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<document id="BA7E63F1E2DD3E8D11E63AE740E75DC2" ID="10.11646/zootaxa.4246.1.1SLASH10606" ID-CLB-Dataset="36212" ID-DOI="10.5281/zenodo.437611" ID-GBIF-Dataset="3ff74501-6241-43b2-a52f-0e7580bb2f45" ID-ISSN="1175-5326" ID-Zenodo-Dep="437611" ID-ZooBank="D8594E96-D561-4A37-9073-B138EC7E95A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490260213340" checkinUser="plazi" docAuthor="Zaragoza, Juan A." docDate="2017" docId="03889E18FFC8556EFF5C41F6FDD7FBC2" docLanguage="en" docName="zootaxa.4246.1.1SLASH10606.pdf" docOrigin="Zootaxa 4246 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ephippiochthonius tetrachelatus Preyssler 1790, n. comb." docType="treatment" docVersion="9" lastPageNumber="121" masterDocId="FFB1E660FFBF5516FFCB4161FF86FFD2" masterDocTitle="Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="221" masterPageNumber="1" pageNumber="120" updateTime="1698438337209" updateUser="ExternalLinkService">
<mods:mods id="D442FC2A2CEA85BF7740DAEF2F0D5F7A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="417D36AA80B889E7BDEE27C0A02B1DCA">
<mods:title id="C62C12DF3DBF6FFDAE5B63E0472DA675">Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)</mods:title>
</mods:titleInfo>
<mods:name id="6E374623A06B901ED389DF0DEE337E4E" type="personal">
<mods:role id="57569F0E2AC5791518B749852C4D3BCA">
<mods:roleTerm id="8A926B2DCB2DB5BE54B6DCDE983FE961">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D1A65C6195B89B940042226ED2FD52D3">Zaragoza, Juan A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="089A0A7832E73B0CECF77268B6BD056F">text</mods:typeOfResource>
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<mods:title id="D8D5B996090D8B927556DDEF32D5D901">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="39806DF1AAD942450639714F1A1FBB26">
<mods:date id="B190FB557230E4F6242400CE98562BC2">2017</mods:date>
<mods:detail id="838128D6D5BAC810C5B013377A995A63" type="volume">
<mods:number id="8912741CC5CFA87633E40F9273C6C153">4246</mods:number>
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<mods:detail id="C520AE04A75445C8FC62E9B21E006C7B" type="issue">
<mods:number id="6D067A99A6134C28119AE62B91F911A1">1</mods:number>
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<mods:start id="1302F2244EFF3BBE1786D0B47C6B8047">1</mods:start>
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<mods:classification id="13C72D0573F2902890F1E612F0EEC102">journal article</mods:classification>
<mods:identifier id="94540FAD7F1A1163E332EAC599DBD11F" type="CLB-Dataset">36212</mods:identifier>
<mods:identifier id="BA0835F8C651237C85B78908F0CA89BE" type="DOI">10.5281/zenodo.437611</mods:identifier>
<mods:identifier id="E470C3BC25D6D719E4BC1E0013BEB913" type="GBIF-Dataset">3ff74501-6241-43b2-a52f-0e7580bb2f45</mods:identifier>
<mods:identifier id="4DB1B5D8B651BE3EC69B94BF08351F3B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5A3B40AC30A164AB7AE3E7EEA4B70D4C" type="Zenodo-Dep">437611</mods:identifier>
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</mods:mods>
<treatment id="03889E18FFC8556EFF5C41F6FDD7FBC2" ID-DOI="http://doi.org/10.5281/zenodo.5687121" ID-GBIF-Taxon="127852063" ID-Zenodo-Dep="5687121" LSID="urn:lsid:plazi:treatment:03889E18FFC8556EFF5C41F6FDD7FBC2" httpUri="http://treatment.plazi.org/id/03889E18FFC8556EFF5C41F6FDD7FBC2" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">
<subSubSection id="C33B7C85FFC85561FF5C41F6FE11FF01" pageId="119" pageNumber="120" type="nomenclature">
<paragraph id="8B9E2F0EFFC85561FF5C41F6FCE9FF60" blockId="119.[151,879,151,211]" box="[151,879,151,178]" pageId="119" pageNumber="120">
<heading id="D0D69862FFC85561FF5C41F6FCE9FF60" bold="true" box="[151,879,151,178]" fontSize="11" level="1" pageId="119" pageNumber="120" reason="1">
<emphasis id="B955F31CFFC85561FF5C41F6FCE9FF60" bold="true" box="[151,879,151,178]" pageId="119" pageNumber="120">
<taxonomicName id="4C21548DFFC85561FF5C41F6FD71FF60" ID-CoL="6FPZY" authority="Preyssler, 1790" authorityName="Preyssler" authorityYear="1790" box="[151,759,151,178]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="tetrachelatus" status="comb. nov.">
<emphasis id="B955F31CFFC85561FF5C41F6FDA4FF63" bold="true" box="[151,546,151,177]" italics="true" pageId="119" pageNumber="120">Ephippiochthonius tetrachelatus</emphasis>
(Preyssler, 1790)
</taxonomicName>
,
<taxonomicNameLabel id="A2664E67FFC85561FCCE41F9FCE9FF60" box="[773,879,152,178]" pageId="119" pageNumber="120">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E2F0EFFC85561FF5C41DBFE11FF01" blockId="119.[151,879,151,211]" box="[151,407,186,211]" pageId="119" pageNumber="120">
(
<figureCitation id="131A338BFFC85561FF5441DBFE85FF01" box="[159,259,186,211]" captionStart="FIGURES 1 – 3" captionStartId="8.[151,264,1797,1820]" captionTargetBox="[183,1404,607,1740]" captionTargetId="figure@8.[151,1436,588,1774]" captionTargetPageId="8" captionText="FIGURES 1 – 3. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., chela of male from Královice-Hájek, U. Markéty, Prague, Czech Republic. 1, antiaxial view, slightly dorsal; 2, dorsal view; 3, ventral view. Abbreviations: ap: apodeme; as: antiaxial sensory setae; bc: basal condyle; ca: antiaxial condyle; cp: paraxial condyle; eh: edge; fa: antiaxial lyrifissure of fixed chelal finger; fb: basal lyrifissure of fixed chelal finger; fd 1, fd 2, fd 3: dorsal lyrifissures of the fixed chelal finger; hd: distal lyrifissure of chelal hand; hp: proximal lyrifissure of chelal hand; ip: intercondylar protuberance; ma 1, ma 2: antiaxial lyrifissures of movable chelal finger; mv 1, mv 2: ventral lyrifissures of movable chelal finger; sp: subdistal protuberance; td: accessory tooth." httpUri="https://zenodo.org/record/437612/files/figure.png" pageId="119" pageNumber="120">Figs 1–3</figureCitation>
,
<figureCitation id="131A338BFFC85561FEC441DAFEBAFF01" box="[271,316,187,211]" captionStart="FIGURES 4 – 7" captionStartId="9.[151,264,1366,1389]" captionTargetBox="[198,1410,200,1356]" captionTargetId="figure@9.[154,1439,177,1392]" captionTargetPageId="9" captionText="FIGURES 4 – 7. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., males from Královice-Hájek, U. Markéty, Prague, Czech Republic (4 ‒ 6) and male from Sea Cliff, Nassau, New York, U. S. A. (7). 4, left chelicera; 5, external genital area; 6, basal portion of movable chelal finger; 7. Basal portions of chelal fingers. Abbreviations: ap: apodeme; bc: basal condyle; ca: antiaxial condyle; fo: fixed chelal finger foramen; ip: intercondylar protuberance; ls: lateral setae; ldb, ldst, ldt, lvb, lve, lvt: lyrifissures associated with cheliceral setae." httpUri="https://zenodo.org/record/437613/files/figure.png" pageId="119" pageNumber="120">4–7</figureCitation>
