464 lines
48 KiB
XML
464 lines
48 KiB
XML
<document id="4A3067D55FFC655617DD25CD2F349C96" ID-DOI="10.5281/zenodo.206798" ID-GBIF-Dataset="f362f4e9-0219-45a1-a498-6f04a5483c9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="206798" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460246210568" checkinUser="plazi" docAuthor="Vila-Farré, Miquel, Sluys, Ronald, Almagro, Ío, Handberg-Thorsager, Mette & Romero, Rafael" docDate="2011" docId="038887CDFF9B7743FF2EBAB26E0C84D6" docLanguage="en" docName="zt02779p038.pdf" docOrigin="Zootaxa 2779" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Phagocata hellenica Vila-Farre & Sluys, sp. nov." docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="FFB1FFB5FF8C7759FFB9BC666D2E866C" masterDocTitle="Freshwater planarians (Platyhelminthes, Tricladida) from the Iberian Peninsula and Greece: diversity and notes on ecology" masterLastPageNumber="38" masterPageNumber="1" pageNumber="24" updateTime="1698241874557" updateUser="plazi">
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<mods:title id="D5BD94EDE4AC61C62CACB950399C7853">Freshwater planarians (Platyhelminthes, Tricladida) from the Iberian Peninsula and Greece: diversity and notes on ecology</mods:title>
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<mods:namePart id="1DF3C7685281C3D8BE29113E8E818779">Vila-Farré, Miquel</mods:namePart>
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<mods:namePart id="AC8531A28C1751AE840C2922A735E6E2">Sluys, Ronald</mods:namePart>
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<mods:namePart id="443D79215B915780118B06B15C4CD31E">Almagro, Ío</mods:namePart>
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<mods:namePart id="1BEED00BF85F84E44D58B8AA2F3D2F04">Handberg-Thorsager, Mette</mods:namePart>
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<mods:namePart id="925064F64FA7FDF5491DB1CA6F6E44F6">Romero, Rafael</mods:namePart>
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<mods:title id="D6D523BAC9B0A5705DF884BDAC3B511D">Zootaxa</mods:title>
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<mods:date id="502F994C0BD6BABDC312FDE9D3663B5D">2011</mods:date>
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<mods:number id="078647470C4C1FDB0DF08110F40454B2">2779</mods:number>
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<mods:identifier id="8A05A2DF7ED65F05F0FACE3029873D79" type="DOI">10.5281/zenodo.206798</mods:identifier>
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<treatment id="038887CDFF9B7743FF2EBAB26E0C84D6" ID-DOI="http://doi.org/10.5281/zenodo.5687266" ID-GBIF-Taxon="119390536" ID-Zenodo-Dep="5687266" LSID="urn:lsid:plazi:treatment:038887CDFF9B7743FF2EBAB26E0C84D6" httpUri="http://treatment.plazi.org/id/038887CDFF9B7743FF2EBAB26E0C84D6" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24">
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<subSubSection id="C33B6550FF9B774EFF2EBAB26CB88163" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8B9E36DBFF9B774EFF2EBAB26FC08082" blockId="23.[151,750,1748,1807]" box="[151,750,1748,1774]" pageId="23" pageNumber="24">
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<heading id="D0D681B7FF9B774EFF2EBAB26FC08082" bold="true" box="[151,750,1748,1774]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B955EAC9FF9B774EFF2EBAB26FC08082" bold="true" box="[151,750,1748,1774]" pageId="23" pageNumber="24">
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<taxonomicName id="4C214D58FF9B774EFF2EBAB26FAD8082" authority="Vila-Farre & Sluys" authorityName="Vila-Farre & Sluys" box="[151,643,1748,1774]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="23" pageNumber="24" phylum="Platyhelminthes" rank="species" species="hellenica" status="sp. nov.">
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<emphasis id="B955EAC9FF9B774EFF2EBAB26CA38082" bold="true" box="[151,397,1748,1774]" italics="true" pageId="23" pageNumber="24">Phagocata hellenica</emphasis>
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Vila-Farré & Sluys
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26657B2FF9B774EFD28BAB26FC08082" box="[657,750,1748,1774]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9E36DBFF9B774EFF2EBA916CB88163" blockId="23.[151,750,1748,1807]" box="[151,406,1783,1807]" pageId="23" pageNumber="24">
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(
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<tableCitation id="C6A30360FF9B774EFF27BA916DDD8163" box="[158,243,1783,1807]" captionStart="TABLE 1" captionStartId="8.[151,235,154,175]" captionTargetBox="[151,1437,266,763]" captionTargetPageId="8" captionText="TABLE 1. List of localities with new species of freshwater planarians in the Iberian Peninsula and Greece. Latitude and longitude are expressed in decimal degrees (Datum European, 1979). Latitude and longitude for “ Canal of Argos, near Argos and Néa Kíos ” are approximations." httpUri="http://table.plazi.org/id/DF5E6653FF847751FF2EBCFC6CFF8683" pageId="23" pageNumber="24" tableUuid="DF5E6653FF847751FF2EBCFC6CFF8683">Table 1</tableCitation>
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,
