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<document id="6938B20F7A5E8EFD9A5D468504BC9951" ID="10.11646/zootaxa.4246.1.1SLASH10606" ID-CLB-Dataset="36212" ID-DOI="10.5281/zenodo.437611" ID-GBIF-Dataset="3ff74501-6241-43b2-a52f-0e7580bb2f45" ID-ISSN="1175-5326" ID-Zenodo-Dep="437611" ID-ZooBank="D8594E96-D561-4A37-9073-B138EC7E95A7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1490260213340" checkinUser="plazi" docAuthor="Zaragoza, Juan A." docDate="2017" docId="03889E18FF6D55C0FF5C4658FE3CFE7E" docLanguage="en" docName="zootaxa.4246.1.1SLASH10606.pdf" docOrigin="Zootaxa 4246 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Occidenchthonius villacarrillo Zaragoza &amp; Perez 2013, n. comb." docType="treatment" docVersion="9" lastPageNumber="215" masterDocId="FFB1E660FFBF5516FFCB4161FF86FFD2" masterDocTitle="Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="221" masterPageNumber="1" pageNumber="211" updateTime="1698438337209" updateUser="ExternalLinkService">
<mods:mods id="2C5FDA10BADDC0675EC33F59FB1B4463" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="939AA624D353992815953480E076FA6F">
<mods:title id="905A3B1F21D77DA4AC73E4E1D926FD6C">Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae)</mods:title>
</mods:titleInfo>
<mods:name id="CFD72E7897E674599F6D959DB888249C" type="personal">
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<mods:roleTerm id="D9CD7AC1E2BFCAF1703E6B53E361E931">Author</mods:roleTerm>
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<mods:namePart id="9C6114D7B8B6D5C2C2AF3445F06D422F">Zaragoza, Juan A.</mods:namePart>
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<mods:title id="6CB558A423CD9E5EB926ED8B96195F06">Zootaxa</mods:title>
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<mods:date id="1BC377853224E4F62744E574846014AC">2017</mods:date>
<mods:detail id="570CF18DAFBC3EB9CED360A65B536318" type="volume">
<mods:number id="7AA494A5C13B45EDF6438D0C4B4BC994">4246</mods:number>
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<mods:classification id="53108E88C6D818AC66DFCA21CF8949FF">journal article</mods:classification>
<mods:identifier id="A725DE1B5D517940CAD80C386FCDCBEA" type="CLB-Dataset">36212</mods:identifier>
<mods:identifier id="0E0D6B813707AF8F7489795A5926A267" type="DOI">10.5281/zenodo.437611</mods:identifier>
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</mods:mods>
<treatment id="03889E18FF6D55C0FF5C4658FE3CFE7E" ID-DOI="http://doi.org/10.5281/zenodo.5687215" ID-GBIF-Taxon="127852090" ID-Zenodo-Dep="5687215" LSID="urn:lsid:plazi:treatment:03889E18FF6D55C0FF5C4658FE3CFE7E" httpUri="http://treatment.plazi.org/id/03889E18FF6D55C0FF5C4658FE3CFE7E" lastPageId="214" lastPageNumber="215" pageId="210" pageNumber="211">
<subSubSection id="C33B7C85FF6D55C4FF5C4658FE81F8A7" pageId="210" pageNumber="211" type="nomenclature">
<paragraph id="8B9E2F0EFF6D55C4FF5C4658FC4EF881" blockId="210.[151,968,1849,1909]" box="[151,968,1849,1875]" pageId="210" pageNumber="211">
<heading id="D0D69862FF6D55C4FF5C4658FC4EF881" bold="true" box="[151,968,1849,1875]" fontSize="11" level="1" pageId="210" pageNumber="211" reason="1">
<emphasis id="B955F31CFF6D55C4FF5C4658FC4EF881" bold="true" box="[151,968,1849,1875]" pageId="210" pageNumber="211">
<taxonomicName id="4C21548DFF6D55C4FF5C4658FCD6F881" ID-CoL="48BRN" authority="Zaragoza &amp; Perez, 2013" authorityName="Zaragoza &amp; Perez" authorityYear="2013" box="[151,848,1849,1875]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="210" pageNumber="211" phylum="Arthropoda" rank="species" species="villacarrillo" status="comb. nov.">
<emphasis id="B955F31CFF6D55C4FF5C4658FD81F881" bold="true" box="[151,519,1849,1875]" italics="true" pageId="210" pageNumber="211">Occidenchthonius villacarrillo</emphasis>
(
<bibRefCitation id="EFB052FFFF6D55C4FDDC4658FCCFF881" author="Zaragoza" box="[535,841,1849,1875]" pageId="210" pageNumber="211" refString="Zaragoza, J. A. &amp; Perez, T. (2013) Hypogean pseudoscorpions (Arachnida) from Jaen province (Andalusia, Spain), with descriptions of four new species and a new synonymy. Zootaxa, 3700 (2), 201 - 225." type="journal article" year="2013">Zaragoza &amp; Pérez, 2013</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2664E67FF6D55C4FC954658FC4EF881" box="[862,968,1849,1875]" pageId="210" pageNumber="211">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9E2F0EFF6D55C4FF5C463DFE81F8A7" blockId="210.[151,968,1849,1909]" box="[151,263,1884,1909]" pageId="210" pageNumber="211">
(
<figureCitation id="131A338BFF6D55C4FF54463DFF79F8A7" box="[159,255,1884,1909]" captionStart="FIGURE 321" captionStartId="211.[151,250,1985,2007]" captionTargetBox="[268,1383,223,1847]" captionTargetId="figure@211.[162,1437,179,1903]" captionTargetPageId="211" captionText="FIGURE 321. Occidenchthonius villacarrillo (Zaragoza &amp; Pérez, 2013) n. comb., female holotype, left chela, antiaxial view (modified after Zaragoza &amp; Pérez, 2013)." httpUri="https://zenodo.org/record/437682/files/figure.png" pageId="210" pageNumber="211">Fig. 321</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF6D55C4FF5C46C5FCB5F80B" pageId="210" pageNumber="211" type="reference_group">
<paragraph id="8B9E2F0EFF6D55C4FF5C46C5FCB5F80B" blockId="210.[151,1073,1956,2009]" pageId="210" pageNumber="211">