,
<figureCitation id="131A338BFFC85561FE8C41DAFE09FF01" box="[327,399,187,211]" captionStart="FIGURES 32 – 39" captionStartId="26.[151,264,1857,1880]" captionTargetBox="[225,1359,193,1840]" captionTargetId="figure@26.[225,1359,193,1841]" captionTargetPageId="26" captionText="FIGURES 32 – 39. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., SEM images. Female, Královice-Hájek, U. Markéty, Prague, Czech Republic (32 – 37) and female, Madeira (38, 39). 32, habitus, antiaxial view, slightly dorsal; 33, detail of chelal fingers, antiaxial view (some adhesive remains between fingers); 34, distal portion of chelal hand and bases of fingers, paraxial view; 35, distal portion of chelal hand and base of movable finger, ventral view; 36, distal portion of chelal hand, dorsal view; 37, medial portion of chelal hand, dorsal view; 38, basal portion of movable chelal finger, antiaxial view; 39, distal portion of movable chelal finger, ventral view. See Material and methods for abbreviations." httpUri="https://zenodo.org/record/437621/files/figure.png" pageId="119" pageNumber="120">32–39</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFC85561FF5C4063FB55FEC9" box="[151,1235,258,283]" pageId="119" pageNumber="120" type="materials_examined">
<paragraph id="8B9E2F0EFFC85561FF5C4063FB55FEC9" blockId="119.[151,1437,258,2017]" box="[151,1235,258,283]" pageId="119" pageNumber="120">
<materialsCitation id="3B492553FFC85561FF5C4063FB55FEC9" ID-GBIF-Occurrence="1449740838" box="[151,1235,258,283]" country="Czech Republic" elevation="209" latitude="50.075554" location="Karlovo" longLatPrecision="18" longitude="14.419723" pageId="119" pageNumber="120" specimenCount="1" stateProvince="Prague" typeStatus="holotype">
<emphasis id="B955F31CFFC85561FF5C4063FEB3FEC9" bold="true" box="[151,309,258,283]" pageId="119" pageNumber="120">
<typeStatus id="549A91ACFFC85561FF5C4063FF54FEC9" box="[151,210,258,283]" pageId="119" pageNumber="120">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3366F9EFFC85561FEF74063FE77FEC9" box="[316,497,258,283]" name="Czech Republic" pageId="119" pageNumber="120">Czech Republic</collectingCountry>
,
<collectingRegion id="49E5E1ECFFC85561FE374062FDCBFEC9" box="[508,589,259,283]" country="Czech Republic" name="Praha" pageId="119" pageNumber="120">Prague</collectingRegion>
,
<location id="8EFE79D5FFC85561FD934063FD32FEC9" LSID="urn:lsid:plazi:treatment:03889E18FFC8556EFF5C41F6FDD7FBC2:8EFE79D5FFC85561FD934063FD32FEC9" box="[600,692,258,283]" country="Czech Republic" latitude="50.075554" longLatPrecision="18" longitude="14.419723" name="Karlovo" pageId="119" pageNumber="120" stateProvince="Prague">Karlovo</location>
náměstí (
<geoCoordinate id="EE1549C9FFC85561FCEA4063FC21FEC9" box="[801,935,258,283]" direction="north" orientation="latitude" pageId="119" pageNumber="120" precision="15" value="50.075554">50°04′32″N</geoCoordinate>
,
<geoCoordinate id="EE1549C9FFC85561FC784063FBB3FEC9" box="[947,1077,258,283]" direction="east" orientation="longitude" pageId="119" pageNumber="120" precision="15" value="14.419723">14°25′11″E</geoCoordinate>
;
<elevation id="000CC83DFFC85561FB8B4062FB42FEC9" box="[1088,1220,258,283]" metricMagnitude="2" metricUnit="m" metricValue="2.09" pageId="119" pageNumber="120" unit="m" value="209.0">
<quantity id="4CD982EBFFC85561FB8B4062FB01FEC8" box="[1088,1159,259,283]" metricMagnitude="2" metricUnit="m" metricValue="2.09" pageId="119" pageNumber="120" unit="m" value="209.0">209 m</quantity>
a.s.l.
</elevation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFC85561FF0C4047FA1AFE55" pageId="119" pageNumber="120" type="distribution">
<paragraph id="8B9E2F0EFFC85561FF0C4047FA1AFE55" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
<emphasis id="B955F31CFFC85561FF0C4047FEE6FEED" bold="true" box="[199,352,294,319]" pageId="119" pageNumber="120">Distribution.</emphasis>
<collectingRegion id="49E5E1ECFFC85561FEA54046FE4FFEEC" box="[366,457,295,318]" country="American Samoa" name="Western" pageId="119" pageNumber="120">Western</collectingRegion>
Palaearctic Region, chiefly Mediterranean; introduced to
<collectingRegion id="49E5E1ECFFC85561FB4F4046FB5EFEEC" box="[1156,1240,295,318]" country="American Samoa" name="Eastern" pageId="119" pageNumber="120">Eastern</collectingRegion>
<collectingCountry id="F3366F9EFFC85561FB2D4047FABBFEED" box="[1254,1341,294,319]" name="Canada" pageId="119" pageNumber="120">Canada</collectingCountry>
,
<collectingCountry id="F3366F9EFFC85561FA844046FA1AFEED" box="[1359,1436,295,319]" name="United States of America" pageId="119" pageNumber="120">U.S.A.</collectingCountry>
(including
<collectingRegion id="49E5E1ECFFC85561FED2402BFEE8FEB1" box="[281,366,330,355]" country="United States of America" name="Hawaii" pageId="119" pageNumber="120">Hawaii</collectingRegion>
),
<collectingCountry id="F3366F9EFFC85561FE42402BFE41FEB1" box="[393,455,330,355]" name="Cuba" pageId="119" pageNumber="120">Cuba</collectingCountry>
,
<collectingCountry id="F3366F9EFFC85561FE11402BFDCAFEB1" box="[474,588,330,355]" name="Argentina" pageId="119" pageNumber="120">Argentina</collectingCountry>
,
<collectingCountry id="F3366F9EFFC85561FD94402BFD51FEB1" box="[607,727,330,355]" name="Seychelles" pageId="119" pageNumber="120">Seychelles</collectingCountry>
and southwestern
<collectingCountry id="F3366F9EFFC85561FC0B402BFBA1FEB1" box="[960,1063,330,355]" name="Australia" pageId="119" pageNumber="120">Australia</collectingCountry>
(
<bibRefCitation id="EFB052FFFFC85561FBF6402BFB59FEB1" author="Gardini" box="[1085,1247,330,355]" pageId="119" pageNumber="120" refString="Gardini, G. (2013) A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas. Zootaxa, 3655 (1), 1 - 151. https: // doi. org / 10.11646 / zootaxa. 3655.1.1" type="journal article" year="2013">Gardini 2013</bibRefCitation>
; Harvey 2013). Records for the Iberian Peninsula and
<collectingRegion id="49E5E1ECFFC85561FD8B400FFD75FE55" box="[576,755,366,391]" country="Spain" name="Illes Baleares" pageId="119" pageNumber="120">Balearic Islands</collectingRegion>
are given in
<bibRefCitation id="EFB052FFFFC85561FC48400EFBBBFE55" author="Zaragoza" box="[899,1085,367,391]" pageId="119" pageNumber="120" refString="Zaragoza, J. A. (2007) Catalogo de los pseudoescorpiones de la Peninsula Iberica e Islas Baleares (Arachnida: Pseudoscorpiones). Revista Iberica Aracnologia, 13, 3 - 91." type="journal article" year="2007">Zaragoza (2007)</bibRefCitation>
and
<bibRefCitation id="EFB052FFFFC85561FBB9400EFA1EFE55" author="Zaragoza" box="[1138,1432,366,391]" pageId="119" pageNumber="120" refString="Zaragoza, J. A. &amp; Vadell, M. (2013) Chthonius (Chthonius) campaneti, a new pseudoscorpion species from Coves del Campanet, Mallorca (Spain). Revista Iberica de Aracnologia, 23, 33 - 41." type="journal article" year="2013">Zaragoza &amp; Vadell (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFC85561FF0C40F3FA1BFD3D" pageId="119" pageNumber="120" type="diagnosis">