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<figureCitation id="131A2A5EFF9B774EFEB8BA916C7E8163" box="[257,336,1783,1807]" captionStart="FIGURE 15" captionStartId="24.[151,250,1687,1710]" captionTargetBox="[151,1436,381,1636]" captionTargetId="figure@24.[151,1436,381,1643]" captionTargetPageId="24" captionText="FIGURE 15. Phagocata hellenica, specimens from the Canal of Argos, near Argos and Néa Kíos, Greece. (A) living animal, (B – C) holotype, V. Pl. 6878.4, (B) sagittal section of the copulatory apparatus (anterior to the right), (C) sagittal reconstruction of the copulatory apparatus (anterior to the right)." httpUri="https://zenodo.org/record/206812/files/figure.png" pageId="23" pageNumber="24">Fig. 15</figureCitation>
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A–C)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6550FF9B774EFF2EBB586FCA818B" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8B9E36DBFF9B774EFF2EBB586E318117" blockId="23.[151,1436,1854,2023]" pageId="23" pageNumber="24">
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<emphasis id="B955EAC9FF9B774EFF2EBB586CAD813B" bold="true" box="[151,387,1854,1879]" pageId="23" pageNumber="24">Material examined.</emphasis>
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<typeStatus id="549A8879FF9B774EFE35BB596F0A813B" box="[396,548,1855,1879]" pageId="23" pageNumber="24" type="holotype">HOLOTYPE</typeStatus>
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.
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<collectionCode id="ED30AE1EFF9B774EFD89BB266F408134" box="[560,622,1856,1880]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="23" pageNumber="24">ZMA</collectionCode>
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V.Pl. 6878.4, Canal of Argos, near Argos and Néa Kíos, Peloponnese,
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<collectingCountry id="F336764BFF9B774EFF2EBB056DC78117" box="[151,233,1891,1915]" name="Greece" pageId="23" pageNumber="24">Greece</collectingCountry>
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, 0
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<date id="FF9F101BFF9B774EFEBABB056C878117" box="[259,425,1891,1916]" pageId="23" pageNumber="24" value="2008-08-08">8 August 2008</date>
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, sagittal sections on seven slides.
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</paragraph>
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<paragraph id="8B9E36DBFF9B774EFF7EBBE16FCA818B" blockId="23.[151,1436,1854,2023]" pageId="23" pageNumber="24">
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<typeStatus id="549A8879FF9B774EFF7EBBE16C4B81F3" box="[199,357,1927,1951]" pageId="23" pageNumber="24" type="paratype">PARATYPES</typeStatus>
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.
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<collectionCode id="ED30AE1EFF9B774EFED6BBEE6C8381CC" box="[367,429,1928,1952]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="23" pageNumber="24">ZMA</collectionCode>
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V.Pl. 6878.1, ibid., horizontal sections on three slides; V.Pl. 6878.2, ibid., sagittal sections on five slides; V.Pl. 6878.3, ibid., sagittal sections on four slides; V.Pl. 6878.4, ibid., sagittal sections on four slides; V.Pl. 6878.5, ibid., horizontal sections on two slides.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6550FF947741FF7EBCFE694D86C3" box="[199,1123,151,176]" pageId="24" pageNumber="25" type="etymology">
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<paragraph id="8B9E36DBFF947741FF7EBCFE694D86C3" blockId="24.[151,1436,151,319]" box="[199,1123,151,176]" pageId="24" pageNumber="25">
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<emphasis id="B955EAC9FF947741FF7EBCFE6C6086DC" bold="true" box="[199,334,152,176]" pageId="24" pageNumber="25">Etymology.</emphasis>
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The specific epithet is based on the Greek
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<emphasis id="B955EAC9FF947741FC89BCFE6E8C86DC" box="[816,930,152,176]" italics="true" pageId="24" pageNumber="25">hellenikos</emphasis>
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, meaning Greek.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33B6550FF947741FF7EBCDD6C7C8753" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph id="8B9E36DBFF947741FF7EBCDD6C7C8753" blockId="24.[151,1436,151,319]" pageId="24" pageNumber="25">
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<emphasis id="B955EAC9FF947741FF7EBCDD6C6F86B8" bold="true" box="[199,321,187,212]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
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With respect to anatomical features,
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<taxonomicName id="4C214D58FF947741FD50BCDA6EFC86B8" box="[745,978,188,212]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="hellenica">
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<emphasis id="B955EAC9FF947741FD50BCDA6EFC86B8" box="[745,978,188,212]" italics="true" pageId="24" pageNumber="25">Phagocata hellenica</emphasis>
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</taxonomicName>
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can be distinguished from its congeners by the combined presence of (1) a short common vas deferens opening at the tip of a short projection into the very proximal part of the ejaculatory duct and (2) a plump plug of cells projecting into a slightly more distal part of the ejaculatory duct.