<treatmentCitationGroup id="AB310820FF6D55C4FF5C46C5FCB5F80B" pageId="210" pageNumber="211">
<taxonomicName id="4C21548DFF6D55C4FF5C46C5FCF9F868" ID-CoL="V3HN" authority="Zaragoza &amp; Perez 2013: 213" authorityName="Zaragoza &amp; Perez" authorityPageNumber="213" authorityYear="2013" box="[151,895,1956,1978]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="210" pageNumber="211" phylum="Arthropoda" rank="species" species="villacarrillo" subGenus="Ephippiochthonius">
<emphasis id="B955F31CFF6D55C4FF5C46C5FF79F868" box="[151,255,1956,1978]" italics="true" pageId="210" pageNumber="211">Chthonius</emphasis>
(
<emphasis id="B955F31CFF6D55C4FEC646C5FE4AF868" box="[269,460,1956,1978]" italics="true" pageId="210" pageNumber="211">Ephippiochthonius</emphasis>
)
<emphasis id="B955F31CFF6D55C4FE1046C5FDD0F868" box="[475,598,1956,1978]" italics="true" pageId="210" pageNumber="211">villacarrillo</emphasis>
<treatmentCitation id="0A80091FFF6D55C4FD9746C4FCF9F868" author="Zaragoza" box="[604,895,1956,1978]" page="213" pageId="210" pageNumber="211" year="2013">
<bibRefCitation id="EFB052FFFF6D55C4FD9746C4FCC9F868" author="Zaragoza" box="[604,847,1956,1978]" pageId="210" pageNumber="211" refString="Zaragoza, J. A. &amp; Perez, T. (2013) Hypogean pseudoscorpions (Arachnida) from Jaen province (Andalusia, Spain), with descriptions of four new species and a new synonymy. Zootaxa, 3700 (2), 201 - 225." type="journal article" year="2013">Zaragoza &amp; Pérez 2013</bibRefCitation>
: 213
</treatmentCitation>
</taxonomicName>
–216, figs 21–26.
<taxonomicName id="4C21548DFF6D55C4FF5C46A2FCB6F80B" ID-CoL="V3HN" authority="Zaragoza 2013a: 68" authorityName="Zaragoza" authorityPageNumber="68" authorityYear="2013" box="[151,816,1987,2009]" class="Arachnida" family="Chthoniidae" genus="Chthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="210" pageNumber="211" phylum="Arthropoda" rank="species" species="villacarrillo" subGenus="Ephippiochthonius">
<emphasis id="B955F31CFF6D55C4FF5C46A2FF79F80B" box="[151,255,1987,2009]" italics="true" pageId="210" pageNumber="211">Chthonius</emphasis>
(
<emphasis id="B955F31CFF6D55C4FEC646A2FE4AF80B" box="[269,460,1987,2009]" italics="true" pageId="210" pageNumber="211">Ephippiochthonius</emphasis>
)
<emphasis id="B955F31CFF6D55C4FE1046A2FDD0F80B" box="[475,598,1987,2009]" italics="true" pageId="210" pageNumber="211">villacarrillo</emphasis>
:
<treatmentCitation id="0A80091FFF6D55C4FDA846A5FCB6F80B" author="Zaragoza" box="[611,816,1987,2009]" page="68" pageId="210" pageNumber="211" year="2013">
<bibRefCitation id="EFB052FFFF6D55C4FDA846A5FC8CF80B" author="Zaragoza" box="[611,778,1987,2009]" pageId="210" pageNumber="211" refString="Zaragoza, J. A. (2013 a) Los pseudoscorpiones (Arachnida) subterraneos de Jaen. In: Grupo de Espeleologia de Villacarrillo (G. E. V.) (Ed.), Los invertebrados de habitats subterraneos de Jaen, 2013, pp. 65 - 71. [Perez Fernandez, T. y Perez Ruiz, A. (Coord.)]" type="journal article" year="2013" yearSuffix="a">Zaragoza 2013a</bibRefCitation>
: 68
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF5E7F86FF6C55C5FF5C46E5FE56F86A" pageId="211" pageNumber="212">
<paragraph id="8B9E2F0EFF6C55C5FF5C46E5FE56F86A" blockId="211.[151,1436,1924,2038]" pageId="211" pageNumber="212">
<emphasis id="B955F31CFF6C55C5FF5C46E5FEA8F84B" bold="true" box="[151,302,1924,1946]" pageId="211" pageNumber="212">FIGURE 320.</emphasis>
<taxonomicName id="4C21548DFF6C55C5FEFE46E5FD59F848" authority="Zaragoza, 1985" authorityName="Zaragoza" authorityYear="1985" box="[309,735,1924,1946]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="verai" status="comb. nov.">
<emphasis id="B955F31CFF6C55C5FEFE46E5FDAEF848" box="[309,552,1924,1946]" italics="true" pageId="211" pageNumber="212">Occidenchthonius verai</emphasis>
(Zaragoza, 1985)
</taxonomicName>
<taxonomicNameLabel id="A2664E67FF6C55C5FD2346E4FCC2F848" box="[744,836,1924,1946]" pageId="211" pageNumber="212">n. comb.</taxonomicNameLabel>
, female paratype, right chela, antiaxial view. See Material and methods for abbreviations.
</paragraph>
</caption>
<caption id="DF5E7F86FF6C55C5FF5C46A0FDBDF824" httpUri="https://zenodo.org/record/437682/files/figure.png" pageId="211" pageNumber="212" targetBox="[268,1383,223,1847]" targetPageId="211">
<paragraph id="8B9E2F0EFF6C55C5FF5C46A0FDBDF824" blockId="211.[151,1436,1924,2038]" pageId="211" pageNumber="212">
<emphasis id="B955F31CFF6C55C5FF5C46A0FEABF804" bold="true" box="[151,301,1985,2007]" pageId="211" pageNumber="212">FIGURE 321.</emphasis>
<taxonomicName id="4C21548DFF6C55C5FEFF46A0FCF9F805" authority="Zaragoza &amp; Perez, 2013" authorityName="Zaragoza &amp; Perez" authorityYear="2013" box="[308,895,1985,2007]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="211" pageNumber="212" phylum="Arthropoda" rank="species" species="villacarrillo" status="comb. nov.">
<emphasis id="B955F31CFF6C55C5FEFF46A0FDE8F805" box="[308,622,1985,2007]" italics="true" pageId="211" pageNumber="212">Occidenchthonius villacarrillo</emphasis>
(Zaragoza &amp; Pérez, 2013)
</taxonomicName>
<taxonomicNameLabel id="A2664E67FF6C55C5FC4D46A3FC64F805" box="[902,994,1985,2007]" pageId="211" pageNumber="212">n. comb.</taxonomicNameLabel>
, female holotype, left chela, antiaxial view (modified after Zaragoza &amp; Pérez, 2013).