<paragraph id="8B9E2F0EFFC85561FF0C40F3FA1BFD3D" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
<emphasis id="B955F31CFFC85561FF0C40F3FEC8FE79" bold="true" box="[199,334,402,427]" pageId="119" pageNumber="120">Diagnosis (</emphasis>
<specimenCount id="9D27E487FFC85561FE8640F2FEE0FE78" box="[333,358,403,426]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
<specimenCount id="9D27E487FFC85561FEAD40F2FEF1FE78" box="[358,375,403,426]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
<emphasis id="B955F31CFFC85561FEB340F3FE0EFE79" bold="true" box="[376,392,402,427]" pageId="119" pageNumber="120">).</emphasis>
A medium-sized, epigean
<taxonomicName id="4C21548DFFC85561FD0840F3FC11FE79" box="[707,919,402,427]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="119" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B955F31CFFC85561FD0840F3FC11FE79" box="[707,919,402,427]" italics="true" pageId="119" pageNumber="120">Ephippiochthonius</emphasis>
</taxonomicName>
species of the
<taxonomicName id="4C21548DFFC85561FB8140F3FB5AFE79" box="[1098,1244,402,427]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="B955F31CFFC85561FB8140F3FB5AFE79" box="[1098,1244,402,427]" italics="true" pageId="119" pageNumber="120">tetrachelatus</emphasis>
</taxonomicName>
-group. Movable cheliceral finger without isolated subapical tooth (
<emphasis id="B955F31CFFC85561FD0640D7FD65FE1D" box="[717,739,438,463]" italics="true" pageId="119" pageNumber="120">di</emphasis>
) and with spinneret in both sexes, lyrifissure
<emphasis id="B955F31CFFC85561FB2D40D7FA8CFE1D" box="[1254,1290,438,463]" italics="true" pageId="119" pageNumber="120">ldb</emphasis>
present; two pairs of eyes with lenses, posterior margin of carapace with 2 macrosetae or rarely with 2 macrosetae and with 1 lateral microseta on one side; pedipalp with femoral chaetotaxy 3:6:2:5:1; chelal hand not depressed or very weakly at level of
<emphasis id="B955F31CFFC85561FEB64343FE15FDE9" box="[381,403,546,571]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FE514343FE3DFDE9" box="[410,443,546,571]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
, with a long, low and rounded hump distad of
<emphasis id="B955F31CFFC85561FBCA4343FB91FDE9" box="[1025,1047,546,571]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FBD44343FBC6FDE9" box="[1055,1088,546,571]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
and moderate slope between trichobothria
<emphasis id="B955F31CFFC85561FEF94327FECEFD8D" box="[306,328,582,607]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FE9B4327FEF7FD8D" box="[336,369,582,607]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
and
<emphasis id="B955F31CFFC85561FE654327FE4EFD8D" box="[430,456,582,607]" italics="true" pageId="119" pageNumber="120">eb</emphasis>
; fixed chelal finger with 14–22 teeth; distal half of movable chelal finger with 6–8 pointed teeth with dental canals, proximal half without a marginal lamina, with 5–9 low vestigial teeth; pedipalpal femur (
<specimenCount id="9D27E487FFC85561FF2043EEFE87FD75" box="[235,257,655,679]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
) 5.4–7.5, (
<specimenCount id="9D27E487FFC85561FE4B43EEFE14FD75" box="[384,402,655,679]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
) 5.3–7.0 times longer than broad, length (
<specimenCount id="9D27E487FFC85561FCBF43EEFC0CFD75" box="[884,906,655,679]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
)
<quantity id="4CD982EBFFC85561FC5043EFFBC7FD74" box="[923,1089,654,679]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" metricValueMax="6.8999999999999995" metricValueMin="4.699999999999999" pageId="119" pageNumber="120" unit="mm" value="0.58" valueMax="0.69" valueMin="0.47">0.47–0.69 mm</quantity>
, (
<specimenCount id="9D27E487FFC85561FB9343EEFBECFD75" box="[1112,1130,655,679]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
)
<quantity id="4CD982EBFFC85561FBB543EFFAA2FD74" box="[1150,1316,654,679]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" metricValueMax="7.6" metricValueMin="5.2" pageId="119" pageNumber="120" unit="mm" value="0.64" valueMax="0.76" valueMin="0.52">0.52–0.76 mm</quantity>
; chela (
<specimenCount id="9D27E487FFC85561FAB643EEFA15FD75" box="[1405,1427,655,679]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
) 4.4–5.8, (
<specimenCount id="9D27E487FFC85561FECE43D2FE91FD19" box="[261,279,691,715]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
) 4.2–5.8 times longer than deep, length (
<specimenCount id="9D27E487FFC85561FD3843D2FC8FFD19" box="[755,777,691,715]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
)
<quantity id="4CD982EBFFC85561FCD143D3FC47FD18" box="[794,961,690,715]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.5" metricValueMin="6.5" pageId="119" pageNumber="120" unit="mm" value="0.8" valueMax="0.95" valueMin="0.65">0.65–0.95 mm</quantity>
, (
<specimenCount id="9D27E487FFC85561FC1243D2FC6DFD19" box="[985,1003,691,715]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
)
<quantity id="4CD982EBFFC85561FC3443D2FB21FD18" box="[1023,1191,691,715]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="10.4" metricValueMin="7.1" pageId="119" pageNumber="120" unit="mm" value="0.875" valueMax="1.04" valueMin="0.71">0.71–1.04 mm</quantity>
; ratio movable chelal finger/chelal hand (
<specimenCount id="9D27E487FFC85561FEB943B6FE0EFD3D" box="[370,392,727,751]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
) 1.3–1.4, (
<specimenCount id="9D27E487FFC85561FDCF43B6FD90FD3D" box="[516,534,727,751]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
) 1.2–1.3; all chelal lyrifissure patterns present with their standard complements.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFC85561FF0C439BFCB2FC15" pageId="119" pageNumber="120" type="materials_examined">
<paragraph id="8B9E2F0EFFC85561FF0C439BFCB2FC15" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
<materialsCitation id="3B492553FFC85561FF0C439BFE8EFCE5" ID-GBIF-Occurrence="1449740828" collectingDate="1998-05-23" collectionCode="FSC" collectorName="Dolni Pocernice" country="Czech Republic" location="Chodov" municipality="Non-type" pageId="119" pageNumber="120" specimenCount="2" specimenCount-male="2" stateProvince="Prague">
<emphasis id="B955F31CFFC85561FF0C439BFDA3FCC1" bold="true" box="[199,549,762,787]" pageId="119" pageNumber="120">
<collectingMunicipality id="6BFAB574FFC85561FF0C439BFEB3FCC1" box="[199,309,762,787]" pageId="119" pageNumber="120">Non-type</collectingMunicipality>
material examined.