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</paragraph>
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</subSubSection>
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||
<caption id="DF5E6653FF947741FF2EBAF16FA08086" httpUri="https://zenodo.org/record/206812/files/figure.png" pageId="24" pageNumber="25" targetBox="[151,1436,381,1636]" targetPageId="24">
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<paragraph id="8B9E36DBFF947741FF2EBAF16FA08086" blockId="24.[151,1436,1687,1770]" pageId="24" pageNumber="25">
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<emphasis id="B955EAC9FF947741FF2EBAF16C0E80C0" bold="true" box="[151,288,1687,1710]" pageId="24" pageNumber="25">FIGURE 15.</emphasis>
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<taxonomicName id="4C214D58FF947741FE9EBAF16CD680C1" box="[295,504,1687,1709]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="24" pageNumber="25" phylum="Platyhelminthes" rank="species" species="hellenica">
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<emphasis id="B955EAC9FF947741FE9EBAF16CD680C1" box="[295,504,1687,1709]" italics="true" pageId="24" pageNumber="25">Phagocata hellenica</emphasis>
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</taxonomicName>
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, specimens from the Canal of Argos, near Argos and Néa Kíos, Greece. (A) living animal, (B–C) holotype, V.Pl. 6878.4, (B) sagittal section of the copulatory apparatus (anterior to the right), (C) sagittal reconstruction of the copulatory apparatus (anterior to the right).
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</paragraph>
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</caption>
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<subSubSection id="C33B6550FF947740FF7EBB716FDA82C9" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
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<paragraph id="8B9E36DBFF947741FF7EBB71694C8116" blockId="24.[151,1436,1815,1988]" pageId="24" pageNumber="25">
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<emphasis id="B955EAC9FF947741FF7EBB716C77815C" bold="true" box="[199,345,1815,1840]" pageId="24" pageNumber="25">Description.</emphasis>
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Preserved specimens measure up to 7.1 x
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<quantity id="4CD99B3EFF947741FC8BBB716EA58143" box="[818,907,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="25" unit="mm" value="2.2">2.2 mm</quantity>
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. The dorsal surface is transparent and therefore the intestinal branches can be seen (
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<figureCitation id="131A2A5EFF947741FD91BB5A6F568138" box="[552,632,1852,1876]" captionStart="FIGURE 15" captionStartId="24.[151,250,1687,1710]" captionTargetBox="[151,1436,381,1636]" captionTargetId="figure@24.[151,1436,381,1643]" captionTargetPageId="24" captionText="FIGURE 15. Phagocata hellenica, specimens from the Canal of Argos, near Argos and Néa Kíos, Greece. (A) living animal, (B – C) holotype, V. Pl. 6878.4, (B) sagittal section of the copulatory apparatus (anterior to the right), (C) sagittal reconstruction of the copulatory apparatus (anterior to the right)." httpUri="https://zenodo.org/record/206812/files/figure.png" pageId="24" pageNumber="25">Fig. 15</figureCitation>
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A). The ventral surface is also devoid of pigment. A small narrowing of the body width is present at the level of the eyes; the tail is rounded or bluntly pointed.
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</paragraph>
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<paragraph id="8B9E36DBFF947741FF7CBBE16EEE81A8" blockId="24.[151,1436,1815,1988]" pageId="24" pageNumber="25">The eyes (maximum eye cup diameter 58 µm) are located relatively far from the frontal margin. The number of retinal cells in each pigment cup could not be determined with certainty.</paragraph>
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<paragraph id="8B9E36DBFF957740FF7CBCF16E9C87E6" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">
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The end of the anterior intestinal branch forms an extension that reaches anterior to the eyes. The cylindrical pharynx is situated in the middle of the body and measures about 1/6th of the body length. The outer epithelium of the pharynx is underlain by a layer of longitudinal muscle fibres, followed by a layer of circular muscles. The inner epithelium is underlain by a thick layer of circular muscles, followed by a layer of longitudinal muscle fibres. The mouth is situated in the posterior portion of the pharyngeal pocket and opens to the exterior underneath the copulatory bursa. In specimen V.Pl. 6878.4, the mouth is situated
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<quantity id="4CD99B3EFF957740FC96BD296EBD870B" box="[815,915,335,359]" metricMagnitude="-3" metricUnit="m" metricValue="3.28" pageId="25" pageNumber="26" unit="mm" value="3.28">3.28 mm</quantity>
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from the anterior end of the body and 106 μm from the gonopore (the total body length of this specimen is
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<quantity id="4CD99B3EFF957740FC84BD146E8B87E6" box="[829,933,370,394]" metricMagnitude="-3" metricUnit="m" metricValue="5.68" pageId="25" pageNumber="26" unit="mm" value="5.68">5.68 mm</quantity>
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).
