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<mods:title id="807308FFF0AB19D44C6359AA5C8EBC2E">Phylogenetic status of some unique species representing Blapstinina Mulsant &amp; Rey (Tenebrionidae: Blaptinae: Opatrini), and implications for continued study of the subtribe</mods:title>
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<mods:namePart id="9DA706AF9842641B53FBE692E16271F9">Lumen, Ryan</mods:namePart>
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<mods:affiliation id="B5FCB27FE2BA815BC52960BE712BABB9">Zoological Museum, Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679 Warszawa, Poland &amp; https: // orcid. org / 0000 - 0002 - 3958 - 7596</mods:affiliation>
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<mods:namePart id="92914E185B8B8DC1C7BC891626A1CB6B">Kanda, Kojun</mods:namePart>
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<mods:affiliation id="3E5063ECA6A351064B4EFD6C2B3501D3">USDA Systematic Entomology Laboratory, c / o Smithsonian Institution, National Museum of Natural History, Washington, DC, U. S. A. https: // orcid. org / 0000 - 0001 - 5561 - 8471</mods:affiliation>
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<mods:affiliation id="924D0DDB7E34148D5248351615B9A603">Purdue University, Department of Entomology, 901 W. State Street, West Lafayette, IN 47907, USA https: // orcid. org / 0000 - 0002 - 1286 - 950 X</mods:affiliation>
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<treatment id="038187A07D08151F7986CA01FBF9FF30" ID-DOI="http://doi.org/10.5281/zenodo.6204799" ID-GBIF-Taxon="193277087" ID-Zenodo-Dep="6204799" LSID="urn:lsid:plazi:treatment:038187A07D08151F7986CA01FBF9FF30" httpUri="http://treatment.plazi.org/id/038187A07D08151F7986CA01FBF9FF30" lastPageId="7" lastPageNumber="526" pageId="5" pageNumber="524">
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<paragraph id="8B9736B67D08151D7986CA01FD27F8D6" blockId="5.[151,651,1851,1878]" box="[151,651,1851,1878]" pageId="5" pageNumber="524">
<heading id="D0DF81DA7D08151D7986CA01FD27F8D6" bold="true" box="[151,651,1851,1878]" fontSize="11" level="1" pageId="5" pageNumber="524" reason="1">
<emphasis id="B95CEAA47D08151D7986CA01FD27F8D6" bold="true" box="[151,651,1851,1878]" pageId="5" pageNumber="524">
<taxonomicName id="4C284D357D08151D7986CA01FE5BF8D5" authority="(Solier)" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="1851" box="[151,503,1851,1878]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="punctulatus" status="comb. nov.">
<emphasis id="B95CEAA47D08151D7986CA01FE3DF8D6" bold="true" box="[151,401,1851,1878]" italics="true" pageId="5" pageNumber="524">Lodinus punctulatus</emphasis>
(Solier)
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,
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<paragraph id="8B9736B67D08151D7986CABAFD9FF818" blockId="5.[151,1436,1920,2042]" box="[151,563,1920,1944]" pageId="5" pageNumber="524">
<treatmentCitationGroup id="AB3811987D08151D7986CABAFD9FF818" box="[151,563,1920,1944]" pageId="5" pageNumber="524">
<treatmentCitation id="0A8910A77D08151D7986CABAFD9FF818" author="Solier, A. J." box="[151,563,1920,1944]" page="233" pageId="5" pageNumber="524" year="1851">
<taxonomicName id="4C284D357D08151D7986CABAFD9FF818" authority="Solier, 1851: 233" authorityName="Solier" authorityPageNumber="233" authorityYear="1851" box="[151,563,1920,1944]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D08151D7986CABAFED2F818" box="[151,382,1920,1944]" italics="true" pageId="5" pageNumber="524">Blapstinus punctulatus</emphasis>
<bibRefCitation id="EFB94B477D08151D7894CABAFD9FF818" author="Solier, A. J." box="[389,563,1920,1944]" pageId="5" pageNumber="524" pagination="5 - 285" refId="ref10194" refString="Solier, A. J. (1851) Insectos. Coleopteros. In: Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella y publicada bajo los auspicios del supremo gobierno por Claudio Gay. Zoologia. Tomo Quinto. Museo de Historia Natural de Santiago, Paris, pp. 5 - 285." type="book chapter" year="1851">Solier, 1851: 233</bibRefCitation>
</taxonomicName>
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<paragraph id="8B9736B67D08151D7986CAA5FB7DF837" blockId="5.[151,1436,1920,2042]" box="[151,1233,1950,1975]" pageId="5" pageNumber="524">
<treatmentCitationGroup id="AB3811987D08151D7986CAA5FB7DF837" box="[151,1233,1950,1975]" pageId="5" pageNumber="524">
<emphasis id="B95CEAA47D08151D7986CAA5FF09F837" box="[151,165,1951,1975]" italics="true" pageId="5" pageNumber="524">=</emphasis>
<treatmentCitation id="0A8910A77D08151D79B4CAA5FD0DF837" author="Mulsant, E. &amp; Rey, C." box="[165,673,1950,1975]" page="131" pageId="5" pageNumber="524" year="1859">
<taxonomicName id="4C284D357D08151D79B4CAA5FD0DF837" authority="Mulsant and Rey, 1859: 131" authorityName="Mulsant and Rey" authorityPageNumber="131" authorityYear="1859" box="[165,673,1950,1975]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B95CEAA47D08151D79B4CAA5FED0F836" box="[165,380,1951,1975]" italics="true" pageId="5" pageNumber="524">Lodinus nigroaeneus</emphasis>
<bibRefCitation id="EFB94B477D08151D7892CAA5FD0DF837" author="Mulsant, E. &amp; Rey, C." box="[387,673,1950,1975]" pageId="5" pageNumber="524" pagination="65 - 157" refId="ref9986" refString="Mulsant, E. &amp; Rey, C. (1859) Essai d'une division des derniers Melasomes. Opuscules Entomologiques, 9, 65 - 157. https: // doi. org / 10.5962 / bhl. title. 2682" type="journal article" year="1859">Mulsant and Rey, 1859: 131</bibRefCitation>
</taxonomicName>
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(syn. by Gemminger in
<bibRefCitation id="EFB94B477D08151D7A88CAA5FB66F837" author="Gemminger, M. &amp; Harold, E. von" box="[921,1226,1951,1975]" pageId="5" pageNumber="524" pagination="1801 - 2179" refId="ref8577" refString="Gemminger, M. &amp; Harold, E. von (1870) a. n. In: Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. VII. Tenebrionidae, Nilionidae, Pythidae, Melandryidae, Lagriidae, Pedilidae, Anthicidae, Pyrochroidae, Mordellidae, Rhipidophoridae, Cantharidae, Oedemeridae. E. H. Gummi, Monachium [Munchen], pp. 1801 - 2179." type="book chapter" year="1870">Gemminger and Harold, 1870</bibRefCitation>
)
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</paragraph>
<paragraph id="8B9736B67D08151D7986CA84FCF1F855" blockId="5.[151,1436,1920,2042]" box="[151,861,1981,2005]" pageId="5" pageNumber="524">
<treatmentCitationGroup id="AB3811987D08151D7986CA84FCF1F855" box="[151,861,1981,2005]" pageId="5" pageNumber="524">
=
<treatmentCitation id="0A8910A77D08151D79B4CA87FD96F855" author="Curtis, J." box="[165,570,1981,2005]" page="469" pageId="5" pageNumber="524" year="1845">
<taxonomicName id="4C284D357D08151D79B4CA87FD96F855" authority="Curtis, 1845: 469" authorityName="Curtis" authorityPageNumber="469" authorityYear="1845" box="[165,570,1981,2005]" class="Insecta" family="Tenebrionidae" genus="Alphitobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B95CEAA47D08151D79B4CA87FE28F855" box="[165,388,1981,2005]" italics="true" pageId="5" pageNumber="524">Alphitobius punctatus</emphasis>
<bibRefCitation id="EFB94B477D08151D7898CA87FD96F855" author="Curtis, J." box="[393,570,1981,2005]" pageId="5" pageNumber="524" pagination="441 - 476" refId="ref8290" refString="Curtis, J. (1845) Descriptions, &amp; c. of the insects collected by Captain P. P. King in the survey of the Straits of Magellan. Transactions of the Linnean Society of London, 19, 441 - 476. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis, 1845: 469</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(syn. by
<treatmentCitation id="0A8910A77D08151D7B89CA87FCFEF855" author="Gebien, H." box="[664,850,1981,2005]" page="405" pageId="5" pageNumber="524" year="1938">
<bibRefCitation id="EFB94B477D08151D7B89CA87FCFEF855" author="Gebien, H." box="[664,850,1981,2005]" pageId="5" pageNumber="524" pagination="283 - 314" refId="ref8544" refString="Gebien, H. (1938) Katalog der Tenebrioniden. Teil II. Mitteilungen der Munchener Entomologischen Gesellschaft, 28, 283 - 314. [kat. 402 - 433]" type="journal article" year="1938">Gebien 1938: 405</bibRefCitation>
</treatmentCitation>
).
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</paragraph>
<paragraph id="8B9736B67D08151E7986CADAFAB2FEC2" blockId="5.[151,1436,1920,2042]" lastBlockId="6.[151,1437,150,2037]" lastPageId="6" lastPageNumber="525" pageId="5" pageNumber="524">
<emphasis id="B95CEAA47D08151D7986CADAFF63F87A" bold="true" box="[151,207,2016,2042]" pageId="5" pageNumber="524">Note</emphasis>
: While proposing synonymy between
<taxonomicName id="4C284D357D08151D7B97CAE5FBBDF87A" authority="Curtis, 1845" authorityName="Curtis" authorityYear="1845" box="[646,1041,2015,2042]" class="Insecta" family="Tenebrionidae" genus="Alphitobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B95CEAA47D08151D7B97CAE5FCD1F879" box="[646,893,2015,2041]" italics="true" pageId="5" pageNumber="524">Alphitobius punctatus</emphasis>
<bibRefCitation id="EFB94B477D08151D7A94CAE5FBBDF87A" author="Curtis, J." box="[901,1041,2015,2042]" pageId="5" pageNumber="524" pagination="441 - 476" refId="ref8290" refString="Curtis, J. (1845) Descriptions, &amp; c. of the insects collected by Captain P. P. King in the survey of the Straits of Magellan. Transactions of the Linnean Society of London, 19, 441 - 476. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis, 1845</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C284D357D08151E7D5BCAE5FF7DFF31" authority="Solier, 1851" authorityName="Solier" authorityYear="1851" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" lastPageId="6" lastPageNumber="525" order="Coleoptera" pageId="5" pageNumber="524" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D08151D7D5BCAE5FAE7F879" box="[1098,1355,2015,2041]" italics="true" pageId="5" pageNumber="524">Blapstinus punctulatus</emphasis>
<bibRefCitation id="EFB94B477D08151E7C42CAE5FF7DFF31" author="Solier, A. J." lastPageId="6" lastPageNumber="525" pageId="5" pageNumber="524" pagination="5 - 285" refId="ref10194" refString="Solier, A. J. (1851) Insectos. Coleopteros. In: Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella y publicada bajo los auspicios del supremo gobierno por Claudio Gay. Zoologia. Tomo Quinto. Museo de Historia Natural de Santiago, Paris, pp. 5 - 285." type="book chapter" year="1851">Solier, 1851</bibRefCitation>
</taxonomicName>
, to avoid secondary homonymy with
<taxonomicName id="4C284D357D0B151E7B99CDADFC50FF32" authority="(Fabricius)" baseAuthorityName="Fabricius" box="[648,1020,151,178]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B95CEAA47D0B151E7B99CDADFCD9FF30" box="[648,885,151,177]" italics="true" pageId="6" pageNumber="525">Blapstinus punctatus</emphasis>
(Fabricius)
</taxonomicName>
,
<bibRefCitation id="EFB94B477D0B151E7D1BCDADFB03FF31" author="Gebien, H." box="[1034,1199,151,177]" pageId="6" pageNumber="525" pagination="283 - 314" refId="ref8544" refString="Gebien, H. (1938) Katalog der Tenebrioniden. Teil II. Mitteilungen der Munchener Entomologischen Gesellschaft, 28, 283 - 314. [kat. 402 - 433]" type="journal article" year="1938">Gebien (1938)</bibRefCitation>
rejected
<taxonomicName id="4C284D357D0B151E7C0ACDADFE30FF55" authority="Curtis, 1845" authorityName="Curtis" authorityYear="1845" class="Insecta" family="Tenebrionidae" genus="Alphitobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="B95CEAA47D0B151E7C0ACDADFEA8FF54" italics="true" pageId="6" pageNumber="525">Alphitobius punctatus</emphasis>
<bibRefCitation id="EFB94B477D0B151E781FCD81FE30FF55" author="Curtis, J." box="[270,412,186,213]" pageId="6" pageNumber="525" pagination="441 - 476" refId="ref8290" refString="Curtis, J. (1845) Descriptions, &amp; c. of the insects collected by Captain P. P. King in the survey of the Straits of Magellan. Transactions of the Linnean Society of London, 19, 441 - 476. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis, 1845</bibRefCitation>
</taxonomicName>
and replaced it with its oldest available synonym—
<taxonomicName id="4C284D357D0B151E7AECCD81FA34FF55" authority="Solier, 1851" authorityName="Solier" authorityYear="1851" box="[1021,1432,186,213]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E7AECCD81FAACFF55" box="[1021,1280,187,213]" italics="true" pageId="6" pageNumber="525">Blapstinus punctulatus</emphasis>
<bibRefCitation id="EFB94B477D0B151E7C1BCD81FA34FF55" author="Solier, A. J." box="[1290,1432,186,213]" pageId="6" pageNumber="525" pagination="5 - 285" refId="ref10194" refString="Solier, A. J. (1851) Insectos. Coleopteros. In: Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella y publicada bajo los auspicios del supremo gobierno por Claudio Gay. Zoologia. Tomo Quinto. Museo de Historia Natural de Santiago, Paris, pp. 5 - 285." type="book chapter" year="1851">Solier, 1851</bibRefCitation>
</taxonomicName>
. According to the regulations of ICZN (1999), particularly art. 59.3, a junior secondary homonym replaced before 1961 is permanently invalid unless the substitute name is not in use and the relevant taxa are no longer considered congeneric. As a result the valid name of the below redescribed species is
<taxonomicName id="4C284D357D0B151E7AC7CC1DFAB5FEC2" authority="(Solier)" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="1851" box="[982,1305,295,322]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E7AC7CC1DFB16FEC1" box="[982,1210,295,321]" italics="true" pageId="6" pageNumber="525">Lodinus punctulatus</emphasis>
(Solier)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C332653D7D0B151E79D6CC71FC24FE09" pageId="6" pageNumber="525" type="description">
<paragraph id="8B9736B67D0B151E79D6CC71FC24FE09" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
Descriptions of
<taxonomicName id="4C284D357D0B151E786ACC71FDB2FEE5" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="1851" box="[379,542,331,357]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E786ACC71FDB2FEE5" box="[379,542,331,357]" italics="true" pageId="6" pageNumber="525">B. punctulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C284D357D0B151E7B44CC71FD50FEE4" box="[597,764,331,357]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B95CEAA47D0B151E7B44CC71FDC9FEE4" box="[597,613,331,356]" italics="true" pageId="6" pageNumber="525">L</emphasis>
.
<emphasis id="B95CEAA47D0B151E7B62CC71FD50FEE4" box="[627,764,331,356]" italics="true" pageId="6" pageNumber="525">nigroaeneus</emphasis>
</taxonomicName>
do not include
<typeStatus id="549388147D0B151E7ABDCC71FBA2FEE5" box="[940,1038,331,357]" pageId="6" pageNumber="525" type="holotype">holotype</typeStatus>
designations. As a result,
<typeStatus id="549388147D0B151E7C27CC71FA31FEE5" box="[1334,1437,331,357]" pageId="6" pageNumber="525" type="lectotype">lectotype</typeStatus>
designations are needed to fix the taxonomic status of these names.
</paragraph>
</subSubSection>
<subSubSection id="C332653D7D0B151E79D6CCA8FCB1FCDD" pageId="6" pageNumber="525" type="materials_examined">
<paragraph id="8B9736B67D0B151E79D6CCA8FB2AFE75" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
<emphasis id="B95CEAA47D0B151E79D6CCA8FE2AFE2D" bold="true" box="[199,390,402,429]" italics="true" pageId="6" pageNumber="525">Studied material</emphasis>
.
<emphasis id="B95CEAA47D0B151E7884CCA9FDEAFE2C" bold="true" box="[405,582,402,429]" pageId="6" pageNumber="525">Type material:</emphasis>
<taxonomicName id="4C284D357D0B151E7B5CCCA9FCE2FE2D" authorityName="Solier" authorityYear="1851" box="[589,846,403,429]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E7B5CCCA9FCE2FE2D" box="[589,846,403,429]" italics="true" pageId="6" pageNumber="525">Blapstinus punctulatus</emphasis>
</taxonomicName>
:
<materialsCitation id="3B403CEB7D0B151E7A4CCCA8FD81FE75" ID-GBIF-Occurrence="3462300309" collectionCode="MNHN, TYPE" country="Chile" county="Solier" location="Punc" municipality="Type" pageId="6" pageNumber="525" specimenCount="1" typeStatus="lectotype">
<emphasis id="B95CEAA47D0B151E7A4CCCA8FB73FE2C" bold="true" box="[861,1247,402,428]" pageId="6" pageNumber="525">
<typeStatus id="549388147D0B151E7A4CCCA8FBA5FE2C" box="[861,1033,402,428]" pageId="6" pageNumber="525" type="lectotype">LECTOTYPE</typeStatus>
(here designated)
</emphasis>
, male (
<collectionCode id="ED39AE737D0B151E7C25CCA9FA3DFE2D" box="[1332,1425,403,429]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="525">MNHN</collectionCode>
): “
<location id="8EF7606D7D0B151E79B0CC8DFF70FE51" LSID="urn:lsid:plazi:treatment:038187A07D08151F7986CA01FBF9FF30:8EF7606D7D0B151E79B0CC8DFF70FE51" box="[161,220,439,465]" country="Chile" county="Solier" municipality="Type" name="Punc" pageId="6" pageNumber="525">Punc</location>
\tulatus” reverse side of the same label “
<collectingMunicipality id="6BF3ACCC7D0B151E7BA9CC8DFD58FE51" box="[696,756,439,465]" pageId="6" pageNumber="525">Type</collectingMunicipality>
\
<collectingCounty id="62F64E3A7D0B151E7BECCC8DFCEFFE51" box="[765,835,439,465]" pageId="6" pageNumber="525">Solier</collectingCounty>
”, “
<taxonomicName id="4C284D357D0B151E7A74CC8DFC6DFE51" box="[869,961,439,465]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus">Lodinus</taxonomicName>
\punctulatus\
<collectingCountry id="F33F76267D0B151E7D44CC8DFB21FE51" box="[1109,1165,439,465]" name="Chile" pageId="6" pageNumber="525">Chili</collectingCountry>
<emphasis id="B95CEAA47D0B151E7D89CC8DFB58FE51" box="[1176,1268,439,465]" italics="true" pageId="6" pageNumber="525">illegible</emphasis>
”, “
<taxonomicName id="4C284D357D0B151E7C0ACC8DFA3FFE51" authorityName="Sturm" authorityYear="1826" box="[1307,1427,439,465]" class="Insecta" family="Tenebrionidae" genus="Blapstinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="532" phylum="Arthropoda" rank="genus">Blapstinus</taxonomicName>
\ punctulatus\
<collectingCountry id="F33F76267D0B151E7832CCE1FEF7FE75" box="[291,347,475,501]" name="Chile" pageId="6" pageNumber="525">Chili</collectingCountry>
\D. M-?”, „
<collectionCode id="ED39AE737D0B151E78C4CCE1FD8FFE75" box="[469,547,475,501]" pageId="6" pageNumber="525">TYPE</collectionCode>
</materialsCitation>
.
<emphasis id="B95CEAA47D0B151E7B29CCE0FC9CFE74" bold="true" box="[568,816,474,500]" pageId="6" pageNumber="525">
<typeStatus id="549388147D0B151E7B29CCE0FC9CFE74" box="[568,816,474,500]" pageId="6" pageNumber="525" type="paralectotype">PARALECTOTYPE</typeStatus>
</emphasis>
, male: same data as lectotype.