</emphasis>
<collectingCountry id="F3366F9EFFC85561FDE6439BFD62FCC1" box="[557,740,762,787]" name="Czech Republic" pageId="119" pageNumber="120">Czech Republic</collectingCountry>
,
<collectingRegion id="49E5E1ECFFC85561FD3B439AFCC4FCC1" box="[752,834,763,787]" country="Czech Republic" name="Praha" pageId="119" pageNumber="120">Prague</collectingRegion>
:
<location id="8EFE79D5FFC85561FC85439BFC2EFCC1" LSID="urn:lsid:plazi:treatment:03889E18FFC8556EFF5C41F6FDD7FBC2:8EFE79D5FFC85561FC85439BFC2EFCC1" box="[846,936,762,787]" country="Czech Republic" municipality="Non-type" name="Chodov" pageId="119" pageNumber="120" stateProvince="Prague">Chodov</location>
,
<specimenCount id="9D27E487FFC85561FC7E439AFC66FCC0" box="[949,992,763,787]" pageId="119" pageNumber="120" type="male">2 ♂</specimenCount>
(
<collectionCode id="ED30B7CBFFC85561FC3B439AFBA1FCC1" box="[1008,1063,763,787]" country="Canada" httpUri="http://grbio.org/cool/q309-g0ck" name="Fredericton Stock Culture Collection" pageId="119" pageNumber="120">FSC</collectionCode>
),
<date id="FF9F09CEFFC85561FBF7439AFB37FCC1" box="[1084,1201,763,787]" pageId="119" pageNumber="120" value="1998-05-23">
<collectingDate id="EFDBF026FFC85561FBF7439AFB37FCC1" box="[1084,1201,763,787]" pageId="119" pageNumber="120" value="1998-05-23">23.V.1998</collectingDate>
</date>
, under stones;
<collectorName id="26D44AD8FFC85561FA97439BFE8EFCE5" pageId="119" pageNumber="120">Dolní Počernice</collectorName>
</materialsCitation>
, 2 ♂, 3 ♀ (1 ♂, 2 ♀ FSC; 1 ♂, 1 ♀ DEUA),
<date id="FF9F09CEFFC85561FCE6427EFC44FCE5" box="[813,962,799,823]" pageId="119" pageNumber="120" value="1998-08-31">31.VIII.1998</date>
, under stones; Čiklova street, 2 ♂ (FSC, DEUA),
<date id="FF9F09CEFFC85561FF374222FEF9FC89" box="[252,383,835,859]" pageId="119" pageNumber="120" value="1998-09-01">01.IX.1998</date>
, leaf litter; Malostranský cemetery, 2 ♂ (FSC),
<date id="FF9F09CEFFC85561FC5C4222FB9CFC89" box="[919,1050,835,859]" pageId="119" pageNumber="120" value="1998-09-21">21.IX.1998</date>
, leaf litter; Husovy sady, 1 ♂, 1 ♀ (FSC),
<date id="FF9F09CEFFC85561FF254206FEF7FCAD" box="[238,369,871,895]" pageId="119" pageNumber="120" value="1998-09-21">21.IX.1998</date>
, leaf litter; Královice-Hájek (U. Markéty), 5 ♂, 4 ♀ (2 ♂, 2 ♀ FSC; 3 ♂, 2 ♀ DEUA),
<date id="FF9F09CEFFC85561FF5C42EAFE8AFC71" box="[151,268,907,931]" pageId="119" pageNumber="120" value="1999-05-01">01.V.1999</date>
, under the stones; Tyršův vrch, 1 ♂, 1 ♀ (FSC),
<date id="FF9F09CEFFC85561FC8242EAFC53FC71" box="[841,981,907,931]" pageId="119" pageNumber="120" value="1999-12-13">13.XII.1999</date>
, leaf litter; Botanic garden, 1 ♀ (FSC),
<date id="FF9F09CEFFC85561FF5C42CEFEAAFC15" box="[151,300,943,967]" pageId="119" pageNumber="120" value="2000-08-29">29.VIII.2000</date>
, under a piece of wood; all leg. F. Šťáhlavský.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFC8556EFF0C42B3FB45FCC7" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120" type="description">
<paragraph id="8B9E2F0EFFC85561FF0C42B3FC7BFBDD" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
<emphasis id="B955F31CFFC85561FF0C42B3FEE1FC39" bold="true" box="[199,359,978,1003]" pageId="119" pageNumber="120">Description (</emphasis>
<specimenCount id="9D27E487FFC85561FEA342B2FE07FC38" box="[360,385,979,1002]" pageId="119" pageNumber="120" type="male">♂</specimenCount>
<specimenCount id="9D27E487FFC85561FE4B42B2FE17FC38" box="[384,401,979,1002]" pageId="119" pageNumber="120" type="female">♀</specimenCount>
<emphasis id="B955F31CFFC85561FE5942B3FE24FC39" bold="true" box="[402,418,978,1003]" pageId="119" pageNumber="120">).</emphasis>
Medium-sized epigean species. Integument pigmented; marked hispid granulation on lateral surfaces of carapace, on cheliceral hand and on bases of chelal fingers.