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</paragraph>
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<paragraph id="8B9E36DBFF957740FF7CBDF1684D849E" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">
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The oval-shaped testes follicles occupy approximately one-half of the dorsoventral diameter of the body and extend from a short distance behind the ovaries into the posterior end of the body. The testes are mainly ventral but some are dorsal. At the level of the penis papilla, the vasa deferentia curve dorsally and, subsequently, recurve before penetrating the wall of the penis bulb. Once they have penetrated the penis bulb, the vasa deferentia open separately into the ejaculatory duct. Within the penis papilla, each vas deferens is surrounded by a thick layer of circular muscles fibres. The ducts fuse to form a very short common vas deferens, which opens at the tip of a small projection into the proximal part of the ejaculatory duct. The ejaculatory duct runs centrally through the penis and opens at the tip of the papilla. The ejaculatory duct is divided histologically into two sections (
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<figureCitation id="131A2A5EFF957740FB09BEF4682E84C6" box="[1200,1280,658,682]" captionStart="FIGURE 15" captionStartId="24.[151,250,1687,1710]" captionTargetBox="[151,1436,381,1636]" captionTargetId="figure@24.[151,1436,381,1643]" captionTargetPageId="24" captionText="FIGURE 15. Phagocata hellenica, specimens from the Canal of Argos, near Argos and Néa Kíos, Greece. (A) living animal, (B – C) holotype, V. Pl. 6878.4, (B) sagittal section of the copulatory apparatus (anterior to the right), (C) sagittal reconstruction of the copulatory apparatus (anterior to the right)." httpUri="https://zenodo.org/record/206812/files/figure.png" pageId="25" pageNumber="26">Fig. 15</figureCitation>
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B, C). A short proximal, anterior portion is lined with very tall cells, forming a plug that fills almost the entire lumen of the canal. The second, distal section is lined by the usual epithelium, which is pierced by abundant erythrophilic glands.
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</paragraph>
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<paragraph id="8B9E36DBFF957740FF7CBE99691C85EE" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">The epithelium of the large conical penis papilla is underlain with a layer of thick circular muscle fibres; the epithelium is thinner at the basal wall of the papilla. The weak penis bulb is formed by intermingled longitudinal and circular muscle layers. The dorsal section of the male atrium is lined with thick nucleated epithelium, underlain by a thin layer of circular muscle fibres, followed by a layer of longitudinal fibres.</paragraph>
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<paragraph id="8B9E36DBFF957740FF7CBFF76FDA82C9" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">
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The paired, globular ovaries occupy about 1/3rd of the dorsoventral diameter of the body and are located a short distance behind the brain. The oviducts curve mediodorsally at the level of the penis papilla, where they unite to form a common oviduct (
|
||
<figureCitation id="131A2A5EFF957740FE0CBFBF6F29859D" box="[437,519,985,1009]" captionStart="FIGURE 15" captionStartId="24.[151,250,1687,1710]" captionTargetBox="[151,1436,381,1636]" captionTargetId="figure@24.[151,1436,381,1643]" captionTargetPageId="24" captionText="FIGURE 15. Phagocata hellenica, specimens from the Canal of Argos, near Argos and Néa Kíos, Greece. (A) living animal, (B – C) holotype, V. Pl. 6878.4, (B) sagittal section of the copulatory apparatus (anterior to the right), (C) sagittal reconstruction of the copulatory apparatus (anterior to the right)." httpUri="https://zenodo.org/record/206812/files/figure.png" pageId="25" pageNumber="26">Fig. 15</figureCitation>
|
||
C). This opens into the posterodorsal section of the male atrium. Shell glands discharge into the common oviduct and also into the distal portion of the oviducts. The large copulatory bursa lies between the hind wall of the pharyngeal pocket and the copulatory apparatus and is lined with tall, vacuolated cells. The long bursal canal curves dorsally to the male atrium. The lining epithelium of the wide bursal canal bears distinct cilia and consists of nucleated cells. The canal is surrounded by a subepithelial layer of circular muscle fibres, followed by a thin layer of longitudinal muscle fibres.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33B6550FF957743FF7EB8D76E0C84D6" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26" type="discussion">
|
||
<paragraph id="8B9E36DBFF957740FF7EB8D76E548311" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">
|
||
<emphasis id="B955EAC9FF957740FF7EB8D76C6282A6" bold="true" box="[199,332,1201,1226]" pageId="25" pageNumber="26">Discussion.</emphasis>
|
||
The most characteristic anatomical traits of these specimens is a plug of cells in the ejaculatory duct and the short common vas deferens that opens at the tip of a short projection into the most anterior part of the ejaculatory duct. Therefore, we will restrict our comparative discussion to species that have such a plug (cf. Sluys
|
||
<emphasis id="B955EAC9FF957740FF2EB9796DFF835A" box="[151,209,1310,1334]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
1995):
|
||
<taxonomicName id="4C214D58FF957740FE95B9786FBB8359" authority="Vejdosky, 1883" authorityName="Vejdosky" authorityYear="1883" box="[300,661,1309,1334]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="albissima">
