallus" status="sp. nov.">
<emphasis id="B94D671D5025FF99FDE9FC9C69A3895A" box="[574,748,795,819]" italics="true" pageId="9" pageNumber="182">B. helicophallus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EDA665025FF99FD25FC9C680A895A" box="[754,837,796,819]" pageId="9" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
which is more likely to be related to some Middle Eastern species (unpublished observations).
</paragraph>
</subSubSection>
<subSubSection id="C323E8845025FF99FFA6FCDC697489BA" pageId="9" pageNumber="182" type="etymology">
<paragraph id="8B86BB0F5025FF99FFA6FCDC6BA789FA" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">
<emphasis id="B94D671D5025FF99FFA6FCDC6BBD891A" bold="true" box="[113,242,860,883]" pageId="9" pageNumber="182">Etymology.</emphasis>
Named in reference to the shape of the aedeagus, twisted like a helix; noun in apposition.
</paragraph>
<paragraph id="8B86BB0F5025FF99FFA6FC1C697489BA" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">
<emphasis id="B94D671D5025FF99FFA6FC1C6ADB89DA" bold="true" box="[113,404,924,947]" pageId="9" pageNumber="182">Collection circumstances.</emphasis>
Most species were collected at light. See notes on
<taxonomicName id="4C39C08C5025FF99FC62FC1C6F0889DA" authority="Hernando &amp; Ribera &amp; Csic, 2014" authorityName="Hernando &amp; Ribera &amp; Csic" authorityYear="2014" box="[949,1095,923,947]" class="Insecta" family="Limnichidae" genus="Pelochares" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="182" phylum="Arthropoda" rank="species" species="sabaeanus" status="sp. nov.">
<emphasis id="B94D671D5025FF99FC62FC1C6F0889DA" box="[949,1095,923,947]" italics="true" pageId="9" pageNumber="182">P. sabaeanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27EDA665025FF99FFA6FC3C6B8B89BA" box="[113,196,956,979]" pageId="9" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
for description of the
<typeStatus id="548205AD5025FF99FE66FC3C6A9089BA" box="[433,479,956,979]" pageId="9" pageNumber="182">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C323E8845025FF99FFA6FC5C6B9C8E7A" pageId="9" pageNumber="182" type="distribution">
<paragraph id="8B86BB0F5025FF99FFA6FC5C6B9C8E7A" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">
<emphasis id="B94D671D5025FF99FFA6FC5C6BB3899A" bold="true" box="[113,252,988,1011]" pageId="9" pageNumber="182">Distribution</emphasis>
(
<figureCitation id="1302A78A5025FF99FEDBFC5C6A12899A" box="[268,349,988,1011]" captionStart="Fig" captionStartId="15.[114,145,1205,1224]" captionTargetBox="[113,1089,219,1187]" captionTargetId="figure-136@15.[113,1088,221,1187]" captionTargetPageId="15" captionText="Fig. 16. Distribution of the known Arabian Limnichidae. Blue circles Pelochares sinbad sp. nov.; green circles P. sabaeanus sp. nov.; red circles Byrrhinus helicophallus sp. nov.; yellow circles B. socotrensis sp. nov.; purple circle Limnichus arabicus sp. nov. With question marks, specimens of uncertain identity from Ethiopia; inset detailed distribution of B. socotrensis sp. nov. in Socotra." figureDoi="http://doi.org/10.5281/zenodo.5312510" httpUri="https://zenodo.org/record/5312510/files/figure.png" pageId="9" pageNumber="182">Fig. 16</figureCitation>
). So far only known from continental
<collectingCountry id="F32EFB9F5025FF99FD2BFC5C6806899A" box="[764,841,988,1011]" name="Yemen" pageId="9" pageNumber="182">Yemen</collectingCountry>
, likely to be present in