</paragraph>
<paragraph id="8B9E2F0EFFC85561FF0C457AFCD6FB35" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
Carapace equal to or slightly shorter or longer than broad and weakly constricted posteriorly; medial part of anterior margin weakly prominent, rarely with epistome, and strongly dentate; anterior eyes with strongly convex lens (diameter
<quantity id="4CD982EBFFC85561FE8F4503FD8CFBA9" box="[324,522,1122,1147]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="119" pageNumber="120" unit="mm" value="0.05" valueMax="0.06" valueMin="0.04">0.040–0.060 mm</quantity>
),
<quantity id="4CD982EBFFC85561FDEB4502FD65FBA8" box="[544,739,1123,1147]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="119" pageNumber="120" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.030–0.040 mm</quantity>
from anterior margin of carapace, posterior eyes more with more or less weakly convex lens than anterior pair, both pairs of eyes with reflecting tapeta. Chaetotaxy: 18 macrosetae and 2 preocular microsetae on each side, rarely one, setal formula mm
<quantity id="4CD982EBFFC85561FBFF45CBFBF0FB11" box="[1076,1142,1194,1219]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="119" pageNumber="120" unit="mm" value="4.0">4mm</quantity>
:6:4:2:2, anteromedial setae
<quantity id="4CD982EBFFC85561FF5C45AEFEBFFB34" box="[151,313,1231,1255]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" metricValueMax="11.0" metricValueMin="7" pageId="119" pageNumber="120" unit="mm" value="0.09" valueMax="0.11" valueMin="0.07">0.07–0.11 mm</quantity>
long; 4 lyrifissures anteriorly and 2 posteriorly.
</paragraph>
<paragraph id="8B9E2F0EFFC85561FF0C4593FE33FA31" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
Chelicera (
<figureCitation id="131A338BFFC85561FE8F4593FE0DFAD9" box="[324,395,1266,1291]" captionStart="FIGURES 4 – 7" captionStartId="9.[151,264,1366,1389]" captionTargetBox="[198,1410,200,1356]" captionTargetId="figure@9.[154,1439,177,1392]" captionTargetPageId="9" captionText="FIGURES 4 – 7. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., males from Královice-Hájek, U. Markéty, Prague, Czech Republic (4 ‒ 6) and male from Sea Cliff, Nassau, New York, U. S. A. (7). 4, left chelicera; 5, external genital area; 6, basal portion of movable chelal finger; 7. Basal portions of chelal fingers. Abbreviations: ap: apodeme; bc: basal condyle; ca: antiaxial condyle; fo: fixed chelal finger foramen; ip: intercondylar protuberance; ls: lateral setae; ldb, ldst, ldt, lvb, lve, lvt: lyrifissures associated with cheliceral setae." httpUri="https://zenodo.org/record/437613/files/figure.png" pageId="119" pageNumber="120">Fig. 4</figureCitation>
) with 6 setae and 2 lateral microsetae on hand, seta
<emphasis id="B955F31CFFC85561FC384593FB88FAD9" box="[1011,1038,1266,1291]" italics="true" pageId="119" pageNumber="120">vb</emphasis>
short and almost twice as long as microsetae; hand with 5 dorsal and 1 ventral lyrifissure, lyrifissure
<emphasis id="B955F31CFFC85561FC744477FC65FAFD" box="[959,995,1302,1327]" italics="true" pageId="119" pageNumber="120">ldb</emphasis>
present. Fixed finger with 6–9 teeth decreasing in size proximally, 1–2 distal teeth distinctly larger than others and 4–6 proximal microtubercles. Movable finger without an isolated subapical tooth (
<emphasis id="B955F31CFFC85561FD3F443FFC8CFAA5" box="[756,778,1374,1399]" italics="true" pageId="119" pageNumber="120">di</emphasis>
), with 5–6 teeth decreasing in size proximally, the distal tooth larger than others, 3–5 proximal microtubercles; spinneret prominent in female, lower and more rounded in male; seta
<emphasis id="B955F31CFFC85561FEDD44C7FEADFA6D" box="[278,299,1446,1471]" italics="true" pageId="119" pageNumber="120">gl</emphasis>
0.53–0.66 from base of movable finger. Rallum with 11 blades. Serrula exterior with 14 blades, serrula interior 12 blades.
</paragraph>
<paragraph id="8B9E2F0EFFC85561FF0C448FFDF2F9A0" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
Chaetotaxy of tergites 4:4:4:4:6:6:6:6:1T2T1:4:1T2T1:0, tergites IX and XI each with 2 sublateral tactile setae. Chaetotaxy of sternites 10–11:(3)8–10(3):(2)7–8(2):8–9:6:6:6:6: 2T1T2:0:2, lateral setae on sternite III microsetal in size, sternite X with 2 submedial tactile setae (
<figureCitation id="131A338BFFC85561FD764757FC87F99D" box="[701,769,1590,1615]" captionStart="FIGURES 4 – 7" captionStartId="9.[151,264,1366,1389]" captionTargetBox="[198,1410,200,1356]" captionTargetId="figure@9.[154,1439,177,1392]" captionTargetPageId="9" captionText="FIGURES 4 – 7. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., males from Královice-Hájek, U. Markéty, Prague, Czech Republic (4 ‒ 6) and male from Sea Cliff, Nassau, New York, U. S. A. (7). 4, left chelicera; 5, external genital area; 6, basal portion of movable chelal finger; 7. Basal portions of chelal fingers. Abbreviations: ap: apodeme; bc: basal condyle; ca: antiaxial condyle; fo: fixed chelal finger foramen; ip: intercondylar protuberance; ls: lateral setae; ldb, ldst, ldt, lvb, lve, lvt: lyrifissures associated with cheliceral setae." httpUri="https://zenodo.org/record/437613/files/figure.png" pageId="119" pageNumber="120">Fig. 5</figureCitation>
); in addition, genital notch of male flanked by 6–9 setae on each side and 4+4 internal glandular setae.