|
||
<emphasis id="B955EAC9FF957740FE95B9786CE9835A" box="[300,455,1310,1334]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FE95B9786C60835A" box="[300,334,1310,1334]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. albissima
|
||
</emphasis>
|
||
(Vejdoský, 1883)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FD1DB9786E65835A" box="[676,843,1310,1334]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="armeniaca">
|
||
<emphasis id="B955EAC9FF957740FD1DB9786E65835A" box="[676,843,1310,1334]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FD1DB9786FE8835A" box="[676,710,1310,1334]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. armeniaca
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FCE2B9786EC0835A" box="[859,1006,1310,1334]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="undulata">
|
||
<emphasis id="B955EAC9FF957740FCE2B9786EC0835A" box="[859,1006,1310,1334]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FCE2B9786E53835A" box="[859,893,1310,1334]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. undulata
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FC47B97868498359" authority="Stankovic, 1926" authorityName="Stankovic" authorityYear="1926" box="[1022,1383,1309,1334]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="bosniaca">
|
||
<emphasis id="B955EAC9FF957740FC47B97869BC835A" box="[1022,1170,1310,1334]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FC47B978690E835A" box="[1022,1056,1310,1334]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. bosniaca
|
||
</emphasis>
|
||
(Stankoviċ, 1926)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FACFB97B6CFF8335" authority="Komarek, 1919" authorityName="Komarek" authorityYear="1919" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="illyrica">
|
||
<emphasis id="B955EAC9FF957740FACFB97B6DC68335" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FACFB97B68B68359" box="[1398,1432,1309,1333]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. illyrica
|
||
</emphasis>
|
||
(Komárek, 1919)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FE5CB9276EAE8335" authority="Stankovic, 1926" authorityName="Stankovic" authorityYear="1926" box="[485,896,1345,1370]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="macedonica">
|
||
<emphasis id="B955EAC9FF957740FE5CB9276F8F8335" box="[485,673,1345,1369]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FE5CB9276F298335" box="[485,519,1345,1369]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. macedonica
|
||
</emphasis>
|
||
(Stankoviċ, 1926)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FC2FB927691A8335" box="[918,1076,1345,1369]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="maculata">
|
||
<emphasis id="B955EAC9FF957740FC2FB927691A8335" box="[918,1076,1345,1369]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FC2FB9276E978335" box="[918,953,1345,1369]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. maculata
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FBF3B9276C7E8311" authority="Stankovic & Komarek, 1927" authorityName="Stankovic & Komarek" authorityYear="1927" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="dalmatica">
|
||
<emphasis id="B955EAC9FF957740FBF3B92769C18335" box="[1098,1263,1345,1369]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FBF3B92769428335" box="[1098,1132,1345,1369]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. dalmatica
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB04B2AFF957740FABFB9276C668311" author="Stankovic" pageId="25" pageNumber="26" refString="Stankovic, S. & Komarek, J. (1927) Die Sußwasser-Tricladen des Westbalkans und die zoogeographischen probleme dieser gegend. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 53, 591 - 674 + 9 pls." type="journal article" year="1927">Stankoviċ & Komárek, 1927</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FEE5B9006CD28312" box="[348,508,1382,1406]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF957740FEE5B9006CD28312" box="[348,508,1382,1406]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FEE5B9006C508312" box="[348,382,1382,1406]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FDB2B9006E588311" authority="Reisinger, 1960" authorityName="Reisinger" authorityYear="1960" box="[523,886,1381,1406]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="stankovici">
|
||
<emphasis id="B955EAC9FF957740FDB2B9006F878312" box="[523,681,1382,1406]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FDB2B9006F038312" box="[523,557,1382,1406]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. stankovici
|
||
</emphasis>
|
||
(Reisinger, 1960)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E36DBFF957740FF7CB9EC69E78039" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">
|
||
In
|
||
<taxonomicName id="4C214D58FF957740FF5DB9EF6C5283CD" box="[228,380,1417,1441]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="albissima">
|
||
<emphasis id="B955EAC9FF957740FF5DB9EF6C5283CD" box="[228,380,1417,1441]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FF5DB9EF6C2883CD" box="[228,262,1417,1441]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. albissima
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D58FF957740FE0DB9EF6F7783CD" box="[436,601,1417,1441]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="armeniaca">
|
||
<emphasis id="B955EAC9FF957740FE0DB9EF6F7783CD" box="[436,601,1417,1441]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FE0DB9EF6CF883CD" box="[436,470,1417,1441]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. armeniaca
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, the presumed plug of cells merely consists of an elevated, papillated epithelium. Thus, the plug is very reduced, if present at all. The body margin is folded in