<collectingCountry id="F32EFB9F5025FF99FFA6FC7C6B808E7A" box="[113,207,1020,1043]" name="Ethiopia" pageId="9" pageNumber="182">Ethiopia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C323E8845025FF99FFA6FB9C69F58E9A" pageId="9" pageNumber="182" type="discussion">
<paragraph id="8B86BB0F5025FF99FFA6FB9C69F58E9A" blockId="9.[113,1095,220,1267]" pageId="9" pageNumber="182">
<emphasis id="B94D671D5025FF99FFA6FB9C6BAF8E5A" bold="true" box="[113,224,1052,1075]" pageId="9" pageNumber="182">Remarks.</emphasis>
The genitalia of the
<specimenCount id="9D3F70865025FF99FE1CFB9C691F8E5A" box="[459,592,1052,1075]" pageId="9" pageNumber="182" type="female">two females</specimenCount>
from
<collectingCountry id="F32EFB9F5025FF99FD40FB9C69BE8E5A" box="[663,753,1052,1075]" name="Ethiopia" pageId="9" pageNumber="182">Ethiopia</collectingCountry>
appear identical to that of the specimens from
<collectingCountry id="F32EFB9F5025FF99FEC9FBBC6A238E3A" box="[286,364,1084,1107]" name="Yemen" pageId="9" pageNumber="182">Yemen</collectingCountry>
. There are small differences in the external morphology, which could well be due to intraspecific variability, but in the absence of males any identification remains uncertain. In
<bibRefCitation id="EFA8C6FE5025FF99FF2DFBFC6A908EFA" author="SPANGLER P. J. &amp; STAINES C. L. &amp; SPANGLER P. M. &amp; STAINES S. L." box="[250,479,1148,1171]" pageId="9" pageNumber="182" pagination="151 - 165" refId="ref7928" refString="SPANGLER P. J., STAINES C. L., SPANGLER P. M. &amp; STAINES S. L. 2001: A checklist of the Limnichidae and the Lutrochidae (Coleoptera) of the world. Insecta Mundi 15: 151 - 165." type="journal article" year="2001">SPANGLER et al. (2001)</bibRefCitation>
<taxonomicName id="4C39C08C5025FF99FE33FBFC68648EFA" authority="Champion, 1923" authorityName="Champion" authorityYear="1923" box="[484,811,1148,1171]" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="182" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B94D671D5025FF99FE33FBFC69388EFA" box="[484,631,1148,1171]" italics="true" pageId="9" pageNumber="182">B. marginatus</emphasis>
<bibRefCitation id="EFA8C6FE5025FF99FDABFBFC68648EFA" author="CHAMPION G. C." box="[636,811,1148,1171]" pageId="9" pageNumber="182" pagination="269 - 277" refId="ref7713" refString="CHAMPION G. C. 1923: Some Indian Coleoptera (12). Entomologist's Monthly Magazine 59: 219 - 224, 247 - 253, 269 - 277." type="journal article" year="1923">Champion, 1923</bibRefCitation>
</taxonomicName>
is recorded from
<collectingCountry id="F32EFB9F5025FF99FC37FBFC6F728EFA" box="[992,1085,1148,1171]" name="Pakistan" pageId="9" pageNumber="182">Pakistan</collectingCountry>
, but we have not been able to trace the origin of this record and it is likely a mistake.
<taxonomicName id="4C39C08C5025FF99FC01FB1B6BA58EBA" authorityName="Champion" authorityYear="1923" class="Insecta" family="Limnichidae" genus="Byrrhinus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="182" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B94D671D5025FF99FC01FB1B6BA58EBA" italics="true" pageId="9" pageNumber="182">Byrrhinus marginatus</emphasis>
</taxonomicName>
was described from the central Himalayas (Kumaon,
<bibRefCitation id="EFA8C6FE5025FF99FCEFFB3C68AD8EBA" author="CHAMPION G. C." box="[824,994,1212,1235]" pageId="9" pageNumber="182" pagination="269 - 277" refId="ref7713" refString="CHAMPION G. C. 1923: Some Indian Coleoptera (12). Entomologist's Monthly Magazine 59: 219 - 224, 247 - 253, 269 - 277." type="journal article" year="1923">CHAMPION 1923</bibRefCitation>
), and no other records have been published after its description.
</paragraph>
</subSubSection>
</treatment>
</document>