</paragraph>
<paragraph id="8B9E2F0EFFC85561FF0C471FFB93F90D" blockId="119.[151,1437,258,2017]" pageId="119" pageNumber="120">
Pedipalpal coxa with 5 setae (including 2 on manducatory process); coxa
<date id="FF9F09CEFFC85561FBD8471EFBB5F945" box="[1043,1075,1663,1687]" pageId="119" pageNumber="120">I 3</date>
+ 3 marginal microsetae, distal marginal seta of pedipalpal disk distinctly longer than the marginal seta of coxa I;
<date id="FF9F09CEFFC85561FB9747C2FB01F969" box="[1116,1159,1699,1723]" pageId="119" pageNumber="120">II 4</date>
+ 7–12 bipinnate coxal spines,
<date id="FF9F09CEFFC85561FF2047A6FE9AF90D" box="[235,284,1734,1759]" pageId="119" pageNumber="120">III 5</date>
+ 3–7 bipinnate coxal spines and
<date id="FF9F09CEFFC85561FD5147A6FD49F90D" box="[666,719,1734,1759]" pageId="119" pageNumber="120">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="8B9E2F0EFFC8556EFF0C478BFC03FDCA" blockId="119.[151,1437,258,2017]" lastBlockId="120.[151,1437,151,1040]" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">
Pedipalp with femoral chaetotaxy 3:6:2:5:1, rarely 3 setae in the posterior dorsal row of one chela and
<quantity id="4CD982EBFFC85561FA8A478AFAEAF8D1" box="[1345,1388,1770,1795]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="119" pageNumber="120" unit="in" value="2.0">2 in</quantity>
that of the other. Chela (
<figureCitation id="131A338BFFC85561FE49466FFE6EF8F5" box="[386,488,1806,1831]" captionStart="FIGURES 1 – 3" captionStartId="8.[151,264,1797,1820]" captionTargetBox="[183,1404,607,1740]" captionTargetId="figure@8.[151,1436,588,1774]" captionTargetPageId="8" captionText="FIGURES 1 – 3. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., chela of male from Královice-Hájek, U. Markéty, Prague, Czech Republic. 1, antiaxial view, slightly dorsal; 2, dorsal view; 3, ventral view. Abbreviations: ap: apodeme; as: antiaxial sensory setae; bc: basal condyle; ca: antiaxial condyle; cp: paraxial condyle; eh: edge; fa: antiaxial lyrifissure of fixed chelal finger; fb: basal lyrifissure of fixed chelal finger; fd 1, fd 2, fd 3: dorsal lyrifissures of the fixed chelal finger; hd: distal lyrifissure of chelal hand; hp: proximal lyrifissure of chelal hand; ip: intercondylar protuberance; ma 1, ma 2: antiaxial lyrifissures of movable chelal finger; mv 1, mv 2: ventral lyrifissures of movable chelal finger; sp: subdistal protuberance; td: accessory tooth." httpUri="https://zenodo.org/record/437612/files/figure.png" pageId="119" pageNumber="120">Figs 1–3</figureCitation>
) with hand not depressed or very weakly at level of
<emphasis id="B955F31CFFC85561FB92466FFBE9F8F5" box="[1113,1135,1806,1831]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FBBC466FFB1EF8F5" box="[1143,1176,1806,1831]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
, with a long, low and rounded hump distad of
<emphasis id="B955F31CFFC85561FE674653FE44F899" box="[428,450,1842,1867]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FE014653FE6DF899" box="[458,491,1842,1867]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
and moderate slope between trichobothria
<emphasis id="B955F31CFFC85561FC1F4653FC6CF899" box="[980,1002,1842,1867]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FC394653FB95F899" box="[1010,1043,1842,1867]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
and
<emphasis id="B955F31CFFC85561FB804653FBE0F899" box="[1099,1126,1842,1867]" italics="true" pageId="119" pageNumber="120">eb</emphasis>
; dorsal-antiaxial surface of the chelal hand between the hump and trichobothria
<emphasis id="B955F31CFFC85561FCE34637FCC5F8BD" box="[808,835,1878,1903]" italics="true" pageId="119" pageNumber="120">eb</emphasis>
/
<emphasis id="B955F31CFFC85561FC814637FCE9F8BD" box="[842,879,1878,1903]" italics="true" pageId="119" pageNumber="120">esb</emphasis>
flattened, limited by a marked edge; ventral intercondylar bow-like protuberance (
<emphasis id="B955F31CFFC85561FD89461DFDDEF841" box="[578,600,1916,1939]" italics="true" pageId="119" pageNumber="120">ip</emphasis>
) present (
<figureCitation id="131A338BFFC85561FD03461BFCF3F841" box="[712,885,1914,1939]" captionStart="FIGURES 32 – 39" captionStartId="26.[151,264,1857,1880]" captionTargetBox="[225,1359,193,1840]" captionTargetId="figure@26.[225,1359,193,1841]" captionTargetPageId="26" captionText="FIGURES 32 – 39. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., SEM images. Female, Královice-Hájek, U. Markéty, Prague, Czech Republic (32 – 37) and female, Madeira (38, 39). 32, habitus, antiaxial view, slightly dorsal; 33, detail of chelal fingers, antiaxial view (some adhesive remains between fingers); 34, distal portion of chelal hand and bases of fingers, paraxial view; 35, distal portion of chelal hand and base of movable finger, ventral view; 36, distal portion of chelal hand, dorsal view; 37, medial portion of chelal hand, dorsal view; 38, basal portion of movable chelal finger, antiaxial view; 39, distal portion of movable chelal finger, ventral view. See Material and methods for abbreviations." httpUri="https://zenodo.org/record/437621/files/figure.png" pageId="119" pageNumber="120">Figs 34, 35, 39</figureCitation>
); chaetotaxy 4:5:3, seta
<emphasis id="B955F31CFFC85561FB4C461BFB2DF845" box="[1159,1195,1914,1943]" italics="true" pageId="119" pageNumber="120">ph3</emphasis>
lacking, setae
<emphasis id="B955F31CFFC85561FA99461BFAF6F845" box="[1362,1392,1914,1943]" italics="true" pageId="119" pageNumber="120">ih1</emphasis>
,
<emphasis id="B955F31CFFC85561FAB5461BFA1AF845" box="[1406,1436,1914,1943]" italics="true" pageId="119" pageNumber="120">ih3</emphasis>
and
<emphasis id="B955F31CFFC85561FF0D46C0FF62F86C" box="[198,228,1953,1982]" italics="true" pageId="119" pageNumber="120">ih4</emphasis>
approximately level with trichobothria
<emphasis id="B955F31CFFC85561FD6B46C0FD30F868" box="[672,694,1953,1978]" italics="true" pageId="119" pageNumber="120">ib</emphasis>
/
<emphasis id="B955F31CFFC85561FD7546C0FD59F868" box="[702,735,1953,1978]" italics="true" pageId="119" pageNumber="120">isb</emphasis>
; distal end of hand and bases of chelal fingers with sclerotized condylar complex. Fixed finger with 14–22 mostly pointed and weakly reclined teeth progressively decreasing in size and with dental canals, 1‒2 distal ones small, 2–4 most proximal teeth rounded, 2–4 microtubercles at base; tip of finger with an accessory tooth (
<emphasis id="B955F31CFFC7556EFDE841DDFDBFFF07" box="[547,569,188,213]" italics="true" pageId="120" pageNumber="121">td</emphasis>
) on antiaxial face; tip of fixed chelal finger of male with a deep hollow on paraxial face and subdistal protuberance (
<emphasis id="B955F31CFFC7556EFDBA4180FD0CFF2A" box="[625,650,225,248]" italics="true" pageId="120" pageNumber="121">sp</emphasis>
); one pair of very short antiaxial sensory setae (
<emphasis id="B955F31CFFC7556EFB784180FB4AFF2A" box="[1203,1228,225,248]" italics="true" pageId="120" pageNumber="121">as</emphasis>
) at the base, level with or proximad of lyrifissure
<emphasis id="B955F31CFFC7556EFDC94065FD9EFECF" box="[514,536,260,285]" italics="true" pageId="120" pageNumber="121">fb</emphasis>
, distance between them distinctly shorter than finger depth at base; 3–4 teeth, rarely 5, at level of