|
||
<taxonomicName id="4C214D58FF957740FBDAB9C869DD83AA" box="[1123,1267,1454,1478]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="undulata">
|
||
<emphasis id="B955EAC9FF957740FBDAB9C869DD83AA" box="[1123,1267,1454,1478]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FBDAB9C869AB83AA" box="[1123,1157,1454,1478]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. undulata
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, but not in
|
||
<taxonomicName id="4C214D58FF957740FACFB9C86DD08385" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="hellenica">
|
||
<emphasis id="B955EAC9FF957740FACFB9C86DD08385" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FACFB9C868B683AA" box="[1398,1432,1454,1478]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. hellenica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C214D58FF957740FE90B9B76C958385" box="[297,443,1489,1513]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="bosniaca">
|
||
<emphasis id="B955EAC9FF957740FE90B9B76C958385" box="[297,443,1489,1513]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FE90B9B76C628385" box="[297,332,1489,1513]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. bosniaca
|
||
</emphasis>
|
||
</taxonomicName>
|
||
the vasa deferentia run through a very long intrapenial papilla-like structure. In contrast, the projection into the lumen of the penis papilla is very short in Greek
|
||
<taxonomicName id="4C214D58FF957740FC05B990697E8062" box="[956,1104,1526,1550]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="hellenica">
|
||
<emphasis id="B955EAC9FF957740FC05B990697E8062" box="[956,1104,1526,1550]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FC05B9906EF08062" box="[956,990,1526,1550]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. hellenica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
specimens. The penis papilla is very elongated in
|
||
<taxonomicName id="4C214D58FF957740FEC1BA7F6CD8805D" box="[376,502,1561,1585]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="illyrica">
|
||
<emphasis id="B955EAC9FF957740FEC1BA7F6CD8805D" box="[376,502,1561,1585]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FEC1BA7F6CB5805D" box="[376,411,1561,1585]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. illyrica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D58FF957740FD92BA7F6FF1805D" box="[555,735,1561,1585]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="macedonica">
|
||
<emphasis id="B955EAC9FF957740FD92BA7F6FF1805D" box="[555,735,1561,1585]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FD92BA7F6F63805D" box="[555,589,1561,1585]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. macedonica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, in contrast to the shorter papilla in the Greek animals.
|
||
<taxonomicName id="4C214D58FF957740FAF0BA7F6C18803A" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="maculata">
|
||
<emphasis id="B955EAC9FF957740FAF0BA7F6C18803A" italics="true" pageId="25" pageNumber="26">Phagocata maculata</emphasis>
|
||
</taxonomicName>
|
||
is a pigmented species, whereas the
|
||
<taxonomicName id="4C214D58FF957740FD6BBA586E48803A" box="[722,870,1598,1622]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="hellenica">
|
||
<emphasis id="B955EAC9FF957740FD6BBA586E48803A" box="[722,870,1598,1622]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FD6BBA586FDA803A" box="[722,756,1598,1622]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. hellenica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
animals are devoid of pigment.
|
||
</paragraph>
|
||
<paragraph id="8B9E36DBFF957740FF7CBA07698B818D" blockId="25.[151,1437,151,2018]" pageId="25" pageNumber="26">
|
||
The organization and histology of the reproductive organs of our specimens of
|
||
<taxonomicName id="4C214D58FF957740FBC8BA0768248015" box="[1137,1290,1633,1657]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="hellenica">
|
||
<emphasis id="B955EAC9FF957740FBC8BA0768248015" box="[1137,1290,1633,1657]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FBC8BA0769BD8015" box="[1137,1171,1633,1657]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. hellenica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
corresponds closely to the scheme of the copulatory apparatuses of
|
||
<taxonomicName id="4C214D58FF957740FCAEBAE06E9980F2" box="[791,951,1670,1694]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="dalmatica">
|
||
<emphasis id="B955EAC9FF957740FCAEBAE06E9980F2" box="[791,951,1670,1694]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FCAEBAE06E1480F2" box="[791,826,1670,1694]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. dalmatica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C214D58FF957740FC71BAE0694580F2" box="[968,1131,1670,1694]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF957740FC71BAE0694580F2" box="[968,1131,1670,1694]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FC71BAE06EC480F2" box="[968,1002,1670,1694]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B955EAC9FF957740FB1EBAE0687F80F2" box="[1191,1361,1670,1694]" italics="true" pageId="25" pageNumber="26">
|
||
<taxonomicName id="4C214D58FF957740FB1EBAE0686280F2" box="[1191,1356,1670,1694]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="stankovici">
|
||
<collectingCountry id="F336764BFF957740FB1EBAE069E780F2" box="[1191,1225,1670,1694]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. stankovici
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
In his monograph on Lake Ohrid planarians,
|
||