</paragraph>
</caption>
<subSubSection id="C33B7C85FF6B55C2FF5C41F6FEAFFF07" pageId="212" pageNumber="213" type="materials_examined">
<paragraph id="8B9E2F0EFF6B55C2FF5C41F6FEAFFF07" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
<materialsCitation id="3B492553FF6B55C2FF5C41F6FEAFFF07" ID-GBIF-Occurrence="1449740737" collectorName="Cueva de la Morciguilla" country="Spain" elevation="693" latitude="38.09111" location="Jaen province" longLatPrecision="19" longitude="-3.0016668" pageId="212" pageNumber="213" specimenCount="1" stateProvince="Andalusia" typeStatus="holotype">
<emphasis id="B955F31CFF6B55C2FF5C41F6FEB1FF62" bold="true" box="[151,311,151,176]" pageId="212" pageNumber="213">
<typeStatus id="549A91ACFF6B55C2FF5C41F6FF54FF62" box="[151,210,151,176]" pageId="212" pageNumber="213">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3366F9EFF6B55C2FEF541F6FE07FF62" box="[318,385,151,176]" name="Spain" pageId="212" pageNumber="213">Spain</collectingCountry>
,
<collectingRegion id="49E5E1ECFF6B55C2FE4641F6FD87FF62" box="[397,513,151,176]" country="Spain" name="Andalucia" pageId="212" pageNumber="213">Andalusia</collectingRegion>
,
<location id="8EFE79D5FF6B55C2FDC541F9FD2BFF62" LSID="urn:lsid:plazi:treatment:03889E18FF6D55C0FF5C4658FE3CFE7E:8EFE79D5FF6B55C2FDC541F9FD2BFF62" box="[526,685,151,176]" country="Spain" latitude="38.09111" longLatPrecision="19" longitude="-3.0016668" name="Jaen province" pageId="212" pageNumber="213" stateProvince="Andalusia">Jaén province</location>
,
<taxonomicName id="4C21548DFF6B55C2FD7141F6FCC2FF62" box="[698,836,151,176]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="212" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo">Villacarrillo</taxonomicName>
,
<collectorName id="26D44AD8FF6B55C2FC9B41F9FBE5FF62" box="[848,1123,151,176]" pageId="212" pageNumber="213">Cueva de la Morciguilla</collectorName>
(
<geoCoordinate id="EE1549C9FF6B55C2FBBF41F6FB7BFF62" box="[1140,1277,151,176]" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="15" value="38.09111">38°05′28″N</geoCoordinate>
,
<geoCoordinate id="EE1549C9FF6B55C2FAC241F6FA1EFF62" box="[1289,1432,151,176]" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="15" value="-3.0016668">03°00′06″W</geoCoordinate>
;
<elevation id="000CC83DFF6B55C2FF5C41DDFE9CFF07" box="[151,282,188,213]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="212" pageNumber="213" unit="m" value="693.0">
<quantity id="4CD982EBFF6B55C2FF5C41DDFF58FF06" box="[151,222,188,213]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="212" pageNumber="213" unit="m" value="693.0">693 m</quantity>
a.s.l.
</elevation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF6B55C2FF0C41BEFDA9FF2A" box="[199,559,223,248]" pageId="212" pageNumber="213" type="distribution">
<paragraph id="8B9E2F0EFF6B55C2FF0C41BEFDA9FF2A" blockId="212.[151,1437,151,2032]" box="[199,559,223,248]" pageId="212" pageNumber="213">
<emphasis id="B955F31CFF6B55C2FF0C41BEFEE6FF2A" bold="true" box="[199,352,223,248]" pageId="212" pageNumber="213">Distribution.</emphasis>
<collectingCountry id="F3366F9EFF6B55C2FEAC41BEFE2DFF2A" box="[359,427,223,248]" name="Spain" pageId="212" pageNumber="213">Spain</collectingCountry>
:
<collectingRegion id="49E5E1ECFF6B55C2FE7E41BEFDACFF2A" box="[437,554,223,248]" country="Spain" name="Andalucia" pageId="212" pageNumber="213">Andalusia</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF6B55C2FF0C4065FCBBFD8C" pageId="212" pageNumber="213" type="diagnosis">
<paragraph id="8B9E2F0EFF6B55C2FF0C4065FCBBFD8C" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
<emphasis id="B955F31CFF6B55C2FF0C4065FECCFECF" bold="true" box="[199,330,260,285]" pageId="212" pageNumber="213">Diagnosis (</emphasis>
<specimenCount id="9D27E487FF6B55C2FE814064FED9FECE" box="[330,351,261,284]" pageId="212" pageNumber="213" type="female">♀</specimenCount>
<emphasis id="B955F31CFF6B55C2FE944065FEE9FECF" bold="true" box="[351,367,260,285]" pageId="212" pageNumber="213">).</emphasis>
A small, hypogean, weakly troglomorphic
<taxonomicName id="4C21548DFF6B55C2FC984065FB9BFECF" box="[851,1053,260,285]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="212" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B955F31CFF6B55C2FC984065FB9BFECF" box="[851,1053,260,285]" italics="true" pageId="212" pageNumber="213">Occidenchthonius</emphasis>
</taxonomicName>
species. Chelicera with 6 setae on hand, movable cheliceral finger with isolated subapical tooth (
<emphasis id="B955F31CFF6B55C2FC9E4046FCECFE92" box="[853,874,295,320]" italics="true" pageId="212" pageNumber="213">di</emphasis>
) and spinneret prominent in female, unknown for male; lyrifissure
<emphasis id="B955F31CFF6B55C2FE9C402BFEFDFEB1" box="[343,379,330,355]" italics="true" pageId="212" pageNumber="213">ldb</emphasis>
absent; without eyes or eye-spots, anterior margin of carapace without preocular microsetae, posterior margin with 2 macrosetae; chelal hand strongly depressed at level of
<emphasis id="B955F31CFF6B55C2FBC1400FFBA6FE55" box="[1034,1056,366,391]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FBEC400FFBCEFE55" box="[1063,1096,366,391]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
, with a short,prominent hump distad of
<emphasis id="B955F31CFF6B55C2FECA40F0FE91FE78" box="[257,279,401,426]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FED540F0FEB9FE78" box="[286,319,401,426]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
and moderate slope between trichobothria
<emphasis id="B955F31CFF6B55C2FCE340F0FCB8FE78" box="[808,830,401,426]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FC8E40F0FCE0FE78" box="[837,870,401,426]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FC5540F0FC3FFE78" box="[926,953,401,426]" italics="true" pageId="212" pageNumber="213">eb</emphasis>
; trichobothrium
<emphasis id="B955F31CFF6B55C2FBB340F2FB14FE78" box="[1144,1170,403,426]" italics="true" pageId="212" pageNumber="213">ist</emphasis>
distad of lyrifissure
<emphasis id="B955F31CFF6B55C2FAB540F0FA12FE78" box="[1406,1428,401,426]" italics="true" pageId="212" pageNumber="213">fb</emphasis>
; fixed chelal finger with 18 teeth; distal half of movable chelal finger with 12 pointed teeth with dental canals, basal half of movable chelal finger with 9 rounded, partially fused, vestigial teeth, without canals, on weakly raised lamina; pedipalpal femur (
<specimenCount id="9D27E487FF6B55C2FE0D409CFE5EFDC7" box="[454,472,509,533]" pageId="212" pageNumber="213" type="female">♀</specimenCount>
) 5.5 times longer than broad, length
<quantity id="4CD982EBFF6B55C2FC4B409DFC6EFDC6" box="[896,1000,508,533]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="212" pageNumber="213" unit="mm" value="0.56">0.56 mm</quantity>
; chela (
<specimenCount id="9D27E487FF6B55C2FB88409CFBD3FDC7" box="[1091,1109,509,533]" pageId="212" pageNumber="213" type="female">♀</specimenCount>
) 5.2 times longer than deep, length
<quantity id="4CD982EBFF6B55C2FF2E4341FECAFDE5" box="[229,332,544,568]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="212" pageNumber="213" unit="mm" value="0.78">0.78 mm</quantity>
; ratio movable chelal finger/chelal hand (
<specimenCount id="9D27E487FF6B55C2FCEF4341FCB0FDEA" box="[804,822,544,568]" pageId="212" pageNumber="213" type="female">♀</specimenCount>
) 1.4; lacking lyrifissures
<emphasis id="B955F31CFF6B55C2FB914341FB02FDEE" box="[1114,1156,544,572]" italics="true" pageId="212" pageNumber="213">ma1</emphasis>
and
<emphasis id="B955F31CFF6B55C2FB704341FB63FDEE" box="[1211,1253,544,572]" italics="true" pageId="212" pageNumber="213">ma2</emphasis>
, all other chelal lyrifissure patterns present with their standard complements.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF6B55C2FF0C4308FC6AFD77" pageId="212" pageNumber="213" type="materials_examined">
<paragraph id="8B9E2F0EFF6B55C2FF0C4308FC6AFD77" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
<materialsCitation id="3B492553FF6B55C2FF0C4308FC6AFD77" ID-GBIF-Occurrence="1449740731" collectingDate="2013-02-24" collectionCode="DEUA" collectorName="Cueva de la Morciguilla" country="Spain" elevation="693" latitude="38.09111" location="Jaen province" longLatPrecision="19" longitude="-3.0016668" pageId="212" pageNumber="213" specimenCount="2" specimenCount-female="1" stateProvince="Andalusia" typeStatus="holotype">
<emphasis id="B955F31CFF6B55C2FF0C4308FE72FD50" bold="true" box="[199,500,617,642]" pageId="212" pageNumber="213">
<typeStatus id="549A91ACFF6B55C2FF0C4308FE84FD50" box="[199,258,617,642]" pageId="212" pageNumber="213">Type</typeStatus>
material examined.
</emphasis>
<specimenCount id="9D27E487FF6B55C2FE36430BFD94FD53" box="[509,530,618,641]" pageId="212" pageNumber="213" type="female">♀</specimenCount>
<typeStatus id="549A91ACFF6B55C2FDD74308FDF8FD50" box="[540,638,617,642]" pageId="212" pageNumber="213" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED30B7CBFF6B55C2FD5B430BFD65FD50" box="[656,739,618,642]" pageId="212" pageNumber="213">DEUA</collectionCode>
),
<collectingCountry id="F3366F9EFF6B55C2FD324308FCBAFD50" box="[761,828,617,642]" name="Spain" pageId="212" pageNumber="213">Spain</collectingCountry>
,
<collectingRegion id="49E5E1ECFF6B55C2FC814308FC38FD50" box="[842,958,617,642]" country="Spain" name="Andalucia" pageId="212" pageNumber="213">Andalusia</collectingRegion>
,
<location id="8EFE79D5FF6B55C2FC07430BFBEAFD50" LSID="urn:lsid:plazi:treatment:03889E18FF6D55C0FF5C4658FE3CFE7E:8EFE79D5FF6B55C2FC07430BFBEAFD50" box="[972,1132,617,642]" country="Spain" latitude="38.09111" longLatPrecision="19" longitude="-3.0016668" name="Jaen province" pageId="212" pageNumber="213" stateProvince="Andalusia">Jaén province</location>
,
<taxonomicName id="4C21548DFF6B55C2FBB14308FA82FD50" box="[1146,1284,617,642]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="212" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo">Villacarrillo</taxonomicName>
,
<collectorName id="26D44AD8FF6B55C2FAD9430BFE9DFD77" pageId="212" pageNumber="213">Cueva de la Morciguilla</collectorName>
(
<geoCoordinate id="EE1549C9FF6B55C2FEE143EDFE35FD77" box="[298,435,652,677]" direction="north" orientation="latitude" pageId="212" pageNumber="213" precision="15" value="38.09111">38°05′28″N</geoCoordinate>
,
<geoCoordinate id="EE1549C9FF6B55C2FE7543EDFDCAFD77" box="[446,588,652,677]" direction="west" orientation="longitude" pageId="212" pageNumber="213" precision="15" value="-3.0016668">03°00′06″W</geoCoordinate>
;
<elevation id="000CC83DFF6B55C2FD9343EDFD5CFD77" box="[600,730,652,677]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="212" pageNumber="213" unit="m" value="693.0">
<quantity id="4CD982EBFF6B55C2FD9343EDFD19FD77" box="[600,671,652,677]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="212" pageNumber="213" unit="m" value="693.0">693 m</quantity>
a.s.l.
</elevation>
),
<date id="FF9F09CEFF6B55C2FD3A43ECFCEEFD77" box="[753,872,653,677]" pageId="212" pageNumber="213" value="2013-02-24">
<collectingDate id="EFDBF026FF6B55C2FD3A43ECFCEEFD77" box="[753,872,653,677]" pageId="212" pageNumber="213" value="2013-02-24">
24.
<bibRefCitation id="EFB052FFFF6B55C2FCD843ECFCEEFD77" author="II" box="[787,872,653,677]" pageId="212" pageNumber="213" refString="Gardini, G. (2013) A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas. Zootaxa, 3655 (1), 1 - 151. https: // doi. org / 10.11646 / zootaxa. 3655.1.1" type="journal article" year="2013">II.2013</bibRefCitation>
</collectingDate>
</date>
, leg. G.E.V.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF6B55C0FF0C43D1FEF9FECE" lastPageId="214" lastPageNumber="215" pageId="212" pageNumber="213" type="description">
<paragraph id="8B9E2F0EFF6B55C2FF0C43D1FF6AFCC3" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
<emphasis id="B955F31CFF6B55C2FF0C43D1FEEDFD1B" bold="true" box="[199,363,688,713]" pageId="212" pageNumber="213">Description (</emphasis>
<specimenCount id="9D27E487FF6B55C2FEA043D0FE06FD1A" box="[363,384,689,712]" pageId="212" pageNumber="213" type="female">♀</specimenCount>
;
<specimenCount id="9D27E487FF6B55C2FE5143D0FE29FD1A" box="[410,431,689,712]" pageId="212" pageNumber="213" type="male">♂</specimenCount>
<emphasis id="B955F31CFF6B55C2FE0B43D1FDC6FD1B" bold="true" box="[448,576,688,713]" pageId="212" pageNumber="213">unknown).</emphasis>
Small, hypogean species of weakly troglomorphic facies. Integument depigmented; marked hispid granulation on lateral surfaces of carapace, on cheliceral hand and on bases of chelal fingers.
</paragraph>
<paragraph id="8B9E2F0EFF6B55C2FF0C427CFCE3FC4D" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
Carapace as long as broad, weakly constricted posteriorly; medial part of anterior margin strongly prominent, with rudimentary epistome, and strongly dentate. Without eyes or eye-spots. Chaetotaxy: 18 setae, without preocular microsetae, 2 setae in posterior row, formula 4:6:4:2:2, anteromedial setae
<quantity id="4CD982EBFF6B55C2FBB24205FB67FCA9" box="[1145,1249,868,892]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="212" pageNumber="213" unit="mm" value="0.08">0.08 mm</quantity>
long, sublateral ocular setae
<quantity id="4CD982EBFF6B55C2FEE842E7FE0CFC4D" box="[291,394,902,927]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="212" pageNumber="213" unit="mm" value="0.06">0.06 mm</quantity>
; 4 lyrifissures anteriorly and 2 posteriorly.
</paragraph>
<paragraph id="8B9E2F0EFF6B55C2FF0C42CBFEE5FBA6" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
Cheliceral hand with 6 setae and 1 lateral microseta, seta
<emphasis id="B955F31CFF6B55C2FC8142CBFCE3FC11" box="[842,869,938,963]" italics="true" pageId="212" pageNumber="213">vb</emphasis>
very short (
<quantity id="4CD982EBFF6B55C2FC2042CBFBD9FC10" box="[1003,1119,938,963]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" pageId="212" pageNumber="213" unit="mm" value="0.025">0.025 mm</quantity>
long), microseta
<quantity id="4CD982EBFF6B55C2FAE842CBFA1EFC10" box="[1315,1432,938,962]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="212" pageNumber="213" unit="mm" value="0.02">0.020 mm</quantity>
; hand with 4 dorsal and 1 ventral lyrifissure, lyrifissure
<emphasis id="B955F31CFF6B55C2FCDE42ACFCBFFC34" box="[789,825,973,998]" italics="true" pageId="212" pageNumber="213">ldb</emphasis>
absent. Fixed finger with 11 teeth decreasing in size proximally, two distal teeth distinctly larger than others. Movable finger with an isolated subapical tooth (
<emphasis id="B955F31CFF6B55C2FAF74290FAD7FBD8" box="[1340,1361,1009,1034]" italics="true" pageId="212" pageNumber="213">di</emphasis>
), with 8 teeth decreasing in size proximally, the distal tooth larger than others; spinneret prominent in female, unknown for male; seta
<emphasis id="B955F31CFF6B55C2FEFC4559FECAFB83" box="[311,332,1080,1105]" italics="true" pageId="212" pageNumber="213">gl</emphasis>
0.53 from base of movable finger. Rallum with 11 blades. Serrula exterior with 15 blades, serrula interior 13 blades.
</paragraph>
<paragraph id="8B9E2F0EFF6B55C2FF0C451FFDD8FB0C" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">Chaetotaxy of tergites 4:4:4:4:6:6:6:6:1T2T1:4:1T2T1:0, tergites IX and XI each with 2 sublateral tactile setae. Chaetotaxy of sternites 9:(3)8(3):(2)7(2):8:6:6:6:6:2T1T2:0:2, lateral setae on sternite III macrosetal in size, sternite X with 2 submedial tactile setae.</paragraph>
<paragraph id="8B9E2F0EFF6B55C2FF0C4588FCC7FA9B" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
Pedipalpal coxa with 5 setae (including 2 on manducatory process), distal marginal seta of disk
<quantity id="4CD982EBFF6B55C2FB33458BFADAFAD3" box="[1272,1372,1258,1282]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="212" pageNumber="213" unit="mm" value="0.07">0.07 mm</quantity>
long; coxa
<date id="FF9F09CEFF6B55C2FF1F446FFF75FAF7" box="[212,243,1293,1317]" pageId="212" pageNumber="213">I 3</date>
+ 3 marginal microsetae, distal marginal seta
<quantity id="4CD982EBFF6B55C2FD35446DFCE5FAF7" box="[766,867,1292,1317]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="212" pageNumber="213" unit="mm" value="0.05">0.05 mm</quantity>
long;
<date id="FF9F09CEFF6B55C2FC67446FFC53FAF7" box="[940,981,1293,1317]" pageId="212" pageNumber="213">II 4</date>
+ 9–11 bipinnate coxal spines,
<date id="FF9F09CEFF6B55C2FAF6446FFAE9FAF7" box="[1341,1391,1292,1317]" pageId="212" pageNumber="213">III 5</date>
+ 4 bipinnate coxal spines and
<date id="FF9F09CEFF6B55C2FE0D4450FE7DFA9B" box="[454,507,1328,1353]" pageId="212" pageNumber="213">IV 6</date>
; intercoxal tubercle bisetose.
</paragraph>
<paragraph id="8B9E2F0EFF6B55C2FF0C4435FEB2F822" blockId="212.[151,1437,151,2032]" pageId="212" pageNumber="213">
Pedipalp with femoral chaetotaxy 3:6:3:5:1. Chela (
<figureCitation id="131A338BFF6B55C2FCDF4435FCF3FABE" box="[788,885,1364,1389]" captionStart="FIGURE 321" captionStartId="211.[151,250,1985,2007]" captionTargetBox="[268,1383,223,1847]" captionTargetId="figure@211.[162,1437,179,1903]" captionTargetPageId="211" captionText="FIGURE 321. Occidenchthonius villacarrillo (Zaragoza &amp; Pérez, 2013) n. comb., female holotype, left chela, antiaxial view (modified after Zaragoza &amp; Pérez, 2013)." httpUri="https://zenodo.org/record/437682/files/figure.png" pageId="212" pageNumber="213">Fig. 321</figureCitation>
) with hand strongly depressed at level of
<emphasis id="B955F31CFF6B55C2FA9C4435FAEBFABF" box="[1367,1389,1364,1389]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FABF4435FA13FABF" box="[1396,1429,1364,1389]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
, with a short, prominent hump distad of
<emphasis id="B955F31CFF6B55C2FDA14416FD06FA42" box="[618,640,1399,1424]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FD4C4416FD2EFA42" box="[647,680,1399,1424]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
and moderate slope between trichobothria
<emphasis id="B955F31CFF6B55C2FB6B4416FB30FA42" box="[1184,1206,1399,1424]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FB764416FB58FA42" box="[1213,1246,1399,1424]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FAD04416FAB0FA42" box="[1307,1334,1399,1424]" italics="true" pageId="212" pageNumber="213">eb</emphasis>
; marked hollow before base of movable finger with thicker cuticle; width equal to depth; chaetotaxy 4:5:4, seta
<emphasis id="B955F31CFF6B55C2FAD344FBFABAFA65" box="[1304,1340,1434,1463]" italics="true" pageId="212" pageNumber="213">ph3</emphasis>
present; distal end of hand and bases of chelal fingers with sclerotized condylar complex. Fixed finger with 18 mostly pointed teeth and with dental canals, first two distal teeth small, third tooth of normal row (
<emphasis id="B955F31CFF6B55C2FB0E4487FB67FA2F" box="[1221,1249,1510,1533]" italics="true" pageId="212" pageNumber="213">mt</emphasis>
) of fixed finger distinctly modified in shape and deviated in orientation with respect to the others, most proximal tooth small and rounded, dental row reaching slightly beyond halfway between trichobothria
<emphasis id="B955F31CFF6B55C2FC35474AFB91F996" box="[1022,1047,1579,1604]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FB85474AFBDAF996" box="[1102,1116,1579,1604]" italics="true" pageId="212" pageNumber="213">b</emphasis>
, 4 proximal microtubercles; tip of fixed finger with an accessory tooth (
<emphasis id="B955F31CFF6B55C2FDB5472FFD12F9B5" box="[638,660,1614,1639]" italics="true" pageId="212" pageNumber="213">td</emphasis>
) on antiaxial face; one pair of long antiaxial sensory setae (
<emphasis id="B955F31CFF6B55C2FAFF4731FACAF9B5" box="[1332,1356,1616,1639]" italics="true" pageId="212" pageNumber="213">as</emphasis>
) at the base, one level with and other markedly distad of lyrifissure
<emphasis id="B955F31CFF6B55C2FC944713FCF3F959" box="[863,885,1650,1675]" italics="true" pageId="212" pageNumber="213">fb</emphasis>
,
<quantity id="4CD982EBFF6B55C2FC4C4713FBCFF958" box="[903,1097,1650,1675]" metricMagnitude="-5" metricUnit="m" metricValue="3" metricValueMax="3.5" metricValueMin="2.5" pageId="212" pageNumber="213" unit="mm" value="0.03" valueMax="0.035" valueMin="0.025">0.025–0.035 mm</quantity>
long, distance between them
<quantity id="4CD982EBFF6B55C2FF5C47F7FE8AF97C" box="[151,268,1686,1710]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="212" pageNumber="213" unit="mm" value="0.03">0.030 mm</quantity>
, fixed finger depth at the base
<quantity id="4CD982EBFF6B55C2FDA047F7FD66F97C" box="[619,736,1686,1710]" metricMagnitude="-5" metricUnit="m" metricValue="4.8" pageId="212" pageNumber="213" unit="mm" value="0.048">0.048 mm</quantity>
; 6 teeth at level of
<emphasis id="B955F31CFF6B55C2FC7047F6FC5CF97C" box="[955,986,1687,1710]" italics="true" pageId="212" pageNumber="213">est</emphasis>
/
<emphasis id="B955F31CFF6B55C2FC2947F6FC77F97C" box="[994,1009,1687,1710]" italics="true" pageId="212" pageNumber="213">it</emphasis>
occupying
<quantity id="4CD982EBFF6B55C2FBBE47F7FB48F97C" box="[1141,1230,1686,1710]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="212" pageNumber="213" unit="mm" value="0.1">0.1 mm</quantity>
, distance between successive apices
<quantity id="4CD982EBFF6B55C2FEAC47D8FDABF903" box="[359,557,1721,1746]" metricMagnitude="-5" metricUnit="m" metricValue="1.65" metricValueMax="1.7999999999999998" metricValueMin="1.5" pageId="212" pageNumber="213" unit="mm" value="0.0165" valueMax="0.018" valueMin="0.015">0.015–0.018 mm</quantity>
. Distal half of movable finger with 12 pointed teeth with dental canals that reaches slightly beyond halfway between trichobothria
<emphasis id="B955F31CFF6B55C2FCCC47BFFC9CF927" box="[775,794,1758,1781]" italics="true" pageId="212" pageNumber="213">st</emphasis>
and
<emphasis id="B955F31CFF6B55C2FC9947BDFCEDF927" box="[850,875,1756,1781]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
, 3 most proximal tooth distinctly reduced in size, distal tooth tiny; basal half of movable chelal finger with 9 rounded, partially fused, vestigial teeth, without canals, on weakly raised lamina; dental row reaching halfway between
<emphasis id="B955F31CFF6B55C2FC534642FC37F8EE" box="[920,945,1827,1852]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FC384642FB87F8EE" box="[1011,1025,1827,1852]" italics="true" pageId="212" pageNumber="213">b</emphasis>
; basal apodeme long and apically indented; coupled sensilla
<emphasis id="B955F31CFF6B55C2FE194629FE6BF88D" box="[466,493,1864,1887]" italics="true" pageId="212" pageNumber="213">pc</emphasis>
halfway between
<emphasis id="B955F31CFF6B55C2FD014626FD65F8B2" box="[714,739,1863,1888]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FCEF4626FCB4F8B2" box="[804,818,1863,1888]" italics="true" pageId="212" pageNumber="213">b</emphasis>
, slightly closer to
<emphasis id="B955F31CFF6B55C2FBD34626FBB7F8B2" box="[1048,1073,1863,1888]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
. Trichobothria as in
<figureCitation id="131A338BFF6B55C2FAFA4626FA11F88D" box="[1329,1431,1863,1888]" captionStart="FIGURE 321" captionStartId="211.[151,250,1985,2007]" captionTargetBox="[268,1383,223,1847]" captionTargetId="figure@211.[162,1437,179,1903]" captionTargetPageId="211" captionText="FIGURE 321. Occidenchthonius villacarrillo (Zaragoza &amp; Pérez, 2013) n. comb., female holotype, left chela, antiaxial view (modified after Zaragoza &amp; Pérez, 2013)." httpUri="https://zenodo.org/record/437682/files/figure.png" pageId="212" pageNumber="213">Fig. 321</figureCitation>
; trichobothrium
<emphasis id="B955F31CFF6B55C2FE8C460DFEE7F851" box="[327,353,1900,1923]" italics="true" pageId="212" pageNumber="213">ist</emphasis>
well distad of
<emphasis id="B955F31CFF6B55C2FDC7460BFDB7F851" box="[524,561,1898,1923]" italics="true" pageId="212" pageNumber="213">esb</emphasis>
and very slightly distad of lyrifissure
<emphasis id="B955F31CFF6B55C2FC2F460BFC7CF851" box="[996,1018,1898,1923]" italics="true" pageId="212" pageNumber="213">fb</emphasis>
; distance between
<emphasis id="B955F31CFF6B55C2FB1F460BFB6CF851" box="[1236,1258,1898,1923]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FB3A460BFA94F851" box="[1265,1298,1898,1923]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
and base of hand approximately equal to that between
<emphasis id="B955F31CFF6B55C2FD4F46EFFD1CF875" box="[644,666,1934,1959]" italics="true" pageId="212" pageNumber="213">ib</emphasis>
/
<emphasis id="B955F31CFF6B55C2FD6A46EFFD44F875" box="[673,706,1934,1959]" italics="true" pageId="212" pageNumber="213">isb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FD3546EFFCA5F875" box="[766,803,1934,1959]" italics="true" pageId="212" pageNumber="213">esb</emphasis>
; distance between
<emphasis id="B955F31CFF6B55C2FBC946EEFB93F874" box="[1026,1045,1935,1958]" italics="true" pageId="212" pageNumber="213">st</emphasis>
and
<emphasis id="B955F31CFF6B55C2FB9B46EFFBEFF875" box="[1104,1129,1934,1959]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
1.8 times longer than that between
<emphasis id="B955F31CFF6B55C2FF3746D0FE93F818" box="[252,277,1969,1994]" italics="true" pageId="212" pageNumber="213">sb</emphasis>
and
<emphasis id="B955F31CFF6B55C2FE8646D0FEDDF818" box="[333,347,1969,1994]" italics="true" pageId="212" pageNumber="213">b</emphasis>
; lacking lyrifissures
<emphasis id="B955F31CFF6B55C2FD8146D3FDF2F81C" box="[586,628,1970,1998]" italics="true" pageId="212" pageNumber="213">ma1</emphasis>
and
<emphasis id="B955F31CFF6B55C2FD6746D3FD50F81C" box="[684,726,1970,1998]" italics="true" pageId="212" pageNumber="213">ma2</emphasis>
, all other chelal lyrifissure patterns present with their standard complements.
</paragraph>
<caption id="DF5E7F86FF6A55C3FF5C46C1FE00F826" httpUri="https://zenodo.org/record/437683/files/figure.png" pageId="213" pageNumber="214" targetBox="[189,1436,240,1874]" targetPageId="213">
<paragraph id="8B9E2F0EFF6A55C3FF5C46C1FE00F826" blockId="213.[151,1436,1952,2036]" pageId="213" pageNumber="214">
<emphasis id="B955F31CFF6A55C3FF5C46C1FEE9F864" bold="true" box="[151,367,1952,1975]" pageId="213" pageNumber="214">FIGURES 322–323.</emphasis>
Known distribution of genera in the Iberian Peninsula and Balearic Islands. 322,
<taxonomicName id="4C21548DFF6A55C3FB1946C0FA13F865" box="[1234,1429,1953,1975]" class="Arachnida" family="Chthoniidae" genus="Cantabrochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="213" pageNumber="33" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B955F31CFF6A55C3FB1946C0FA13F865" box="[1234,1429,1953,1975]" italics="true" pageId="213" pageNumber="214">Cantabrochthonius</emphasis>
</taxonomicName>
: diamonds;
<taxonomicName id="4C21548DFF6A55C3FEC146A1FE4FF804" box="[266,457,1984,2006]" class="Arachnida" family="Chthoniidae" genus="Ephippiochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B955F31CFF6A55C3FEC146A1FE4FF804" box="[266,457,1984,2006]" italics="true" pageId="213" pageNumber="214">Ephippiochthonius</emphasis>
</taxonomicName>
: squares;
<taxonomicName id="4C21548DFF6A55C3FDF346A1FD69F804" box="[568,751,1984,2006]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="213" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B955F31CFF6A55C3FDF346A1FD69F804" box="[568,751,1984,2006]" italics="true" pageId="213" pageNumber="214">Occidenchthonius</emphasis>
</taxonomicName>
: circles. 323,
<taxonomicName id="4C21548DFF6A55C3FC4346A1FBB9F804" box="[904,1087,1984,2006]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="213" pageNumber="127" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B955F31CFF6A55C3FC4346A1FBB9F804" box="[904,1087,1984,2006]" italics="true" pageId="213" pageNumber="214">Occidenchthonius</emphasis>
</taxonomicName>
(coloured areas), Canary Islands displaced and enlarged.
</paragraph>
</caption>
<paragraph id="8B9E2F0EFF6955C0FF0C41F8FEF9FECE" blockId="214.[151,1436,151,429]" pageId="214" pageNumber="215">
<emphasis id="B955F31CFF6955C0FF0C41F8FE61FF62" box="[199,487,151,176]" italics="true" pageId="214" pageNumber="215">Measurements and ratios</emphasis>
. Female
<typeStatus id="549A91ACFF6955C0FD9841F6FD3CFF62" box="[595,698,151,176]" pageId="214" pageNumber="215" type="holotype">holotype</typeStatus>
: Body 1.30. Carapace 0.41/0.41 (1.0). Chelicera 0.36/0.17 (2.1), movable finger 0.19. Pedipalp: femur 0.56/0.10 (5.5), patella 0.23/0.11 (2.0), chela 0.78/0.15 (5.2), hand 0.32 (2.1), movable finger 0.45; ratio movable finger/hand 1.4, femur/movable finger 1.3, femur/carapace 1.4, chela/carapace 1.9, chela/femur 1.4.
</paragraph>
</subSubSection>
<subSubSection id="C33B7C85FF6955C0FF0C4046FE3CFE7E" pageId="214" pageNumber="215" type="discussion">
<paragraph id="8B9E2F0EFF6955C0FF0C4046FE3CFE7E" blockId="214.[151,1436,151,429]" pageId="214" pageNumber="215">
<emphasis id="B955F31CFF6955C0FF0C4046FEBDFE92" bold="true" box="[199,315,295,320]" pageId="214" pageNumber="215">Remarks.</emphasis>
<taxonomicName id="4C21548DFF6955C0FE8A4046FD1FFE92" box="[321,665,295,320]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="villacarrillo">
<emphasis id="B955F31CFF6955C0FE8A4046FD1FFE92" box="[321,665,295,320]" italics="true" pageId="214" pageNumber="215">Occidenchthonius villacarrillo</emphasis>
</taxonomicName>
does not belong to either of the recognized species-groups within the genus. Comparison with the closely related species
<taxonomicName id="4C21548DFF6955C0FD20402CFC02FEB7" box="[747,900,332,357]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="ambrosiae">
<emphasis id="B955F31CFF6955C0FD20402CFC02FEB7" box="[747,900,332,357]" italics="true" pageId="214" pageNumber="215">O. ambrosiae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C21548DFF6955C0FC5E402CFBBBFEB7" box="[917,1085,332,357]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="bullonorum">
<emphasis id="B955F31CFF6955C0FC5E402CFBBBFEB7" box="[917,1085,332,357]" italics="true" pageId="214" pageNumber="215">O. bullonorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C21548DFF6955C0FBB3402CFAB3FEB7" box="[1144,1333,332,357]" class="Arachnida" family="Chthoniidae" genus="Occidenchthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="felgueraorum" status="sp. nov.">
<emphasis id="B955F31CFF6955C0FBB3402CFAB3FEB7" box="[1144,1333,332,357]" italics="true" pageId="214" pageNumber="215">O. felgueraorum</emphasis>
</taxonomicName>
<emphasis id="B955F31CFF6955C0FAF5402DFA06FEB7" bold="true" box="[1342,1408,332,357]" pageId="214" pageNumber="215">
<taxonomicNameLabel id="A2664E67FF6955C0FAF5402DFA06FEB7" box="[1342,1408,332,357]" pageId="214" pageNumber="215" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is made in the key and under the remarks for those species. The above description incorporates in part that of
<bibRefCitation id="EFB052FFFF6955C0FF5C40F4FE30FE7E" author="Zaragoza" box="[151,438,404,429]" pageId="214" pageNumber="215" refString="Zaragoza, J. A. &amp; Perez, T. (2013) Hypogean pseudoscorpions (Arachnida) from Jaen province (Andalusia, Spain), with descriptions of four new species and a new synonymy. Zootaxa, 3700 (2), 201 - 225." type="journal article" year="2013">Zaragoza &amp; Pérez (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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