<emphasis id="B955F31CFFC7556EFEB14048FE1FFE92" box="[378,409,297,320]" italics="true" pageId="120" pageNumber="121">est</emphasis>
/
<emphasis id="B955F31CFFC7556EFE6A4048FE37FE92" box="[417,433,297,320]" italics="true" pageId="120" pageNumber="121">it</emphasis>
occupying
<quantity id="4CD982EBFFC7556EFDFD4049FD16FEED" box="[566,656,296,320]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="120" pageNumber="121" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<quantity id="4CD982EBFFC7556EFBFE4049FB7FFEED" box="[1077,1273,296,320]" metricMagnitude="-5" metricUnit="m" metricValue="3.2" metricValueMax="4.0" metricValueMin="2.4" pageId="120" pageNumber="121" unit="mm" value="0.032" valueMax="0.04" valueMin="0.024">0.024–0.040 mm</quantity>
. Distal half of movable finger with 6–8 pointed teeth with dental canals, the distal one small (rarely absent); proximal half without marginal lamina, with 5–9 low vestigial teeth reaching trichobothrium
<emphasis id="B955F31CFFC7556EFBEE400EFBB8FE5A" box="[1061,1086,367,392]" italics="true" pageId="120" pageNumber="121">sb</emphasis>
or halfway between
<emphasis id="B955F31CFFC7556EFAF8400EFACAFE5A" box="[1331,1356,367,392]" italics="true" pageId="120" pageNumber="121">sb</emphasis>
and
<emphasis id="B955F31CFFC7556EFA4C400EFA13FE5A" box="[1415,1429,367,392]" italics="true" pageId="120" pageNumber="121">b</emphasis>
, mostly with dental canals; basal condyle (
<emphasis id="B955F31CFFC7556EFDB740F5FD10FE7F" box="[636,662,404,429]" italics="true" pageId="120" pageNumber="121">bc</emphasis>
) enlarged and apodeme long and apically indented (
<figureCitation id="131A338BFFC7556EFB3D40F5FAE6FE7E" box="[1270,1376,404,429]" captionStart="FIGURES 4 – 7" captionStartId="9.[151,264,1366,1389]" captionTargetBox="[198,1410,200,1356]" captionTargetId="figure@9.[154,1439,177,1392]" captionTargetPageId="9" captionText="FIGURES 4 – 7. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., males from Královice-Hájek, U. Markéty, Prague, Czech Republic (4 ‒ 6) and male from Sea Cliff, Nassau, New York, U. S. A. (7). 4, left chelicera; 5, external genital area; 6, basal portion of movable chelal finger; 7. Basal portions of chelal fingers. Abbreviations: ap: apodeme; bc: basal condyle; ca: antiaxial condyle; fo: fixed chelal finger foramen; ip: intercondylar protuberance; ls: lateral setae; ldb, ldst, ldt, lvb, lve, lvt: lyrifissures associated with cheliceral setae." httpUri="https://zenodo.org/record/437613/files/figure.png" pageId="120" pageNumber="121">Figs 6, 7</figureCitation>
,
<figureCitation id="131A338BFFC7556EFAA440F5FA08FE7E" box="[1391,1422,404,428]" captionStart="FIGURES 32 – 39" captionStartId="26.[151,264,1857,1880]" captionTargetBox="[225,1359,193,1840]" captionTargetId="figure@26.[225,1359,193,1841]" captionTargetPageId="26" captionText="FIGURES 32 – 39. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., SEM images. Female, Královice-Hájek, U. Markéty, Prague, Czech Republic (32 – 37) and female, Madeira (38, 39). 32, habitus, antiaxial view, slightly dorsal; 33, detail of chelal fingers, antiaxial view (some adhesive remains between fingers); 34, distal portion of chelal hand and bases of fingers, paraxial view; 35, distal portion of chelal hand and base of movable finger, ventral view; 36, distal portion of chelal hand, dorsal view; 37, medial portion of chelal hand, dorsal view; 38, basal portion of movable chelal finger, antiaxial view; 39, distal portion of movable chelal finger, ventral view. See Material and methods for abbreviations." httpUri="https://zenodo.org/record/437621/files/figure.png" pageId="120" pageNumber="121">38</figureCitation>
); coupled sensilla
<emphasis id="B955F31CFFC7556EFE9840D8FEE8FE02" box="[339,366,441,464]" italics="true" pageId="120" pageNumber="121">pc</emphasis>
halfway between
<emphasis id="B955F31CFFC7556EFDF640D6FDCDFE02" box="[573,587,439,464]" italics="true" pageId="120" pageNumber="121">b</emphasis>
and
<emphasis id="B955F31CFFC7556EFD4840D6FD1AFE02" box="[643,668,439,464]" italics="true" pageId="120" pageNumber="121">sb</emphasis>
, mostly closer to
<emphasis id="B955F31CFFC7556EFCA340D6FC07FE02" box="[872,897,439,464]" italics="true" pageId="120" pageNumber="121">sb</emphasis>
. Trichobothria as in
<figureCitation id="131A338BFFC7556EFBA540D6FB54FE02" box="[1134,1234,439,464]" captionStart="FIGURES 1 – 3" captionStartId="8.[151,264,1797,1820]" captionTargetBox="[183,1404,607,1740]" captionTargetId="figure@8.[151,1436,588,1774]" captionTargetPageId="8" captionText="FIGURES 1 – 3. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., chela of male from Královice-Hájek, U. Markéty, Prague, Czech Republic. 1, antiaxial view, slightly dorsal; 2, dorsal view; 3, ventral view. Abbreviations: ap: apodeme; as: antiaxial sensory setae; bc: basal condyle; ca: antiaxial condyle; cp: paraxial condyle; eh: edge; fa: antiaxial lyrifissure of fixed chelal finger; fb: basal lyrifissure of fixed chelal finger; fd 1, fd 2, fd 3: dorsal lyrifissures of the fixed chelal finger; hd: distal lyrifissure of chelal hand; hp: proximal lyrifissure of chelal hand; ip: intercondylar protuberance; ma 1, ma 2: antiaxial lyrifissures of movable chelal finger; mv 1, mv 2: ventral lyrifissures of movable chelal finger; sp: subdistal protuberance; td: accessory tooth." httpUri="https://zenodo.org/record/437612/files/figure.png" pageId="120" pageNumber="121">Figs 1‒2</figureCitation>
; trichobothria
<emphasis id="B955F31CFFC7556EFAB340D6FF66FE27" italics="true" pageId="120" pageNumber="121">ebesb-ist</emphasis>
mostly in a straight line,
<emphasis id="B955F31CFFC7556EFDCE40BCFDA6FE26" box="[517,544,477,500]" italics="true" pageId="120" pageNumber="121">ist</emphasis>
always distad of
<emphasis id="B955F31CFFC7556EFD2240BDFC88FE27" box="[745,782,476,501]" italics="true" pageId="120" pageNumber="121">esb</emphasis>
and well proximad of lyrifissure
<emphasis id="B955F31CFFC7556EFB4540BDFB22FE27" box="[1166,1188,476,501]" italics="true" pageId="120" pageNumber="121">fb</emphasis>
. All chelal lyrifissure patterns present with their standard complements (
<figureCitation id="131A338BFFC7556EFD00409EFCFEFDCA" box="[715,888,511,536]" captionStart="FIGURES 32 – 39" captionStartId="26.[151,264,1857,1880]" captionTargetBox="[225,1359,193,1840]" captionTargetId="figure@26.[225,1359,193,1841]" captionTargetPageId="26" captionText="FIGURES 32 – 39. Ephippiochthonius tetrachelatus (Preyssler, 1790) n. comb., SEM images. Female, Královice-Hájek, U. Markéty, Prague, Czech Republic (32 – 37) and female, Madeira (38, 39). 32, habitus, antiaxial view, slightly dorsal; 33, detail of chelal fingers, antiaxial view (some adhesive remains between fingers); 34, distal portion of chelal hand and bases of fingers, paraxial view; 35, distal portion of chelal hand and base of movable finger, ventral view; 36, distal portion of chelal hand, dorsal view; 37, medial portion of chelal hand, dorsal view; 38, basal portion of movable chelal finger, antiaxial view; 39, distal portion of movable chelal finger, ventral view. See Material and methods for abbreviations." httpUri="https://zenodo.org/record/437621/files/figure.png" pageId="120" pageNumber="121">Figs 33, 36, 37</figureCitation>
).