</paragraph>
<paragraph id="8B9736B67D0B151E79D6CCC5FE36FDE1" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
<taxonomicName id="4C284D357D0B151E79D6CCC5FE18FD98" box="[199,436,511,537]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B95CEAA47D0B151E79D6CCC5FE18FD98" box="[199,436,511,537]" italics="true" pageId="6" pageNumber="525">Lodinus nigroaeneus</emphasis>
</taxonomicName>
:
<materialsCitation id="3B403CEB7D0B151E78D7CCC4FC46FDBD" ID-GBIF-Occurrence="3462300308" collectionCode="MNHN, TYPE" country="Chile" location="Mulsant" pageId="6" pageNumber="525" specimenCount="1" typeStatus="lectotype">
<emphasis id="B95CEAA47D0B151E78D7CCC4FCE7FD98" bold="true" box="[454,843,510,536]" pageId="6" pageNumber="525">
<typeStatus id="549388147D0B151E78D7CCC4FDDEFD98" box="[454,626,510,536]" pageId="6" pageNumber="525" type="lectotype">LECTOTYPE</typeStatus>
(here designated)
</emphasis>
, male (
<collectionCode id="ED39AE737D0B151E7AB5CCC5FBADFD99" box="[932,1025,511,537]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="525">MNHN</collectionCode>
): „nigro-aeneus” reverse side of the same label “
<location id="8EF7606D7D0B151E7831CF19FED3FDBD" LSID="urn:lsid:plazi:treatment:038187A07D08151F7986CA01FBF9FF30:8EF7606D7D0B151E7831CF19FED3FDBD" box="[288,383,547,573]" country="Chile" name="Mulsant" pageId="6" pageNumber="525">Mulsant</location>
type”, “
<taxonomicName id="4C284D357D0B151E78CACF19FD9BFDBD" box="[475,567,547,573]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus">Lodinus</taxonomicName>
/nigroaeneus/
<collectingCountry id="F33F76267D0B151E7BC0CF19FCA5FDBD" box="[721,777,547,573]" name="Chile" pageId="6" pageNumber="525">Chili</collectingCountry>
<emphasis id="B95CEAA47D0B151E7A00CF19FCC1FDBD" box="[785,877,547,573]" italics="true" pageId="6" pageNumber="525">illegible</emphasis>
”, “
<collectionCode id="ED39AE737D0B151E7A83CF19FC4CFDBD" box="[914,992,547,573]" pageId="6" pageNumber="525">TYPE</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B403CEB7D0B151E7AE4CF18FE3AFDE1" ID-GBIF-Occurrence="3462300304" collectionCode="MNHN, TYPE" country="Chile" location="Mulsant" pageId="6" pageNumber="525" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<emphasis id="B95CEAA47D0B151E7AE4CF18FB50FDBC" bold="true" box="[1013,1276,546,572]" pageId="6" pageNumber="525">
<typeStatus id="549388147D0B151E7AE4CF18FB50FDBC" box="[1013,1276,546,572]" pageId="6" pageNumber="525" type="paralectotype">PARALECTOTYPES</typeStatus>
</emphasis>
,
<specimenCount id="9D2EFD3F7D0B151E7C1DCF19FA35FDBD" box="[1292,1433,547,573]" pageId="6" pageNumber="525" type="female">two females</specimenCount>
: same data as lectotype
</materialsCitation>
.
</paragraph>
<paragraph id="8B9736B67D0B151E79D6CF50FCB1FCDD" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
<emphasis id="B95CEAA47D0B151E79D6CF50FE11FD04" bold="true" box="[199,445,618,644]" pageId="6" pageNumber="525">Additional material:</emphasis>
<materialsCitation id="3B403CEB7D0B151E78D4CF51FCB1FCDD" ID-GBIF-Occurrence="3462300310" collectingDate="1955-11-08" collectingDateMax="2011-03-07" collectingDateMin="1955-11-08" collectionCode="PERC, R, MIZ, PAS, V, MNHN" collectorName="M. Dieguez &amp; Coll. P. Ardoin" country="Argentina" county="Cordoba" elevation="350" latitude="-31.3755" location="Maddison" longLatPrecision="6" longitude="-65.4797" municipality="Reserva Chancani" pageId="6" pageNumber="525" specimenCount="33" specimenCount-female="2" specimenCount-male="4" stateProvince="Chubut">
male (
<collectionCode id="ED39AE737D0B151E7B1DCF51FDF5FD05" box="[524,601,619,645]" pageId="6" pageNumber="525">PERC</collectionCode>
): “
<collectingCountry id="F33F76267D0B151E7B67CF51FC8DFD05" box="[630,801,619,645]" name="Argentina" pageId="6" pageNumber="525">ARGENTINA</collectingCountry>
:
<collectingCounty id="62F64E3A7D0B151E7A3DCF51FC3DFD05" box="[812,913,619,645]" pageId="6" pageNumber="525">Cordoba</collectingCounty>
:
<collectingMunicipality id="6BF3ACCC7D0B151E7A8DCF51FBC0FD05" box="[924,1132,619,645]" pageId="6" pageNumber="525">Reserva Chancani</collectingMunicipality>
,
<quantity id="4CD09B537D0B151E7D69CF50FB16FD05" box="[1144,1210,618,645]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="525" unit="m" value="350.0">
<elevation id="0005D1857D0B151E7D69CF50FB16FD05" box="[1144,1210,618,645]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="525" unit="m" value="350.0">350m</elevation>
</quantity>
,
<geoCoordinate id="EE1C50717D0B151E7DD7CF50FA96FD05" box="[1222,1338,618,645]" degrees="31.3755" direction="south" orientation="latitude" pageId="6" pageNumber="525" precision="5" value="-31.3755">31.3755 S</geoCoordinate>
<geoCoordinate id="EE1C50717D0B151E7C50CF50FF1EFD29" degrees="65.4797" direction="west" orientation="longitude" pageId="6" pageNumber="525" precision="5" value="-65.4797">65.4797 W</geoCoordinate>
,
<date id="FF9610767D0B151E79ACCFB5FE8BFD29" box="[189,295,655,681]" pageId="6" pageNumber="525" value="2011-03-07">
<collectingDate id="EFD2E99E7D0B151E79ACCFB5FE8BFD29" box="[189,295,655,681]" pageId="6" pageNumber="525" value="2011-03-07">7.iii.2011</collectingDate>
</date>
DRM 11.007. D.
<collectionCode id="ED39AE737D0B151E78F9CFB5FE52FD29" box="[488,510,655,681]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="525">R</collectionCode>
.
<location id="8EF7606D7D0B151E7B18CFB5FDD3FD29" LSID="urn:lsid:plazi:treatment:038187A07D08151F7986CA01FBF9FF30:8EF7606D7D0B151E7B18CFB5FDD3FD29" box="[521,639,655,681]" country="Argentina" county="Cordoba" latitude="-31.3755" longLatPrecision="6" longitude="-65.4797" municipality="Reserva Chancani" name="Maddison" pageId="6" pageNumber="525" stateProvince="Chubut">Maddison</location>
”;
<specimenCount id="9D2EFD3F7D0B151E7B82CFB5FD45FD29" box="[659,745,655,681]" pageId="6" pageNumber="525" type="male">2 males</specimenCount>
and female (
<collectionCode id="ED39AE737D0B151E7A68CFB5FC02FD29" box="[889,942,655,681]" pageId="6" pageNumber="525">MIZ</collectionCode>
<collectionCode id="ED39AE737D0B151E7AA5CFB5FC47FD29" box="[948,1003,655,681]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="6" pageNumber="525" type="Herbarium">PAS</collectionCode>
): „Chile Santiago \ Baños Morales \
<date id="FF9610767D0B151E7C91CFB5FEBBFD4D" pageId="6" pageNumber="525" value="2001-05-19">
<collectingDate id="EFD2E99E7D0B151E7C91CFB5FEBBFD4D" pageId="6" pageNumber="525" value="2001-05-19">19 Mayo 2001</collectingDate>
</date>
\leg.
<collectionCode id="ED39AE737D0B151E7844CF89FEC5FD4D" box="[341,361,691,717]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="525" type="Museum">V</collectionCode>
. Manuel Diéguez”;
<specimenCount id="9D2EFD3F7D0B151E7B59CF89FD32FD4D" box="[584,670,691,717]" pageId="6" pageNumber="525" type="male">2 males</specimenCount>
and female (
<collectionCode id="ED39AE737D0B151E7A3ECF89FCC8FD4D" box="[815,868,691,717]" pageId="6" pageNumber="525">MIZ</collectionCode>
<collectionCode id="ED39AE737D0B151E7A7ACF89FC0EFD4D" box="[875,930,691,717]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="6" pageNumber="525" type="Herbarium">PAS</collectionCode>
): „Argentina
<collectingRegion id="49ECF8547D0B151E7D27CF89FB24FD4D" box="[1078,1160,691,717]" country="Argentina" name="Chubut" pageId="6" pageNumber="525">Chubut</collectingRegion>
\ Camino a cholila \ 580 MSNM \ 23/
<date id="FF9610767D0B151E7839CFEDFE67FD71" box="[296,459,727,753]" pageId="6" pageNumber="525" value="2000-01-31">
<collectingDate id="EFD2E99E7D0B151E7839CFEDFE67FD71" box="[296,459,727,753]" pageId="6" pageNumber="525" value="2000-01-31">31 Enero 2000</collectingDate>
</date>
\ Leg.
<collectionCode id="ED39AE737D0B151E7B08CFEDFD83FD71" box="[537,559,727,753]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="525" type="Museum">V</collectionCode>
.
<collectorName id="26DD53607D0B151E7B23CFEDFD19FD71" box="[562,693,727,753]" pageId="6" pageNumber="525">M. Diéguez</collectorName>
M.”; male and
<specimenCount id="9D2EFD3F7D0B151E7A75CFEDFC7CFD71" box="[868,976,727,753]" pageId="6" pageNumber="525" type="female">2 females</specimenCount>
(
<collectionCode id="ED39AE737D0B151E7ACFCFEDFBBFFD71" box="[990,1043,727,753]" pageId="6" pageNumber="525">MIZ</collectionCode>
<collectionCode id="ED39AE737D0B151E7D0BCFEDFBFDFD71" box="[1050,1105,727,753]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="6" pageNumber="525" type="Herbarium">PAS</collectionCode>
): „Chile Melipilla \ Pallocabe 300Mts. \
<date id="FF9610767D0B151E7819CFC1FE3EFC95" box="[264,402,763,789]" pageId="6" pageNumber="525" value="2001-09-09">
<collectingDate id="EFD2E99E7D0B151E7819CFC1FE3EFC95" box="[264,402,763,789]" pageId="6" pageNumber="525" value="2001-09-09">9 Sept. 2001</collectingDate>
</date>
\ leg.
<collectionCode id="ED39AE737D0B151E78C4CFC1FE45FC95" box="[469,489,763,789]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="525" type="Museum">V</collectionCode>
. Manuel Diéguez”; male and female (
<collectionCode id="ED39AE737D0B151E7A9ACFC1FC6CFC95" box="[907,960,763,789]" pageId="6" pageNumber="525">MIZ</collectionCode>
<collectionCode id="ED39AE737D0B151E7AD7CFC1FC51FC95" box="[966,1021,763,789]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="6" pageNumber="525" type="Herbarium">PAS</collectionCode>
): “Santiago Thilippes”;
<specimenCount id="9D2EFD3F7D0B151E7C17CFC1FA31FC95" box="[1286,1437,763,789]" pageId="6" pageNumber="525" type="generic">27 specimens</specimenCount>
(
<collectionCode id="ED39AE737D0B151E798ECE25FF78FCB9" box="[159,212,799,825]" pageId="6" pageNumber="525">MIZ</collectionCode>
<collectionCode id="ED39AE737D0B151E79CACE25FEBEFCB9" box="[219,274,799,825]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="6" pageNumber="525" type="Herbarium">PAS</collectionCode>
): „Chile Philippi S”; female (
<collectionCode id="ED39AE737D0B151E7B48CE25FD1AFCB9" box="[601,694,799,825]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="525">MNHN</collectionCode>
): “Brasilien\Nova Teutania\2711B—52
<quantity id="4CD09B537D0B151E7D6FCE25FB0EFCB9" box="[1150,1186,799,825]" metricMagnitude="4" metricUnit="kg" metricValue="2.3" pageId="6" pageNumber="525" unit="t" value="23.0">23t</quantity>
\Fritz Plaumann\
<quantity id="4CD09B537D0B151E7C74CE25FF77FCDD" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="6" pageNumber="525" unit="m" value="550.0" valueMax="800.0" valueMin="300.0">
<elevation id="0005D1857D0B151E7C74CE25FF77FCDD" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="6" pageNumber="525" unit="m" value="550.0" valueMax="800.0" valueMin="300.0">300 800m</elevation>
</quantity>
\
<date id="FF9610767D0B151E79F5CE78FEF6FCDD" box="[228,346,834,861]" pageId="6" pageNumber="525" value="1955-11-08">
<collectingDate id="EFD2E99E7D0B151E79F5CE78FEF6FCDD" box="[228,346,834,861]" pageId="6" pageNumber="525" value="1955-11-08">8-XI-1955</collectingDate>
</date>
”, “Museum Paris \
<collectorName id="26DD53607D0B151E7B3CCE79FD7EFCDD" box="[557,722,835,861]" pageId="6" pageNumber="525">Coll. P. Ardoin</collectorName>
1978”
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C332653D7D0B151E79D6CE5CFACEFA8A" pageId="6" pageNumber="525" type="diagnosis">
<paragraph id="8B9736B67D0B151E79D6CE5CFF61FC6D" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
<emphasis id="B95CEAA47D0B151E79D6CE5CFE96FC00" bold="true" box="[199,314,870,896]" pageId="6" pageNumber="525">Diagnosis</emphasis>
. Newly examined material enables supplementation of available descriptions of this species (
<bibRefCitation id="EFB94B477D0B151E7C47CE5DFF7EFC25" author="Curtis, J." pageId="6" pageNumber="525" pagination="441 - 476" refId="ref8290" refString="Curtis, J. (1845) Descriptions, &amp; c. of the insects collected by Captain P. P. King in the survey of the Straits of Magellan. Transactions of the Linnean Society of London, 19, 441 - 476. https: // doi. org / 10.1111 / j. 1096 - 3642.1842. tb 00371. x" type="journal article" year="1845">Curtis 1845</bibRefCitation>
,
<bibRefCitation id="EFB94B477D0B151E79CCCEB1FECCFC25" author="Solier, A. J." box="[221,352,906,933]" pageId="6" pageNumber="525" pagination="5 - 285" refId="ref10194" refString="Solier, A. J. (1851) Insectos. Coleopteros. In: Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella y publicada bajo los auspicios del supremo gobierno por Claudio Gay. Zoologia. Tomo Quinto. Museo de Historia Natural de Santiago, Paris, pp. 5 - 285." type="book chapter" year="1851">Solier 1851</bibRefCitation>
,
<bibRefCitation id="EFB94B477D0B151E787DCEB1FDC0FC25" author="Mulsant, E. &amp; Rey, C." box="[364,620,906,933]" pageId="6" pageNumber="525" pagination="65 - 157" refId="ref9986" refString="Mulsant, E. &amp; Rey, C. (1859) Essai d'une division des derniers Melasomes. Opuscules Entomologiques, 9, 65 - 157. https: // doi. org / 10.5962 / bhl. title. 2682" type="journal article" year="1859">Mulsant and Rey 1859</bibRefCitation>
,
<bibRefCitation id="EFB94B477D0B151E7B64CEB1FCE0FC26" author="Aballay, F. H. &amp; Flores, G. E. &amp; Silvestro, V. A. &amp; Zanetti, N. I. &amp; Centeno, N. D." box="[629,844,907,934]" pageId="6" pageNumber="525" pagination="703 - 726" refId="ref7819" refString="Aballay, F. H., Flores, G. E., Silvestro, V. A., Zanetti, N. I. &amp; Centeno, N. D. (2016) An illustrated key to, and diagnoses of the species of Tenebrionidae (Coleoptera) associated with decaying carcasses in Argentina. Annales Zoologici, 66, 703 - 726. https: // doi. org / 10.3161 / 00034541 ANZ 2016.66.4.021" type="journal article" year="2016">
Aballay
<emphasis id="B95CEAA47D0B151E7BC5CEB1FCA9FC25" box="[724,773,907,933]" italics="true" pageId="6" pageNumber="525">et al</emphasis>
. 2016
</bibRefCitation>
) with data on male and female terminalia. Therefore, a redescription is provided below and a new diagnosis is provided to accommodate this species with its new congeners.
</paragraph>
<paragraph id="8B9736B67D0B151E79D6CECDFC58FBB5" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
The following features place this species within
<taxonomicName id="4C284D357D0B151E7BECCECDFCFBFB91" box="[765,855,1015,1041]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B95CEAA47D0B151E7BECCECDFCFBFB91" box="[765,855,1015,1041]" italics="true" pageId="6" pageNumber="525">Lodinus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFB94B477D0B151E7A78CECDFBB6FB91" author="Marcuzzi, G." box="[873,1050,1014,1041]" pageId="6" pageNumber="525" pagination="1 - 36" refId="ref9624" refString="Marcuzzi, G. (1954) Studies on the fauna of Curacao and other Caribbean Islands: no. 22. Tenebrionid beetles of Curacao, Aruba, Bonaire, and the Venezuelan Islands. Natuurwetenschappelijke Studiekring voor Suriname, 10, 1 - 36." type="journal article" year="1954">Marcuzzi 1954</bibRefCitation>
, 1982): median lobe of aedeagus widely projecting between parameres; parameres with truncate apex (
<figureCitation id="13132A337D0B151E7AB0C921FC4FFBB5" box="[929,995,1051,1077]" captionStart="FIGURE 1" captionStartId="3.[151,250,1675,1699]" captionTargetBox="[151,1419,525,1640]" captionTargetId="figure-169@3.[151,1436,525,1651]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of subtribe Blapstinina. Figured topology was recovered in maximum likelihood analysis using IQ- Tree. Ultrafast bootstrap values (1,000 replicates) (integers) and posterior probabilities (20 million generations) acquired in Bayesian analysis (decimals) are displayed in red. The three investigated Blapstinina species are displayed in blue. Male genitalia (14) of selected species (dorsal views) are illustrated on the left. Topologies of collapsed clades (remaining Opatrini &amp; Dendarini + Pedininini + Playtonotini) are fully consistent with those presented in Lumen et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.5964431" httpUri="https://zenodo.org/record/5964431/files/figure.png" pageId="6" pageNumber="525">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B9736B67D0B151E79D6C904FACEFA8A" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
<taxonomicName id="4C284D357D0B151E79D6C904FE01FBD8" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="1851" box="[199,429,1086,1112]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E79D6C904FE01FBD8" box="[199,429,1086,1112]" italics="true" pageId="6" pageNumber="525">Lodinus punctulatus</emphasis>
</taxonomicName>
can be easily distinguished from its congeners by differing pronotal structure (
<bibRefCitation id="EFB94B477D0B151E7C20C904FF78FBFC" author="Marcuzzi, G." pageId="6" pageNumber="525" pagination="1 - 36" refId="ref9624" refString="Marcuzzi, G. (1954) Studies on the fauna of Curacao and other Caribbean Islands: no. 22. Tenebrionid beetles of Curacao, Aruba, Bonaire, and the Venezuelan Islands. Natuurwetenschappelijke Studiekring voor Suriname, 10, 1 - 36." type="journal article" year="1954">Marcuzzi 1954</bibRefCitation>
): anterior angles rounded, not prominent (pointed and prominent in remaining species); punctures sparsely distributed on disc (&gt;1.0 diameters apart vs. &lt;
<quantity id="4CD09B537D0B151E7BB2C9BFFD5FFB1F" box="[675,755,1157,1184]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="6" pageNumber="525" unit="in" value="0.75">0.75 in</quantity>
remaining species). Furthermore,
<taxonomicName id="4C284D357D0B151E7D6AC9BCFAB1FB1F" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="1851" box="[1147,1309,1157,1183]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E7D6AC9BCFAB1FB1F" box="[1147,1309,1157,1183]" italics="true" pageId="6" pageNumber="525">L. punctulatus</emphasis>
</taxonomicName>
is the only species within the genus which possesses subparallel parameres in their apical portion (slightly narrowing towards apex in
<emphasis id="B95CEAA47D0B151E79E0C9F7FE6CFB66" box="[241,448,1228,1254]" italics="true" pageId="6" pageNumber="525">
<taxonomicName id="4C284D357D0B151E79E0C9F7FE0FFB66" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="2022" box="[241,419,1228,1254]" class="Insecta" family="Tenebrionidae" genus="Austrocaribius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="venezuelensis">L. venezuelensis</taxonomicName>
</emphasis>
Figs 46 and
<quantity id="4CD09B537D0B151E7B5AC9F6FD29FB66" box="[587,645,1228,1254]" metricMagnitude="0" metricUnit="m" metricValue="1.1684" pageId="6" pageNumber="525" unit="in" value="46.0">46 in</quantity>
<bibRefCitation id="EFB94B477D0B151E7B9DC9F6FC94FB66" author="Marcuzzi, G." box="[652,824,1228,1254]" pageId="6" pageNumber="525" pagination="239 - 269" refId="ref9671" refString="Marcuzzi, G. (1983) Description of the external male genitalia of some neotropical tenebrionidae coleoptera heteromera. Folia Entomologica Hungarica, 44 (2), 239 - 269." type="journal article" year="1983">Marcuzzi 1983</bibRefCitation>
; strongly indented in
<taxonomicName id="4C284D357D0B151E7D3CC9F7FB04FB66" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="2022" box="[1069,1192,1229,1254]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="araguae">
<emphasis id="B95CEAA47D0B151E7D3CC9F7FB04FB66" box="[1069,1192,1229,1254]" italics="true" pageId="6" pageNumber="525">L. araguae</emphasis>
</taxonomicName>
<figureCitation id="13132A337D0B151E7DD6C9F6FAB2FB66" box="[1223,1310,1228,1254]" captionStart="FIGURE 3" captionStartId="9.[151,250,1859,1883]" captionTargetBox="[170,1399,602,1830]" captionTargetId="figure-194@9.[151,1436,597,1836]" captionTargetPageId="9" captionText="FIGURE 3. Morphology of female and male terminalia of selected Blapstinina species. (AC) Lodinus punctulatus comb. nov., (D) Lodinus araguae comb. nov., (E, F) Blapstinus tibialis. (A, B, E) ovipositor, (C, F) genital tubes, (D) parameres. Abbreviations: ag—accessory gland, b—hooked baculus of c4, c1c4—subsequent lobes of coxites, g—gonostylus, sp—sper- matheca, vag—vagina." figureDoi="http://doi.org/10.5281/zenodo.5964437" httpUri="https://zenodo.org/record/5964437/files/figure.png" pageId="6" pageNumber="525">Fig. 3D</figureCitation>
). Lastly, as of this redescription,
<taxonomicName id="4C284D357D0B151E7894C9CAFD8AFA89" authorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" authorityYear="2022" baseAuthorityName="Kamiński &amp; Lumen &amp; Kanda &amp; Smith" baseAuthorityYear="1851" box="[389,550,1263,1289]" class="Insecta" family="Tenebrionidae" genus="Lodinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="525" phylum="Arthropoda" rank="species" species="punctulatus">
<emphasis id="B95CEAA47D0B151E7894C9CAFD8AFA89" box="[389,550,1263,1289]" italics="true" pageId="6" pageNumber="525">L. punctulatus</emphasis>
</taxonomicName>
is the only species of the genus known from south of the equator (
<figureCitation id="13132A337D0B151E7C1EC9D5FAF8FA8A" box="[1295,1364,1263,1290]" captionStart="FIGURE 4" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[205,1382,181,1919]" captionTargetId="figure-23@10.[205,1382,181,1919]" captionTargetPageId="10" captionText="FIGURE 4. Distribution of Lodinus." figureDoi="http://doi.org/10.5281/zenodo.5964439" httpUri="https://zenodo.org/record/5964439/files/figure.png" pageId="6" pageNumber="525">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C332653D7D0B151E79D6C829FDB7F874" pageId="6" pageNumber="525" type="description">
<paragraph id="8B9736B67D0B151E79D6C829FDB7F874" blockId="6.[151,1437,150,2037]" pageId="6" pageNumber="525">
<emphasis id="B95CEAA47D0B151E79D6C829FEC1FAAD" bold="true" box="[199,365,1299,1325]" pageId="6" pageNumber="525">Redescription</emphasis>
. Length
<quantity id="4CD09B537D0B151E78DEC829FDFBFAAD" box="[463,599,1299,1325]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.2" metricValueMin="4.9" pageId="6" pageNumber="525" unit="mm" value="5.55" valueMax="6.2" valueMin="4.9">4.96.2 mm</quantity>
, width
<quantity id="4CD09B537D0B151E7BB6C829FC9CFAAD" box="[679,816,1299,1325]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="6" pageNumber="525" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.12.6 mm</quantity>
.
<emphasis id="B95CEAA47D0B151E7A28C829FCDBFAAD" bold="true" box="[825,887,1299,1325]" pageId="6" pageNumber="525">Body</emphasis>
(
<figureCitation id="13132A337D0B151E7A97C829FC4CFAAD" box="[902,992,1299,1325]" captionStart="FIGURE 2" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[163,1434,203,1471]" captionTargetId="figure-27@7.[151,1436,200,1474]" captionTargetPageId="7" captionText="FIGURE 2. Morphology of studied Blapstinina species. Habitus photographs and diagnostic features. (AC) Lodinus punctulatus comb. nov., (DF) Lodinus araguae comb. nov., (G) Blapstinus tibialis, (H, I) Blapstinus grandis. (B) protarsus, (C) mesotarsus, (D) last 5 antennomeres, (I) apical tip of epipleura." figureDoi="http://doi.org/10.5281/zenodo.5964433" httpUri="https://zenodo.org/record/5964433/files/figure.png" pageId="6" pageNumber="525">Fig. 2A</figureCitation>
): dull brown to black; dorsal and ventral surfaces matte; ovate-oblong; widest at 2/3 of elytral length; weakly convex in lateral view; evenly covered with fine setae dorsally.
<emphasis id="B95CEAA47D0B151E787FC860FE1BFAF4" bold="true" box="[366,439,1370,1396]" pageId="6" pageNumber="525">Head:</emphasis>
Epistoma broadly convex; punctures equal to diameter of an ommatidium (distributed 1 2 diameters apart); setae equal in length to intervals between punctures. Fronto-epistomal suture slightly marked on sides. Epistoma weakly emarginate.
<emphasis id="B95CEAA47D0B151E7B25C89BFD1BFA3B" bold="true" box="[564,695,1441,1467]" pageId="6" pageNumber="525">Prothorax:</emphasis>
Pronotum narrowing anteriorly, apical angles sharp. Lateral margin slightly rounded; marginated. Basal width slightly narrower than humeri; basal margin slightly bisinuate. Disc sparsely but noticeably covered with setae; finely punctate (punctures 13 diameters apart). Lateral sides marginated. Hypomeron dull and rugose; hypomeron impressed on sides. Intercoxal process of prosternum not depressed in lateral view.
<emphasis id="B95CEAA47D0B151E7850CB14FDAEF9C8" bold="true" box="[321,514,1582,1608]" pageId="6" pageNumber="525">Scutellar shield:</emphasis>
Pentagonal, of equal width and length, finely punctate.
<emphasis id="B95CEAA47D0B151E7D7CCB15FAABF9C9" bold="true" box="[1133,1287,1583,1609]" pageId="6" pageNumber="525">Pterothorax:</emphasis>
Elytral striae shallow and punctate (12 diameters apart). Striae not joining prior humeral angle. Intervals flat, punctate (0.51.0 diameters apart), with noticeable microsculpture. Epipleura finely punctate. Metathoracic wings well developed. Metaventrite long, sparsely punctate; sparse setae present.
<emphasis id="B95CEAA47D0B151E7A2BCBA3FCD0F933" bold="true" box="[826,892,1689,1715]" pageId="6" pageNumber="525">Legs:</emphasis>
Protibia apically dilated with lateral projection at apex. Meso and meta tibia straight and dilated apically. Pro- and mesotarsomeres 2 and 3 dilated in males (
<figureCitation id="13132A337D0B151E7C7ECB86FF72F97A" captionStart="FIGURE 2" captionStartId="7.[151,250,1498,1522]" captionTargetBox="[163,1434,203,1471]" captionTargetId="figure-27@7.[151,1436,200,1474]" captionTargetPageId="7" captionText="FIGURE 2. Morphology of studied Blapstinina species. Habitus photographs and diagnostic features. (AC) Lodinus punctulatus comb. nov., (DF) Lodinus araguae comb. nov., (G) Blapstinus tibialis, (H, I) Blapstinus grandis. (B) protarsus, (C) mesotarsus, (D) last 5 antennomeres, (I) apical tip of epipleura." figureDoi="http://doi.org/10.5281/zenodo.5964433" httpUri="https://zenodo.org/record/5964433/files/figure.png" pageId="6" pageNumber="525">Fig. 2B, C</figureCitation>
). Abdominal ventrites: Finely punctate, scattered with setae; intercoxal process narrowly rounded; ventrite 5 with posterior margin evenly rounded, not bordered.
<emphasis id="B95CEAA47D0B151E7A19CA39FCD7F89D" bold="true" box="[776,891,1795,1821]" pageId="6" pageNumber="525">Aedeagus</emphasis>
(
<figureCitation id="13132A337D0B151E7A9DCA39FC63F89E" box="[908,975,1795,1822]" captionStart="FIGURE 1" captionStartId="3.[151,250,1675,1699]" captionTargetBox="[151,1419,525,1640]" captionTargetId="figure-169@3.[151,1436,525,1651]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of subtribe Blapstinina. Figured topology was recovered in maximum likelihood analysis using IQ- Tree. Ultrafast bootstrap values (1,000 replicates) (integers) and posterior probabilities (20 million generations) acquired in Bayesian analysis (decimals) are displayed in red. The three investigated Blapstinina species are displayed in blue. Male genitalia (14) of selected species (dorsal views) are illustrated on the left. Topologies of collapsed clades (remaining Opatrini &amp; Dendarini + Pedininini + Playtonotini) are fully consistent with those presented in Lumen et al. (2020)." figureDoi="http://doi.org/10.5281/zenodo.5964431" httpUri="https://zenodo.org/record/5964431/files/figure.png" pageId="6" pageNumber="525">Fig. 1</figureCitation>
): whole tegmen evenly curved in lateral view. Apical part of aedeagus truncate; median lobe opening circular.
<emphasis id="B95CEAA47D0B151E7ABBCA1DFB28F8C1" bold="true" box="[938,1156,1831,1857]" pageId="6" pageNumber="525">Female terminalia</emphasis>
(
<figureCitation id="13132A337D0B151E7D85CA1DFABDF8C1" box="[1172,1297,1831,1857]" captionStart="FIGURE 3" captionStartId="9.[151,250,1859,1883]" captionTargetBox="[170,1399,602,1830]" captionTargetId="figure-194@9.[151,1436,597,1836]" captionTargetPageId="9" captionText="FIGURE 3. Morphology of female and male terminalia of selected Blapstinina species. (AC) Lodinus punctulatus comb. nov., (D) Lodinus araguae comb. nov., (E, F) Blapstinus tibialis. (A, B, E) ovipositor, (C, F) genital tubes, (D) parameres. Abbreviations: ag—accessory gland, b—hooked baculus of c4, c1c4—subsequent lobes of coxites, g—gonostylus, sp—sper- matheca, vag—vagina." figureDoi="http://doi.org/10.5281/zenodo.5964437" httpUri="https://zenodo.org/record/5964437/files/figure.png" pageId="6" pageNumber="525">Fig. 3AC</figureCitation>
): Ovipositor with paraprocts slightly longer than coxites. Paraproct does not shield valvifer and other lobes (directed basally). Valvifer wide and short; second lobe elongate; third one semicircular (about 0.5 of height of third one); apical lobe rounded, situated apically, sclerotised at outer and inner edges, inner sclerotisation hook-like, gonostyli on dorsal side, directed inwards. Proctiger covering nearly whole ovipositor. Vagina without sclerites. Spermatheca with narrow duct. Spiculum ventrale short.
</paragraph>
</subSubSection>
<subSubSection id="C332653D7D0A151F79D6CDACFBF9FF30" box="[199,1109,150,177]" pageId="7" pageNumber="526" type="distribution">
<paragraph id="8B9736B67D0A151F79D6CDACFBF9FF30" blockId="7.[199,1109,150,177]" box="[199,1109,150,177]" pageId="7" pageNumber="526">
<emphasis id="B95CEAA47D0A151F79D6CDACFEF5FF30" bold="true" box="[199,345,150,176]" pageId="7" pageNumber="526">Distribution</emphasis>
.
<collectingCountry id="F33F76267D0A151F7877CDACFDA0FF30" box="[358,524,150,176]" name="Argentina" pageId="7" pageNumber="526">ARGENTINA</collectingCountry>
,
<collectingCountry id="F33F76267D0A151F7B06CDACFDD1FF30" box="[535,637,150,176]" name="Brazil" pageId="7" pageNumber="526">BRASIL</collectingCountry>
,
<collectingCountry id="F33F76267D0A151F7B98CDACFD53FF30" box="[649,767,150,176]" name="Bolivia" pageId="7" pageNumber="526">BOLIVIA</collectingCountry>
,
<collectingCountry id="F33F76267D0A151F7A1BCDACFCF3FF30" box="[778,863,150,176]" name="Chile" pageId="7" pageNumber="526">CHILE</collectingCountry>
,
<collectingCountry id="F33F76267D0A151F7A7BCDACFC5EFF30" box="[874,1010,150,176]" name="Uruguay" pageId="7" pageNumber="526">URUGUAY</collectingCountry>
(
<figureCitation id="13132A337D0A151F7D13CDACFBEBFF30" box="[1026,1095,150,176]" captionStart="FIGURE 4" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[205,1382,181,1919]" captionTargetId="figure-23@10.[205,1382,181,1919]" captionTargetPageId="10" captionText="FIGURE 4. Distribution of Lodinus." figureDoi="http://doi.org/10.5281/zenodo.5964439" httpUri="https://zenodo.org/record/5964439/files/figure.png" pageId="7" pageNumber="526">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>