<bibRefCitation id="EFB04B2AFF957740FDF7BACF6FF380AD" author="Kenk" box="[590,733,1705,1729]" pageId="25" pageNumber="26" refString="Kenk, R. (1978) The planarians (Turbellaria: Tricladida Paludicola) of Lake Ohrid in Macedonia. Smithsonian Contributions to Zoology 280, 1 - 55." type="journal article" year="1978">Kenk (1978)</bibRefCitation>
|
||
noted that the taxonomic position of these three nominal species is still an open question. After a brief summary of the opinions of various authors,
|
||
<bibRefCitation id="EFB04B2AFF957740FBE4BAAB69C48089" author="Kenk" box="[1117,1258,1741,1765]" pageId="25" pageNumber="26" refString="Kenk, R. (1978) The planarians (Turbellaria: Tricladida Paludicola) of Lake Ohrid in Macedonia. Smithsonian Contributions to Zoology 280, 1 - 55." type="journal article" year="1978">Kenk (1978)</bibRefCitation>
|
||
concluded that: “It seems to me quite probable that all three will ultimately prove to be one species,
|
||
<taxonomicName id="4C214D58FF957740FB1CBA976C868141" authority="Stankovic & Komarec" authorityName="Stankovic & Komarec" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="dalmatica">
|
||
<emphasis id="B955EAC9FF957740FB1CBA9768B28165" box="[1189,1436,1777,1801]" italics="true" pageId="25" pageNumber="26">Phagocata dalmatica</emphasis>
|
||
(Stankoviċ & Komárec)
|
||
</taxonomicName>
|
||
.” The size, shape and body colouration of our specimens is similar to that of
|
||
<taxonomicName id="4C214D58FF957740FB45BB7368B28141" box="[1276,1436,1813,1837]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF957740FB45BB7368B28141" box="[1276,1436,1813,1837]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FB45BB7368308141" box="[1276,1310,1813,1837]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C214D58FF957740FF7FBB5F6C4A813D" box="[198,356,1849,1873]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="stankovici">
|
||
<emphasis id="B955EAC9FF957740FF7FBB5F6C4A813D" box="[198,356,1849,1873]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FF7FBB5F6DC6813D" box="[198,232,1849,1873]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. stankovici
|
||
</emphasis>
|
||
</taxonomicName>
|
||
from Lake Ohrid. The size of the pharynx, arrangement of the testes, and shape and structure of the penis papilla and the common oviduct are also similar. In our specimens, the anterior intestinal branch forms an extension that reaches anterior to the eyes. This is a characteristic feature of
|
||
<taxonomicName id="4C214D58FF957740FC4FBBE769B881F5" box="[1014,1174,1921,1945]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="stankovici">
|
||
<emphasis id="B955EAC9FF957740FC4FBBE769B881F5" box="[1014,1174,1921,1945]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FC4FBBE7693781F5" box="[1014,1049,1921,1945]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. stankovici
|
||
</emphasis>
|
||
</taxonomicName>
|
||
that is absent from
|
||
<taxonomicName id="4C214D58FF957740FACFBBE76C2481D2" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF957740FACFBBE76C2481D2" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FACFBBE768B681F5" box="[1398,1432,1921,1945]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
. Recent electrophoretic results (
|
||
<bibRefCitation id="EFB04B2AFF957740FDCBBBC36E5581D1" author="Krstanovski" box="[626,891,1957,1982]" pageId="25" pageNumber="26" refString="Krstanovski, Z., Tasevska, O., Sywula, T., Sell, J. & Kretowicz, T. (2004) Taxonomic statute of Phagocata stankovici and Phagocata ochridana (Turbellaria: Tricladida: Paludicola) established with enzyme electrophoresis. Balwois 2004. Available from: http: // balwois. com / balwois / administration / full _ paper / ffp- 6 p- 312. pdf (16 August 2010)" type="journal article" year="2004">
|
||
Krstanovski
|
||
<emphasis id="B955EAC9FF957740FCBBBBC16E1481D2" box="[770,826,1958,1982]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
) have been interpreted to favour a separate taxonomic status for
|
||
<taxonomicName id="4C214D58FF957740FEEBBBAC6CDE818E" box="[338,496,1994,2018]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="stankovici">
|
||
<emphasis id="B955EAC9FF957740FEEBBBAC6CDE818E" box="[338,496,1994,2018]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FEEBBBAC6C5A818E" box="[338,372,1994,2018]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. stankovici
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, which is frequently considered a synonym of
|
||
<taxonomicName id="4C214D58FF957740FC47BBAC69B0818E" box="[1022,1182,1994,2018]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="25" pageNumber="26" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF957740FC47BBAC69B0818E" box="[1022,1182,1994,2018]" italics="true" pageId="25" pageNumber="26">
|
||
<collectingCountry id="F336764BFF957740FC47BBAC690E818E" box="[1022,1056,1994,2018]" name="Philippines" pageId="25" pageNumber="26">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9E36DBFF967743FF7CBCF16E208773" blockId="26.[151,1436,151,698]" pageId="26" pageNumber="27">
|
||
The penis bulb is small in the
|
||
<taxonomicName id="4C214D58FF967743FD98BCFE6F9186DC" box="[545,703,152,176]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="dalmatica">
|
||
<emphasis id="B955EAC9FF967743FD98BCFE6F9186DC" box="[545,703,152,176]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FD98BCFE6F6D86DC" box="[545,579,152,176]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. dalmatica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
group. Its musculature is rather weak and found mainly near its periphery. In our specimens, we found reticulated mesenchymal tissue in the penis bulb that was not described in the former species. This tissue may well be present in these species, but may not have been adequately described in the literature, due to its small size or inconspicuousness.