</paragraph>
<paragraph id="8B9E2F0EFFC7556EFF0C4344FB45FCC7" blockId="120.[151,1437,151,1040]" pageId="120" pageNumber="121">
<emphasis id="B955F31CFFC7556EFF0C4344FE6FFDEE" box="[199,489,548,573]" italics="true" pageId="120" pageNumber="121">Measurements and ratios</emphasis>
. The following data are from the description of
<bibRefCitation id="EFB052FFFFC7556EFBE84345FB56FDEE" author="Gardini" box="[1059,1232,548,573]" pageId="120" pageNumber="121" refString="Gardini, G. (2013) A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas. Zootaxa, 3655 (1), 1 - 151. https: // doi. org / 10.11646 / zootaxa. 3655.1.1" type="journal article" year="2013">Gardini (2013)</bibRefCitation>
, who measured a large number of specimens from many populations assigned to this species: Male, followed by female in square brackets: Body 1.3–1.8 [1.4–1.9]. Carapace 0.35–0.50/0.36–0.46 (0.9–1.1) [0.37–0.55/0.38–0.54] Chelicera 0.29– 0.41/0.14–0.19 (1.9–2.2) [0.32–0.45/0.16–0.21], movable finger length 0.13–0.19 [0.16–0.23]. Pedipalp: femur 0.47–0.69/0.08–0.12 (5.4–7.5) [0.52–0.76/ 0.09–0.14 (5.3–7.0)], chela 0.65–0.95/0.12–0.17 (4.4–5.8) [0.71–1.04/ 0.14–0.23 (4.2–5.8)], hand 0.27–0.42 (2.0–2.5) [0.30–0.44 (1.8–2.3)], movable finger 0.38–0.56 [0.40–0.60]; ratio movable finger/hand 1.3–1.4 [1.2–1.3], femur/movable finger 1.2–1.4, femur/carapace 1.3–1.5.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FFC7556EFF0C427EFDD7FBC2" pageId="120" pageNumber="121" type="discussion">
<paragraph id="8B9E2F0EFFC7556EFF0C427EFDD7FBC2" blockId="120.[151,1437,151,1040]" pageId="120" pageNumber="121">
<emphasis id="B955F31CFFC7556EFF0C427EFEBDFCEA" bold="true" box="[199,315,799,824]" pageId="120" pageNumber="121">Remarks.</emphasis>
The presence of
<taxonomicName id="4C21548DFFC7556EFDC74240FD47FCEA" box="[524,705,799,824]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="B955F31CFFC7556EFDC74240FD47FCEA" box="[524,705,799,824]" italics="true" pageId="120" pageNumber="121">E. tetrachelatus</emphasis>
</taxonomicName>
seems less frequent in the Iberian Peninsula than in
<collectingCountry id="F3366F9EFFC7556EFAFE427EFAEEFCEA" box="[1333,1384,799,824]" name="Italy" pageId="120" pageNumber="121">Italy</collectingCountry>
and Central Europe. However, it should be taken into consideration that a disproportionate number of the samples of Iberian
<taxonomicName id="4C21548DFFC7556EFF264206FEF0FC52" box="[237,374,871,896]" class="Arachnida" family="Chthoniidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="120" pageNumber="121" phylum="Arthropoda" rank="family">Chthoniidae</taxonomicName>
studied in both the previous literature and the present work come from hypogean environments in comparison to those collected from epigean environments. Most literature records of
<taxonomicName id="4C21548DFFC7556EFB5A42ECFAC3FC77" box="[1169,1349,908,933]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="tetrachelatus">
<emphasis id="B955F31CFFC7556EFB5A42ECFAC3FC77" box="[1169,1349,908,933]" italics="true" pageId="120" pageNumber="121">E. tetrachelatus</emphasis>
</taxonomicName>
for the Iberian Peninsula need to be verified by restudying the available specimens concerned, and it will not be surprising if some of them are found to correspond to other species [e.g. misidentifications of
<taxonomicName id="4C21548DFFC7556EFBFD42B4FB43FC3F" box="[1078,1221,980,1005]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="andalucia" status="sp. nov.">
<emphasis id="B955F31CFFC7556EFBFD42B4FB43FC3F" box="[1078,1221,980,1005]" italics="true" pageId="120" pageNumber="121">E. andalucia</emphasis>
</taxonomicName>
<emphasis id="B955F31CFFC7556EFB0742B5FA8CFC3F" bold="true" box="[1228,1290,980,1005]" pageId="120" pageNumber="121">
<taxonomicNameLabel id="A2664E67FFC7556EFB0742B5FA8CFC3F" box="[1228,1290,980,1005]" pageId="120" pageNumber="121" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by
<bibRefCitation id="EFB052FFFFC7556EFAFF42B4FEC2FBC2" author="Zaragoza" pageId="120" pageNumber="121" refString="Zaragoza, J. A. &amp; Perez, T. (2013) Hypogean pseudoscorpions (Arachnida) from Jaen province (Andalusia, Spain), with descriptions of four new species and a new synonymy. Zootaxa, 3700 (2), 201 - 225." type="journal article" year="2013">Zaragoza &amp; Pérez (2013)</bibRefCitation>
and
<bibRefCitation id="EFB052FFFFC7556EFEB14299FDC3FBC2" author="Zaragoza" box="[378,581,1016,1040]" pageId="120" pageNumber="121" refString="Zaragoza, J. A. (2013 a) Los pseudoscorpiones (Arachnida) subterraneos de Jaen. In: Grupo de Espeleologia de Villacarrillo (G. E. V.) (Ed.), Los invertebrados de habitats subterraneos de Jaen, 2013, pp. 65 - 71. [Perez Fernandez, T. y Perez Ruiz, A. (Coord.)]" type="journal article" year="2013" yearSuffix="a">Zaragoza (2013a)</bibRefCitation>
].
</paragraph>
</subSubSection>
</treatment>
</document>
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