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<document id="66C83CD599EC6430D56808250F6E9F9D" ID-CLB-Dataset="99513" ID-DOI="10.11646/zootaxa.5165.4.5" ID-GBIF-Dataset="bd8c5821-b532-4539-af24-856a62ba7b27" ID-ISSN="1175-5326" ID-Zenodo-Dep="6854024" ID-ZooBank="F108240E-FE0B-4832-96CB-F0540CB338E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658127318027" checkinUser="plazi" docAuthor="Zhou, Jin-Jin, Achterberg, Kees Van, Tang, Pu &amp; Chen, Xue-Xin" docDate="2022" docId="038887803102054BFF619ADAFDD07B3C" docLanguage="en" docName="zootaxa.5165.4.5.pdf" docOrigin="Zootaxa 5165 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chelonus (Megachelonus) macrodontus Zhou &amp; Achterberg &amp; Tang &amp; Chen 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="540" masterDocId="FFB1FFF831090546FFF69A4DFFCD7D6E" masterDocTitle="A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China" masterLastPageNumber="547" masterPageNumber="527" pageNumber="538" updateTime="1699360222658" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F4D8B600E54DD49B512E21A6074FE7FB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="76F36AFC7F40345E4B46B9BFDAB02FE3">
<mods:title id="34B9635B3EE7C6C9E144C0F6EC761EFC">A review of the subgenera Arichelonus, Megachelonus and Scabrichelonus of the genus Chelonus Panzer (Braconidae: Cheloninae) with description of five new species from China</mods:title>
</mods:titleInfo>
<mods:name id="7C69390416362B78D3C10DC36D302756" type="personal">
<mods:role id="12EFA287B9B071E330350916C865738B">
<mods:roleTerm id="0F685FB399CF8A212338C544A6574915">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ABA0EC36B2822309FDCDF40AE8FB4D07">Zhou, Jin-Jin</mods:namePart>
<mods:nameIdentifier id="CC1E95960C2F037A9641948EA2BB6D9C" type="ORCID">0000-0002-1563-8408</mods:nameIdentifier>
<mods:affiliation id="1081232D21B10010DF9D979EA18E3A86">Hainan Institute, Zhejiang University, Sanya 572000, China &amp; Guangdong Laboratory for Lingnan Modern Agriculture, Guangzhou 510642, China &amp; auzhou @ zju. edu. cn; https: // orcid. org / 0000 - 0002 - 1563 - 8408</mods:affiliation>
<mods:nameIdentifier id="D326B7F07F34E9A14CB685AD0D7724E5" type="email">auzhou@zju.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="9D63F726D5F9342F5651673DC58A774E" type="personal">
<mods:role id="3BCFBE67715BA13D45E7CDB59AEED555">
<mods:roleTerm id="CC66F87E719FEBA938FBDFE34592B978">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D67C25FE716A4E7E2B29BA96F288B760">Achterberg, Kees Van</mods:namePart>
<mods:nameIdentifier id="BACD39BA42E3C363B7344B1C98263FD5" type="ORCID">0000-0002-6495-4853</mods:nameIdentifier>
<mods:affiliation id="902A506041E1009029D9068AD08142E4">Guangdong Laboratory for Lingnan Modern Agriculture, Guangzhou 510642, China &amp; kees @ vanachterberg. org; https: // orcid. org / 0000 - 0002 - 6495 - 4853</mods:affiliation>
<mods:nameIdentifier id="CF7B9BE350AD9175732A861BD6CC5400" type="email">kees@vanachterberg.org</mods:nameIdentifier>
</mods:name>
<mods:name id="08239487F52BE0807DAF1E1368B0E333" type="personal">
<mods:role id="340C22AD3483ABA6748FF9E4B8BCB59F">
<mods:roleTerm id="28C8FDB0BB93289B8C852592535A06D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="119FED7C9AC110E6F56A3189D415A3E3">Tang, Pu</mods:namePart>
<mods:affiliation id="D484F5A75C02E93AB47CCB6B8D00DFFC">Guangdong Laboratory for Lingnan Modern Agriculture, Guangzhou 510642, China &amp; State Key Lab of Rice Biology, Zhejiang University, Hangzhou 310058, China &amp; Ministry of Agriculture Key Lab of Molecular Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China Zhejiang Provincial Key Laboratory of Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
</mods:name>
<mods:name id="C232B23982191AB649D1BB7CCE907A33" type="personal">
<mods:role id="F7ED5768907AE0072D8233C3523E6AA8">
<mods:roleTerm id="67B7D6F8934C9B83638242F666BB150F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="91F5A70BBF5EE6B59F408AD0E09E8D8B">Chen, Xue-Xin</mods:namePart>
<mods:nameIdentifier id="99BA87792A368EFBB2640E6D7EB6742A" type="ORCID">0000-0002-9109-8853</mods:nameIdentifier>
<mods:affiliation id="83516C15CA70BE294C6D0ABB65816A93">Hainan Institute, Zhejiang University, Sanya 572000, China &amp; Guangdong Laboratory for Lingnan Modern Agriculture, Guangzhou 510642, China &amp; State Key Lab of Rice Biology, Zhejiang University, Hangzhou 310058, China &amp; Ministry of Agriculture Key Lab of Molecular Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China Zhejiang Provincial Key Laboratory of Biology of Crop Pathogens and Insects, Zhejiang University, Hangzhou 310058, China Institute of Insect Sciences, College of Agriculture and Biotechnology, Zhejiang University, Hangzhou 310058, China &amp; xxchen @ zju. edu. cn; https: // orcid. org / 0000 - 0002 - 9109 - 8853</mods:affiliation>
<mods:nameIdentifier id="1AB46374E913C15162E8502891724EAB" type="email">xxchen@zju.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="35FCEB09ECA43B1DF483821BA3FA3433">text</mods:typeOfResource>
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<mods:titleInfo id="6DDCBB5CDFA9192FCCD4A7FD1ABB1C56">
<mods:title id="278ECC4D0B0A3A3D98DBE463C045E91C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D393F6B8C23FF96A320DCA86B2FF3211">
<mods:date id="6DE65CEF83C432974DD5CC87B4E70636">2022</mods:date>
<mods:detail id="0A6AFE91784C5142F311A528417A42FE" type="pubDate">
<mods:number id="A84AFB014E1FAC89618905DC3985F110">2022-07-18</mods:number>
</mods:detail>
<mods:detail id="6269150F7DB64F0656436B6562491556" type="volume">
<mods:number id="811713D7CAC83548F3181CB96A17C1AC">5165</mods:number>
</mods:detail>
<mods:detail id="778D46506503F8BB3A2D553278EF8FB3" type="issue">
<mods:number id="E5FB00086412F032C0330037EFC4B78A">4</mods:number>
</mods:detail>
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<mods:start id="4EC30B8CA51E0ABB10BCFD4DDBC3395B">527</mods:start>
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</mods:extent>
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<mods:classification id="93C7FEAAA3424F5B084AF3FE34BB6EED">journal article</mods:classification>
<mods:identifier id="54A28876E338BF2316B21DAD859E4DFC" type="CLB-Dataset">99513</mods:identifier>
<mods:identifier id="850804C1D2B67C5F9E8B739EDFAA6A48" type="DOI">10.11646/zootaxa.5165.4.5</mods:identifier>
<mods:identifier id="9DF3DEF76647D603DAE8EAD1BE3D4619" type="GBIF-Dataset">bd8c5821-b532-4539-af24-856a62ba7b27</mods:identifier>
<mods:identifier id="B2E40DDD9F8243BABF95AB20CD491501" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1E3D42ADD07005C195453ACBD30592C0" type="Zenodo-Dep">6854024</mods:identifier>
<mods:identifier id="1977082563B818C855125A89950056C1" type="ZooBank">F108240E-FE0B-4832-96CB-F0540CB338E3</mods:identifier>
</mods:mods>
<treatment id="038887803102054BFF619ADAFDD07B3C" ID-DOI="http://doi.org/10.5281/zenodo.6854026" ID-GBIF-Taxon="197819136" ID-Zenodo-Dep="6854026" LSID="urn:lsid:plazi:treatment:038887803102054BFF619ADAFDD07B3C" httpUri="http://treatment.plazi.org/id/038887803102054BFF619ADAFDD07B3C" lastPageId="13" lastPageNumber="540" pageId="11" pageNumber="538">
<subSubSection id="C33B651D3102054DFF619ADAFD2F7DDF" box="[151,738,151,178]" pageId="11" pageNumber="538" type="nomenclature">
<paragraph id="8B9E36963102054DFF619ADAFD2F7DDF" blockId="11.[151,738,151,213]" box="[151,738,151,178]" pageId="11" pageNumber="538">
<heading id="D0D681FA3102054DFF619ADAFD2F7DDF" bold="true" box="[151,738,151,178]" fontSize="11" level="1" pageId="11" pageNumber="538" reason="1">
<emphasis id="B955EA843102054DFF619ADAFD2F7DDF" bold="true" box="[151,738,151,178]" pageId="11" pageNumber="538">
<taxonomicName id="4C214D153102054DFF619ADAFDB07DDC" authority="Zhou &amp; Achterberg &amp; Tang &amp; Chen, 2022" authorityName="Zhou &amp; Achterberg &amp; Tang &amp; Chen" authorityYear="2022" box="[151,637,151,178]" class="Insecta" family="Braconidae" genus="Chelonus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="species" species="macrodontus" status="sp. nov." subGenus="Megachelonus">
<emphasis id="B955EA843102054DFF619ADAFEC67DDC" bold="true" box="[151,267,151,178]" italics="true" pageId="11" pageNumber="538">Chelonus</emphasis>
(
<emphasis id="B955EA843102054DFEEA9ADAFE037DDC" bold="true" box="[284,462,151,178]" italics="true" pageId="11" pageNumber="538">Megachelonus</emphasis>
)
<emphasis id="B955EA843102054DFE299ADAFDB07DDC" bold="true" box="[479,637,151,178]" italics="true" pageId="11" pageNumber="538">macrodontus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26657FF3102054DFD729ADAFD2F7DDF" box="[644,738,151,177]" pageId="11" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33B651D3102054DFF619AF6FECB7DBB" box="[151,262,187,213]" pageId="11" pageNumber="538" type="description">
<paragraph id="8B9E36963102054DFF619AF6FECB7DBB" blockId="11.[151,738,151,213]" box="[151,262,187,213]" pageId="11" pageNumber="538">
<figureCitation id="131A2A133102054DFF619AF6FECB7DBB" box="[151,262,187,213]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="11.[151,250,1915,1939]" captionStartId-1="12.[151,250,1942,1966]" captionTargetBox-0="[195,1393,993,1892]" captionTargetBox-1="[179,1408,181,1918]" captionTargetId-0="figure-315@11.[195,1393,993,1892]" captionTargetId-1="figure-22@12.[179,1408,181,1918]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 9. Chelonus (Megachelonus) macrodontus sp. nov., female, holotype, habitus, lateral aspect." captionText-1="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi-0="http://doi.org/10.5281/zenodo.6854048" figureDoi-1="http://doi.org/10.5281/zenodo.6854050" httpUri-0="https://zenodo.org/record/6854048/files/figure.png" httpUri-1="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Figs 9–10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B651D3102054DFF619B4EFEB97C2C" pageId="11" pageNumber="538" type="materials_examined">
<paragraph id="8B9E36963102054DFF619B4EFEB97C2C" blockId="11.[151,1437,259,970]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF619B4EFE417C73" bold="true" box="[151,396,259,285]" pageId="11" pageNumber="538">Material examined.</emphasis>
<materialsCitation id="3B493CCB3102054DFE6B9B4EFEB97C2C" ID-GBIF-Occurrence="3863920303" collectingDate="2005-06-23" collectionCode="ZJUH" collectorName="Jinxian Liu, No." country="China" location="Tongbiguan" pageId="11" pageNumber="538" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="549A88343102054DFE6B9B4EFDC77C73" box="[413,522,259,285]" pageId="11" pageNumber="538" type="holotype">Holotype</typeStatus>
: female,
<collectingCountry id="F33676063102054DFD739B4EFD067C73" box="[645,715,259,285]" name="China" pageId="11" pageNumber="538">China</collectingCountry>
,
<collectingRegion id="49E5F8743102054DFD299B49FC497C70" box="[735,900,260,286]" country="China" name="Yunnan" pageId="11" pageNumber="538">Yunnan Prov.</collectingRegion>
,
<location id="8EFE604D3102054DFC6F9B4EFBEC7C73" LSID="urn:lsid:plazi:treatment:038887803102054BFF619ADAFDD07B3C:8EFE604D3102054DFC6F9B4EFBEC7C73" box="[921,1057,259,285]" country="China" name="Tongbiguan" pageId="11" pageNumber="538" stateProvince="Yunnan">Tongbiguan</location>
,
<date id="FF9F10563102054DFBCE9B4EFB747C70" box="[1080,1209,259,286]" pageId="11" pageNumber="538" value="2005-06-23">
<collectingDate id="EFDBE9BE3102054DFBCE9B4EFB747C70" box="[1080,1209,259,286]" pageId="11" pageNumber="538" value="2005-06-23">23.VI.2005</collectingDate>
</date>
,
<collectorName id="26D453403102054DFB269B4EFA507C70" box="[1232,1437,259,286]" pageId="11" pageNumber="538">Jinxian Liu, No.</collectorName>
200609605(
<collectionCode id="ED30AE533102054DFED79B6AFEA57C2C" box="[289,360,295,322]" pageId="11" pageNumber="538">ZJUH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33B651D3102054DFF319B06FCAD7CE7" pageId="11" pageNumber="538" type="diagnosis">
<paragraph id="8B9E36963102054DFF319B06FCAD7CE7" blockId="11.[151,1437,259,970]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF319B06FE8F7C0B" bold="true" box="[199,322,331,357]" pageId="11" pageNumber="538">Diagnosis.</emphasis>
This species differs from the other species in the subgenus
<emphasis id="B955EA843102054DFC1C9B06FB4C7C0B" box="[1002,1153,331,357]" italics="true" pageId="11" pageNumber="538">Megachlonus</emphasis>
by having a pair of long and large teeth at apex of metasomal carapace (
<figureCitation id="131A2A133102054DFD5F9B22FC9C7CE7" box="[681,849,367,393]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Figs 10G, 10H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B651D3102054BFF319BDEFE457888" lastPageId="13" lastPageNumber="540" pageId="11" pageNumber="538" type="description">
<paragraph id="8B9E36963102054DFF319BDEFBE17CC0" blockId="11.[151,1437,259,970]" box="[199,1068,403,430]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF319BDEFE947CC3" bold="true" box="[199,345,403,429]" pageId="11" pageNumber="538">Description.</emphasis>
<typeStatus id="549A88343102054DFE969BDEFE067CC3" box="[352,459,403,429]" pageId="11" pageNumber="538" type="holotype">Holotype</typeStatus>
, female, length of body
<quantity id="4CD99B733102054DFD2C9BDEFCFF7CC0" box="[730,818,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="538" unit="mm" value="5.3">5.3 mm</quantity>
, of fore wing
<quantity id="4CD99B733102054DFC399BDEFBE57CC0" box="[975,1064,403,430]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="11" pageNumber="538" unit="mm" value="3.9">3.9 mm</quantity>
.
</paragraph>
<paragraph id="8B9E36963102054DFF319BFAFC947FA3" blockId="11.[151,1437,259,970]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF319BFAFEC37CBF" bold="true" box="[199,270,439,465]" pageId="11" pageNumber="538">Head.</emphasis>
Antennomeres 28 (remaining, broken in
<typeStatus id="549A88343102054DFD039BFAFC967CBF" box="[757,859,439,465]" pageId="11" pageNumber="538" type="holotype">holotype</typeStatus>
), basal half of antenna distinctly broadened, then rapidly tapering to apex; third antennomere 1.3 × longer than fourth; third, fourth and terminal antennomeres 4.7 ×, 3.7 × and 3.2 × longer than wide, respectively; 9-20
<superScript id="7C549BDE3102054DFD089BB2FCC67F63" attach="left" box="[766,779,511,525]" fontSize="6" pageId="11" pageNumber="538">th</superScript>
antennomeres flattened, distinctly shorter than wide; head transverse, in dorsal view roundly contracted behind eyes; OOL: OD: POL= 15: 6: 10; longitudinal diameter of eye 2.0 × height of temples; vertex coarsely striate-rugose; frons distinctly concave, reticulate-rugose but smooth near antennal sockets; temple irregularly reticulate-rugose (
<figureCitation id="131A2A133102054DFD099826FCAB7FEB" box="[767,870,619,645]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Fig. 10E</figureCitation>
); face coarsely reticulate-rugose with longitudinal elements, clypeus flat and densely punctate, smooth and shining, length of malar space 1.6 × basal width of mandible (
<figureCitation id="131A2A133102054DFF2898FEFE8A7FA3" box="[222,327,691,717]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Fig. 10D</figureCitation>
); face densely setose in lateral view (
<figureCitation id="131A2A133102054DFD1E98FEFC817FA3" box="[744,844,691,717]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Fig. 10F</figureCitation>
).
</paragraph>
<paragraph id="8B9E36963102054DFF31989AFBE77E33" blockId="11.[151,1437,259,970]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF31989AFE817F9F" bold="true" box="[199,332,727,753]" pageId="11" pageNumber="538">Mesosoma.</emphasis>
Length of mesosoma 1.4 × its height; mesoscutum areolate-rugose, but smooth and shiny medially; scutellum punctate medially, rugose laterally; scutellar suture comparatively deep; mesopleuron evenly convex, coarsely reticulate-rugose (
<figureCitation id="131A2A133102054DFE3F9952FDFD7E57" box="[457,560,799,825]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Fig. 10B</figureCitation>
); propodeum coarsely reticulaterugose, with a distinct transversal carina connected with 4 blunt tubercles, lateral tubercles larger than medial ones (
<figureCitation id="131A2A133102054DFC40990EFBD37E33" box="[950,1054,835,861]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Fig. 10C</figureCitation>
).
</paragraph>
<paragraph id="8B9E36963102054DFF31992AFDA07EA7" blockId="11.[151,1437,259,970]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF31992AFED77EEF" bold="true" box="[199,282,871,897]" pageId="11" pageNumber="538">Wings.</emphasis>
Length of fore wing 2.7 × its width; length of marginal cell 0.8 × pterostigma; vein 2-Rl just present, vein r shorter than 3-SR; r: 3-SR: SRl = 1: 1: 5; SRl more or less curved; parastigma distinctly swollen; 1-CU1: 2- CU1= 1: 2; vein CUlb present (
<figureCitation id="131A2A133102054DFE0E99E2FDAD7EA7" box="[504,608,943,969]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="11" pageNumber="538">Fig. 10A</figureCitation>
).
</paragraph>
<caption id="DF5E661E3102054DFF619D36FB667AFD" ID-DOI="http://doi.org/10.5281/zenodo.6854048" ID-Zenodo-Dep="6854048" box="[151,1195,1915,1940]" httpUri="https://zenodo.org/record/6854048/files/figure.png" pageId="11" pageNumber="538" startId="11.[151,250,1915,1939]" targetBox="[195,1393,993,1892]" targetPageId="11">
<paragraph id="8B9E36963102054DFF619D36FB667AFD" blockId="11.[151,1195,1915,1940]" box="[151,1195,1915,1940]" pageId="11" pageNumber="538">
<emphasis id="B955EA843102054DFF619D36FEDE7AFD" bold="true" box="[151,275,1915,1939]" pageId="11" pageNumber="538">FIGURE 9.</emphasis>
<taxonomicName id="4C214D153102054DFEEF9D36FD607AFD" authority="Zhou &amp; Achterberg &amp; Tang &amp; Chen, 2022" authorityName="Zhou &amp; Achterberg &amp; Tang &amp; Chen" authorityYear="2022" box="[281,685,1915,1940]" class="Insecta" family="Braconidae" genus="Chelonus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="538" phylum="Arthropoda" rank="species" species="macrodontus" status="sp. nov." subGenus="Megachelonus">
<emphasis id="B955EA843102054DFEEF9D36FEB47AFD" box="[281,377,1915,1939]" italics="true" pageId="11" pageNumber="538">Chelonus</emphasis>
(
<emphasis id="B955EA843102054DFE7E9D36FDD17AFD" box="[392,540,1915,1939]" italics="true" pageId="11" pageNumber="538">Megachelonus</emphasis>
)
<emphasis id="B955EA843102054DFDDC9D36FD607AFD" box="[554,685,1915,1939]" italics="true" pageId="11" pageNumber="538">macrodontus</emphasis>
</taxonomicName>
<emphasis id="B955EA843102054DFD429D31FCC87AFA" bold="true" box="[692,773,1916,1940]" pageId="11" pageNumber="538">
<taxonomicNameLabel id="A26657FF3102054DFD429D31FCC87AFA" box="[692,773,1916,1940]" pageId="11" pageNumber="538" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype, habitus, lateral aspect.
</paragraph>
</caption>
<caption id="DF5E661E3105054AFF619DDBFE7B7A98" ID-DOI="http://doi.org/10.5281/zenodo.6854050" ID-Zenodo-Dep="6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="12" pageNumber="539" startId="12.[151,250,1942,1966]" targetBox="[179,1408,181,1918]" targetPageId="12">
<paragraph id="8B9E36963105054AFF619DDBFE7B7A98" blockId="12.[151,1436,1942,2039]" pageId="12" pageNumber="539">
<emphasis id="B955EA843105054AFF619DDBFEED7AC0" bold="true" box="[151,288,1942,1966]" pageId="12" pageNumber="539">FIGURE 10.</emphasis>
<taxonomicName id="4C214D153105054AFED09DDBFD777AC0" authority="Zhou &amp; Achterberg &amp; Tang &amp; Chen, 2022" authorityName="Zhou &amp; Achterberg &amp; Tang &amp; Chen" authorityYear="2022" box="[294,698,1942,1967]" class="Insecta" family="Braconidae" genus="Chelonus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="539" phylum="Arthropoda" rank="species" species="macrodontus" status="sp. nov." subGenus="Megachelonus">
<emphasis id="B955EA843105054AFED09DDBFE4B7AC0" box="[294,390,1942,1966]" italics="true" pageId="12" pageNumber="539">Chelonus</emphasis>
(
<emphasis id="B955EA843105054AFE629DDBFDE57AC0" box="[404,552,1942,1966]" italics="true" pageId="12" pageNumber="539">Megachelonus</emphasis>
)
<emphasis id="B955EA843105054AFDC19DDBFD777AC0" box="[567,698,1942,1966]" italics="true" pageId="12" pageNumber="539">macrodontus</emphasis>
</taxonomicName>
<emphasis id="B955EA843105054AFD369DDAFCDC7AC1" bold="true" box="[704,785,1943,1967]" pageId="12" pageNumber="539">
<taxonomicNameLabel id="A26657FF3105054AFD369DDAFCDC7AC1" box="[704,785,1943,1967]" pageId="12" pageNumber="539" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female, holotype.
<emphasis id="B955EA843105054AFC389DDBFC2D7AC0" bold="true" box="[974,992,1942,1966]" pageId="12" pageNumber="539">A</emphasis>
, fore wing;
<emphasis id="B955EA843105054AFBAA9DDAFBA07AC1" bold="true" box="[1116,1133,1943,1967]" pageId="12" pageNumber="539">B</emphasis>
, mesosoma, lateral aspect;
<emphasis id="B955EA843105054AFA729DDBFA5B7AC0" bold="true" box="[1412,1430,1942,1966]" pageId="12" pageNumber="539">C</emphasis>
, mesosoma, dorsal aspect;
<emphasis id="B955EA843105054AFE579DF6FE7E7ABD" bold="true" box="[417,435,1979,2003]" pageId="12" pageNumber="539">D</emphasis>
, head, anterior aspect;
<emphasis id="B955EA843105054AFD579DF6FD7F7ABD" bold="true" box="[673,690,1979,2003]" pageId="12" pageNumber="539">E</emphasis>
, head, dorsal aspect;
<emphasis id="B955EA843105054AFC789DF6FC507ABD" bold="true" box="[910,925,1979,2003]" pageId="12" pageNumber="539">F</emphasis>
, head, lateral aspect;
<emphasis id="B955EA843105054AFB8D9DF7FB427ABC" bold="true" box="[1147,1167,1978,2002]" pageId="12" pageNumber="539">G</emphasis>
, metasoma, lateral aspect;
<emphasis id="B955EA843105054AFF619D92FF667A99" bold="true" box="[151,171,2015,2039]" pageId="12" pageNumber="539">H</emphasis>
, metasoma, dorsal aspect.
</paragraph>
</caption>
<caption id="DF5E661E3104054BFF619E0FFB217937" ID-DOI="http://doi.org/10.5281/zenodo.6854052" ID-Zenodo-Dep="6854052" box="[151,1260,1090,1114]" httpUri="https://zenodo.org/record/6854052/files/figure.png" pageId="13" pageNumber="540" startId="13.[151,250,1090,1114]" targetBox="[204,1384,181,1066]" targetPageId="13">
<paragraph id="8B9E36963104054BFF619E0FFB217937" blockId="13.[151,1260,1090,1114]" box="[151,1260,1090,1114]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF619E0FFED37934" bold="true" box="[151,286,1090,1114]" pageId="13" pageNumber="540">FIGURE 11.</emphasis>
<taxonomicName id="4C214D153104054BFED39E0FFC847937" authority="Zhang, Chen &amp; He" authorityName="Zhang, Chen &amp; He" authorityYear="2006" box="[293,841,1090,1114]" class="Insecta" family="Braconidae" genus="Chelonus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="540" phylum="Arthropoda" rank="species" species="macros" subGenus="Megachelonus">
<emphasis id="B955EA843104054BFED39E0FFE487934" box="[293,389,1090,1114]" italics="true" pageId="13" pageNumber="540">Chelonus</emphasis>
(
<emphasis id="B955EA843104054BFE659E0FFDEA7934" box="[403,551,1090,1114]" italics="true" pageId="13" pageNumber="540">Megachelonus</emphasis>
)
<emphasis id="B955EA843104054BFDC09E0EFD4D7934" box="[566,640,1091,1114]" italics="true" pageId="13" pageNumber="540">macros</emphasis>
Zhang, Chen &amp; He
</taxonomicName>
, female, holotype, habitus, lateral aspect.
</paragraph>
</caption>
<paragraph id="8B9E36963104054BFF319EC5FC5E79A8" blockId="13.[151,1437,1160,1619]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF319EC5FECB79CC" bold="true" box="[199,262,1160,1186]" pageId="13" pageNumber="540">Legs.</emphasis>
Hind coxa densely short setose and striate-rugose; hind femur robust; length of hind femur, tibia and basitarsus 3.2 ×, 4.5 × and 6.4 × their maximum width, respectively.
</paragraph>
<paragraph id="8B9E36963104054BFF319E9DFB267838" blockId="13.[151,1437,1160,1619]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF319E9DFE867984" bold="true" box="[199,331,1232,1258]" pageId="13" pageNumber="540">Metasoma.</emphasis>
Carapace elongate-oval in dorsal view, coarsely reticulaterugose basally, longitudinal striate-rugose medially, densely reticulaterugose apically; length of carapace 2.4 × its maximum width in dorsal view (
<figureCitation id="131A2A133104054BFA999EB9FF06785C" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="13" pageNumber="540">Fig. 10H</figureCitation>
); ventral cavity (as a narrow channel) open to apex, with a pair of long and large teeth; ovipositor long, slender and the ovipositor sheath 0.16 × as long as fore wing, 0.44 × as long as hind tibia (
<figureCitation id="131A2A133104054BFBC59F71FB137838" box="[1075,1246,1340,1366]" captionStart="FIGURE 10" captionStartId="12.[151,250,1942,1966]" captionTargetBox="[179,1408,181,1918]" captionTargetId="figure-22@12.[179,1408,181,1918]" captionTargetPageId="12" captionText="FIGURE 10. Chelonus (Megachelonus) macrodontus sp. nov. female, holotype. A, fore wing; B, mesosoma, lateral aspect; C, mesosoma, dorsal aspect; D, head, anterior aspect; E, head, dorsal aspect; F, head, lateral aspect; G, metasoma, lateral aspect; H, metasoma, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.6854050" httpUri="https://zenodo.org/record/6854050/files/figure.png" pageId="13" pageNumber="540">Figs 10G, 10H</figureCitation>
).
</paragraph>
<paragraph id="8B9E36963104054BFF319F2DFBFE78AC" blockId="13.[151,1437,1160,1619]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF319F2DFEED7814" bold="true" box="[199,288,1376,1402]" pageId="13" pageNumber="540">Colour.</emphasis>
Black; antennomeres blackish-brown; basal half of clypeus reddish-brown; pterostigma and veins blackish-brown; fore wing more or less infuscate; basal third of carapace yellowish; legs almost entirely yellowishbrown, hind tibia light yellow basally and black apically; ovipositor sheath brown.
</paragraph>
<paragraph id="8B9E36963104054BFF319F81FE457888" blockId="13.[151,1437,1160,1619]" box="[199,392,1484,1510]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF319F81FEC67888" bold="true" box="[199,267,1484,1510]" pageId="13" pageNumber="540">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33B651D3104054BFF319FBDFE697B64" box="[199,420,1520,1546]" pageId="13" pageNumber="540" type="biology_ecology">
<paragraph id="8B9E36963104054BFF319FBDFE697B64" blockId="13.[151,1437,1160,1619]" box="[199,420,1520,1546]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF319FBDFEEA7B64" bold="true" box="[199,295,1520,1546]" pageId="13" pageNumber="540">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C33B651D3104054BFF319C59FDD07B3C" pageId="13" pageNumber="540" type="etymology">
<paragraph id="8B9E36963104054BFF319C59FDD07B3C" blockId="13.[151,1437,1160,1619]" pageId="13" pageNumber="540">
<emphasis id="B955EA843104054BFF319C59FE837B40" bold="true" box="[199,334,1556,1582]" pageId="13" pageNumber="540">Etymology.</emphasis>
From
<emphasis id="B955EA843104054BFE6E9C58FE127B40" box="[408,479,1557,1582]" italics="true" pageId="13" pageNumber="540">macro</emphasis>
(Greek for long) and
<emphasis id="B955EA843104054BFD249C59FCE67B40" box="[722,811,1556,1582]" italics="true" pageId="13" pageNumber="540">odontos</emphasis>
(Greek for teeth), because of the apex of the metasomal carapace bears a pair of long teeth.
</paragraph>
</subSubSection>
</treatment>
</document>
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