|
||
</paragraph>
|
||
<paragraph id="8B9E36DBFF967743FF7CBD486E0C84D6" blockId="26.[151,1436,151,698]" pageId="26" pageNumber="27">
|
||
In addition to these differences, some specimens of
|
||
<taxonomicName id="4C214D58FF967743FCB6BD4B6E9E8729" box="[783,944,301,325]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF967743FCB6BD4B6E9E8729" box="[783,944,301,325]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FCB6BD4B6E1F8729" box="[783,817,301,325]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
have a characteristic infranucleated ephitelium in the posterior section of the atrium and bursal. This does not appear to be present in the fully nucleated epithelium of the atrium and bursal canal of the new material of
|
||
<taxonomicName id="4C214D58FF967743FCFCBD1E6EF787FC" box="[837,985,376,400]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="hellenica">
|
||
<emphasis id="B955EAC9FF967743FCFCBD1E6EF787FC" box="[837,985,376,400]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FCFCBD1E6E4987FC" box="[837,871,376,400]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. hellenica
|
||
</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F336764BFF967743FBA7BD11695F87E3" box="[1054,1137,375,399]" name="Greece" pageId="26" pageNumber="27">Greece</collectingCountry>
|
||
. After careful examination of the description of
|
||
<taxonomicName id="4C214D58FF967743FE3CBDFB6F0987D9" box="[389,551,413,437]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF967743FE3CBDFB6F0987D9" box="[389,551,413,437]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FE3CBDFB6C8987D9" box="[389,423,413,437]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and of a specimen of this species from lake Ohrid in the Zoological Museum Amsterdam collection we noted that our Greek animals differ from
|
||
<taxonomicName id="4C214D58FF967743FC1DBDA5696987B7" box="[932,1095,451,475]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF967743FC1DBDA5696987B7" box="[932,1095,451,475]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FC1DBDA56EE887B7" box="[932,966,451,475]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
and from any other currently known species of the genus
|
||
<taxonomicName id="4C214D58FF967743FE6BBD8E6F65846C" box="[466,587,488,512]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="genus">
|
||
<emphasis id="B955EAC9FF967743FE6BBD8E6F65846C" box="[466,587,488,512]" italics="true" pageId="26" pageNumber="27">Phagocata</emphasis>
|
||
</taxonomicName>
|
||
in two features: the combined presence of (1) a short common vas deferens opening at the tip of a short projection into the very proximal part of the ejaculatory duct and (2) a plump plug of cells that project into a slightly more distal part of the ejaculatory duct. In
|
||
<taxonomicName id="4C214D58FF967743FC76BE5569418427" box="[975,1135,563,587]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF967743FC76BE5569418427" box="[975,1135,563,587]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FC76BE556EDF8427" box="[975,1009,563,587]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, the short projection of the common vas deferens is absent, while the plug of cells in the ejaculatory duct of this species is always elongated and pointed. Therefore, our Greek specimens represent a new species and do not belong to
|
||
<taxonomicName id="4C214D58FF967743FB0DBE1B687984F9" box="[1204,1367,637,661]" class="Rhabditophora" family="Planariidae" genus="Phagocata" kingdom="Animalia" order="Seriata" pageId="26" pageNumber="27" phylum="Platyhelminthes" rank="species" species="ochridana">
|
||
<emphasis id="B955EAC9FF967743FB0DBE1B687984F9" box="[1204,1367,637,661]" italics="true" pageId="26" pageNumber="27">
|
||
<collectingCountry id="F336764BFF967743FB0DBE1B69F884F9" box="[1204,1238,637,661]" name="Philippines" pageId="26" pageNumber="27">Ph</collectingCountry>
|
||
. ochridana
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, even though this species has been reported in mainland
|
||
<collectingCountry id="F336764BFF967743FD72BEC46E3084D6" box="[715,798,674,698]" name="Greece" pageId="26" pageNumber="27">Greece</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |