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<document id="02033DDBF90AB4433AC8DDAC6ED8028D" ID-DOI="10.11646/zootaxa.4759.1.4" ID-GBIF-Dataset="00adaef8-991b-47aa-9c33-12e2d46a5258" ID-ISSN="1175-5326" ID-Zenodo-Dep="3735986" ID-ZooBank="DA789391-F0EB-4CF2-A954-8CBACE37D57C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585730814225" checkinUser="plazi" docAuthor="Costa, Fábio Correia, Cherman, Mariana Alejandra &amp; Iannuzzi, Luciana" docDate="2020" docId="DD4387AED4338B65BCFCFB4BA36EAA69" docLanguage="en" docName="zootaxa.4759.1.4.pdf" docOrigin="Zootaxa 4759 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ovomanonychus Costa &amp; Cherman &amp; Iannuzzi 2020" docType="treatment" docVersion="5" lastPageNumber="70" masterDocId="217AFFD6D4328B60BC6BFFBFA015A80A" masterDocTitle="Ovomanonychus, a new genus of South American Sericoidini (Coleoptera Scarabaeidae: Melolonthinae)" masterLastPageNumber="76" masterPageNumber="65" pageNumber="66" updateTime="1698734145830" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="00233F5857A1034401887BF53315D45A">Ovomanonychus, a new genus of South American Sericoidini (Coleoptera Scarabaeidae: Melolonthinae)</mods:title>
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<mods:namePart id="AD8EF51C60BD9C95ADED37999FEEEAB0">Costa, Fábio Correia</mods:namePart>
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<mods:namePart id="533F09314F31955592881C3C3F720187">Cherman, Mariana Alejandra</mods:namePart>
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<mods:namePart id="3D1E5F6649703A1FF8F59ACC49179F3B">Iannuzzi, Luciana</mods:namePart>
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<mods:title id="E4898CBD249FFA863ED6B3E7CC479D65">Zootaxa</mods:title>
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<mods:date id="F92E5667C5559DEABBF9E1E8613F5FFD">2020</mods:date>
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<treatment id="DD4387AED4338B65BCFCFB4BA36EAA69" ID-DOI="http://doi.org/10.5281/zenodo.3811909" ID-GBIF-Taxon="163420510" ID-Zenodo-Dep="3811909" LSID="urn:lsid:plazi:treatment:DD4387AED4338B65BCFCFB4BA36EAA69" httpUri="http://treatment.plazi.org/id/DD4387AED4338B65BCFCFB4BA36EAA69" lastPageId="5" lastPageNumber="70" pageId="1" pageNumber="66">
<subSubSection id="1DF06533D4338B61BCFCFB4BA371AD05" box="[151,868,1268,1295]" pageId="1" pageNumber="66" type="nomenclature">
<paragraph id="555536B8D4338B61BCFCFB4BA371AD05" blockId="1.[151,868,1268,1295]" box="[151,868,1268,1295]" pageId="1" pageNumber="66">
<heading id="0E1D81D4D4338B61BCFCFB4BA371AD05" bold="true" box="[151,868,1268,1295]" fontSize="11" level="1" pageId="1" pageNumber="66" reason="1">
<emphasis id="679EEAAAD4338B61BCFCFB4BA371AD05" bold="true" box="[151,868,1268,1295]" pageId="1" pageNumber="66">
<taxonomicName id="92EA4D3BD4338B61BCFCFB4BA176AD05" ID-CoL="6B5L" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[151,355,1268,1295]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4338B61BCFCFB4BA176AD05" bold="true" box="[151,355,1268,1295]" italics="true" pageId="1" pageNumber="66">Ovomanonychus</emphasis>
</taxonomicName>
Costa, Cherman, &amp; Iannuzzi, new genus
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1DF06533D4338B61BCFCFA82A379AD5C" box="[151,876,1340,1367]" pageId="1" pageNumber="66" type="type_taxon">
<paragraph id="555536B8D4338B61BCFCFA82A379AD5C" blockId="1.[151,1437,1340,1979]" box="[151,876,1340,1367]" pageId="1" pageNumber="66">
<emphasis id="679EEAAAD4338B61BCFCFA82A13EAD5D" bold="true" box="[151,299,1340,1367]" pageId="1" pageNumber="66">
<typeStatus id="8A51881AD4338B61BCFCFA82A0C7AD5D" box="[151,210,1341,1367]" pageId="1" pageNumber="66">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="92EA4D3BD4338B61BD52FA83A2BFAD5C" authority="Frey, 1976" authorityName="Frey" authorityYear="1976" box="[313,682,1340,1367]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4338B61BD52FA83A23DAD5C" box="[313,552,1340,1366]" italics="true" pageId="1" pageNumber="66">Manonychus rosettae</emphasis>
<bibRefCitation id="317B4B49D4338B61BE44FA82A2BFAD5C" author="Frey, G." box="[559,682,1340,1367]" pageId="1" pageNumber="66" pagination="375 - 388" refId="ref5976" refString="Frey, G. (1976) Neue Sericinae und Macrodactylini aus Brasilien (Coleoptera Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 27, 375 - 388." type="journal article" year="1976">Frey, 1976</bibRefCitation>
</taxonomicName>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="1DF06533D4338B61BCACFADFA431AE90" pageId="1" pageNumber="66" type="diagnosis">
<paragraph id="555536B8D4338B61BCACFADFA431AE90" blockId="1.[151,1437,1340,1979]" pageId="1" pageNumber="66">
<emphasis id="679EEAAAD4338B61BCACFADFA157AD71" bold="true" box="[199,322,1376,1403]" pageId="1" pageNumber="66">Diagnosis.</emphasis>
<taxonomicName id="92EA4D3BD4338B61BD22FADFA211AD70" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[329,516,1376,1402]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4338B61BD22FADFA211AD70" box="[329,516,1376,1402]" italics="true" pageId="1" pageNumber="66">Ovomanonychus</emphasis>
</taxonomicName>
is distinguished from all other Sericoidini by the following combination of characters: ovate body; labrum vertical, with superior margin inflated, hidden below the clypeus, not on same plane; galea with outer margin curved; labium longer than it is wide; antennae with nine antennomeres and club (with three antennomeres) longer than flagellum; absence of membrane along median of anterior pronotal margin; six fused ventrites, except the VI articulate; ventrite I with basal half hidden by metacoxae and ventrite VI one quarter the length of other ventrites; ventrite V and propygidium separated by suture; tarsomeres with sparse setae ventrally; elytra with strong costae; procoxae conical; apical protibial spur in both sexes; mesocoxae three times longer than wide; metacoxae longer than ventrite II; mesotarsomere I longer than mesotarsomere II; claws simple on all tarsi; metatibia with two apical spurs, spurs set below and above the tarsal articulation.
</paragraph>
</subSubSection>
<subSubSection id="1DF06533D4338B64BCACF91BA2ECAF55" lastPageId="4" lastPageNumber="69" pageId="1" pageNumber="66" type="description">
<paragraph id="555536B8D4338B64BCACF91BA27DAF1D" blockId="1.[151,1437,1340,1979]" lastBlockId="4.[151,1437,1213,2032]" lastPageId="4" lastPageNumber="69" pageId="1" pageNumber="66">
<emphasis id="679EEAAAD4338B61BCACF91BA14CAEB5" bold="true" box="[199,345,1700,1727]" pageId="1" pageNumber="66">Description.</emphasis>
Length: 10.012.0 mm; width: 4.0
<quantity id="92129B5DD4338B61BE92F91BA35CAEB5" box="[761,841,1700,1727]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="66" unit="mm" value="5.5">5.5mm</quantity>
. Body reddish brown, ovate sides slightly arcuate at middle of elytra (
<figureCitation id="CDD12A3DD4338B61BD33F977A1A4AEE9" box="[344,433,1736,1763]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="1" pageNumber="66">Fig. 1A</figureCitation>
).
<emphasis id="679EEAAAD4338B61BDAEF977A217AEE8" box="[453,514,1736,1762]" italics="true" pageId="1" pageNumber="66">Head</emphasis>
: Anterior margin rounded; distance between eyes twice or three times the width of one eye; frons densely punctate; frontoclypeal suture strongly impressed; canthus covered by clypeus; clypeus trapezoidal, punctate (
<figureCitation id="CDD12A3DD4338B61BD0EF8AFA1AFAF21" box="[357,442,1808,1835]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="1" pageNumber="66">Fig. 1E</figureCitation>
), strongly reflexed, and separated from labrum by a suture; labrum with superior margin bulged, vertical, hidden below the clypeus, not on same plane; galea of maxilla with outer margin curved (
<figureCitation id="CDD12A3DD4338B61B95DF88BA598AF45" box="[1334,1421,1844,1871]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="1" pageNumber="66">Fig. 1B</figureCitation>
); galea with tooth I projected; lacinia with distal portion forming a toothlike projection; labium densely setose along the lateral margin, longer than wide (
<figureCitation id="CDD12A3DD4338B61BE4DF8C3A26EAF9D" box="[550,635,1916,1943]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="1" pageNumber="66">Fig. 1D</figureCitation>
); palp insertion covered by the labium and placed submedially; ligula and labium fused; antennae with nine antennomeres, club with three antennomeres and longer than flagellum (
<figureCitation id="CDD12A3DD4338B61B952F81FA59BAFB1" box="[1337,1422,1952,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="1" pageNumber="66">Fig. 1E</figureCitation>
).
<emphasis id="679EEAAAD4308B62BCFCF88EA11CAF41" box="[151,265,1841,1867]" italics="true" pageId="2" pageNumber="67">Prothorax</emphasis>
: Pronotum wider than long, glabrous, densely punctate, disc narrowly smooth along midline; absence of membrane along median of anterior pronotal margin (
<figureCitation id="CDD12A3DD4308B62BFCFF8EAA417AF65" box="[932,1026,1877,1903]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="2" pageNumber="67">Fig. 1E</figureCitation>
); anterolateral region slightly depressed; lateral margin sparsely setose; anterior angle rounded; anterolateral margin arcuate, posterolateral margins straight; proepimeron smooth laterally.
<emphasis id="679EEAAAD4308B62BEBFF822A34FAFBD" box="[724,858,1949,1975]" italics="true" pageId="2" pageNumber="67">Pterothorax</emphasis>
: Scutellum subtriangular; mesosternum with foveolate, bristled punctures; internal posterior angle of mesepimeron acute and projected; metasternum bristles long; distance between the mesocoxae and metacoxae equal to length of metacoxae; metepisternum with coarse punctures.
<emphasis id="679EEAAAD4368B64BD7AFB5DA142ACF6" box="[273,343,1250,1276]" italics="true" pageId="4" pageNumber="69">Elytra</emphasis>
: Shiny, with costae parallel to the elytral suture, which is elevated (
<figureCitation id="CDD12A3DD4368B64B82EFB5EA488ACF6" box="[1093,1181,1249,1276]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="4" pageNumber="69">Fig. 1A</figureCitation>
).
<emphasis id="679EEAAAD4368B64B8DAFB5DA4F3ACF1" box="[1201,1254,1250,1275]" italics="true" pageId="4" pageNumber="69">Legs</emphasis>
: Procoxae conical, with sparse punctures on proximal region; profemurs with aggregated punctures extending from the base to the disc; protibia with sculpture and three teeth well developed, tooth II forming an acute angle with tooth III; apical protibial spur in both sexes; protarsomeres cylindrical; mesocoxae contiguous, punctate distally and three times longer than wide; mesofemur with parallel double row of punctures; mesotibia gradually enlarged towards apex, and with two transverse carinae; transverse carina II incomplete (non-reaching inner margin of mesotibia); two apical spurs, spurs set below and above the tarsal articulation; mesotarsomere I longer than mesotarsomere II (
<figureCitation id="CDD12A3DD4368B64B958FA06A59AADDE" box="[1331,1423,1465,1492]" captionStart="FIGURE 2" captionStartId="3.[151,250,1837,1862]" captionTargetBox="[160,1411,196,1810]" captionTargetId="figure@3.[151,1436,181,1811]" captionTargetPageId="3" captionText="FIGURE 2. Ovomanonychus diagnostic characters. A, Ovomanonychus rosettae (Frey, 1976) mesotarsomere; B, O. inajae new species spurs set below and above the tarsal articulation; C, O. rosettae metacoxae longer than ventrite II; D, Manonychus ovalis (Blanchard, 1851) metacoxae length equal to ventrite II; E, O. striatus new species metacoxae densely bristled; F, O. rosettae apical region of phallobase excavated." figureDoi="http://doi.org/10.5281/zenodo.3735992" httpUri="https://zenodo.org/record/3735992/files/figure.png" pageId="4" pageNumber="69">Fig. 2A</figureCitation>
); metacoxae longer than ventrite II, with dense or sparse bristles; metacoxal lateral margin in obtuse angle; metafemur with row of punctures posteriorly and smooth basally; transverse carina II incomplete; metatibial apical spurs with equal or different size (
<figureCitation id="CDD12A3DD4368B64BDFCF99AA1F8AE4A" box="[407,493,1573,1600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1837,1862]" captionTargetBox="[160,1411,196,1810]" captionTargetId="figure@3.[151,1436,181,1811]" captionTargetPageId="3" captionText="FIGURE 2. Ovomanonychus diagnostic characters. A, Ovomanonychus rosettae (Frey, 1976) mesotarsomere; B, O. inajae new species spurs set below and above the tarsal articulation; C, O. rosettae metacoxae longer than ventrite II; D, Manonychus ovalis (Blanchard, 1851) metacoxae length equal to ventrite II; E, O. striatus new species metacoxae densely bristled; F, O. rosettae apical region of phallobase excavated." figureDoi="http://doi.org/10.5281/zenodo.3735992" httpUri="https://zenodo.org/record/3735992/files/figure.png" pageId="4" pageNumber="69">Fig. 2B</figureCitation>
), spurs set below and above the tarsal articulation; apex of metatibia prominent at the tarsal articulation; ventral surface of all tarsomeres with sparse bristles (
<figureCitation id="CDD12A3DD4368B64BFA1F9F6A435AE6E" box="[970,1056,1609,1636]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="4" pageNumber="69">Fig. 1F</figureCitation>
); all claws simple.
<emphasis id="679EEAAAD4368B64B896F9F5A572AE6E" box="[1277,1383,1610,1636]" italics="true" pageId="4" pageNumber="69">Abdomen</emphasis>
: Six fused ventrites, except ventrite VI which is articulate; ventrite I with basal half hidden by metacoxae and ventrite VI one quarter the length the others (
<figureCitation id="CDD12A3DD4368B64BE5BF92EA29DAEA6" box="[560,648,1681,1708]" captionStart="FIGURE 2" captionStartId="3.[151,250,1837,1862]" captionTargetBox="[160,1411,196,1810]" captionTargetId="figure@3.[151,1436,181,1811]" captionTargetPageId="3" captionText="FIGURE 2. Ovomanonychus diagnostic characters. A, Ovomanonychus rosettae (Frey, 1976) mesotarsomere; B, O. inajae new species spurs set below and above the tarsal articulation; C, O. rosettae metacoxae longer than ventrite II; D, Manonychus ovalis (Blanchard, 1851) metacoxae length equal to ventrite II; E, O. striatus new species metacoxae densely bristled; F, O. rosettae apical region of phallobase excavated." figureDoi="http://doi.org/10.5281/zenodo.3735992" httpUri="https://zenodo.org/record/3735992/files/figure.png" pageId="4" pageNumber="69">Fig. 2C</figureCitation>
); lateral keel along the ventrites; ventrites IIVI with transverse row of setose punctures; pygidium and propygidium separated by suture; propygidium punctate.
<emphasis id="679EEAAAD4368B64B81CF909A50FAEDA" box="[1143,1306,1718,1744]" italics="true" pageId="4" pageNumber="69">Male genitalia</emphasis>
: Parameres symmetrical, converging at apex; outer margin slightly curved on distal third; inner margin on distal portion curved; medial portion of proximal margin lobed.
</paragraph>
<caption id="01956630D4308B62BCFCF9C1A32FAF09" ID-DOI="http://doi.org/10.5281/zenodo.3735988" ID-Zenodo-Dep="3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="2" pageNumber="67" startId="2.[151,250,1662,1687]" targetBox="[170,1408,192,1623]" targetPageId="2">
<paragraph id="555536B8D4308B62BCFCF9C1A32FAF09" blockId="2.[151,1437,1662,1795]" pageId="2" pageNumber="67">
<emphasis id="679EEAAAD4308B62BCFCF9C1A107AE9D" bold="true" box="[151,274,1662,1687]" pageId="2" pageNumber="67">FIGURE 1.</emphasis>
<taxonomicName id="92EA4D3BD4308B62BD7CF9C0A1D5AE9D" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[279,448,1663,1687]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4308B62BD7CF9C0A1D5AE9D" box="[279,448,1663,1687]" italics="true" pageId="2" pageNumber="67">Ovomanonychus</emphasis>
</taxonomicName>
diagnostic characters.
<emphasis id="679EEAAAD4308B62BECFF9C1A2A3AE9D" bold="true" box="[676,694,1662,1687]" pageId="2" pageNumber="67">A</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62BED6F9C0A3A2AE9D" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[701,951,1663,1687]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="679EEAAAD4308B62BED6F9C0A3A2AE9D" box="[701,951,1663,1687]" italics="true" pageId="2" pageNumber="67">Ovomanonychus striatus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4308B62BFD7F9C0A42DAE9D" bold="true" box="[956,1080,1662,1687]" pageId="2" pageNumber="67">
<taxonomicNameLabel id="7CAD57D1D4308B62BFD7F9C0A42DAE9D" box="[956,1080,1662,1687]" pageId="2" pageNumber="67" rank="species">new species</taxonomicNameLabel>
</emphasis>
male paratype dorsal habitus;
<emphasis id="679EEAAAD4308B62B90CF9C0A56DAE9D" bold="true" box="[1383,1400,1663,1687]" pageId="2" pageNumber="67">B</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62B9EFF9C0A0F6AEB1" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="striatus">
<emphasis id="679EEAAAD4308B62B9EFF9C0A0F6AEB1" italics="true" pageId="2" pageNumber="67">O. striatus</emphasis>
</taxonomicName>
right maxilla, galea with outer margin curved;
<emphasis id="679EEAAAD4308B62BEAFF91DA2C3AEB1" bold="true" box="[708,726,1698,1723]" pageId="2" pageNumber="67">C</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62BE88F91CA476AEB1" authority="Martinez, 1959" authorityName="Martinez" authorityYear="1959" box="[739,1123,1698,1723]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="birabeni">
<emphasis id="679EEAAAD4308B62BE88F91CA3D4AEB1" box="[739,961,1699,1723]" italics="true" pageId="2" pageNumber="67">Manonychus birabeni</emphasis>
Martínez, 1959
</taxonomicName>
right maxilla, galea with outer margin lobed;
<emphasis id="679EEAAAD4308B62BD41F978A129AED5" bold="true" box="[298,316,1735,1759]" pageId="2" pageNumber="67">D</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62BD23F978A1A6AED5" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[328,435,1735,1759]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="striatus">
<emphasis id="679EEAAAD4308B62BD23F978A1A6AED5" box="[328,435,1735,1759]" italics="true" pageId="2" pageNumber="67">O. striatus</emphasis>
</taxonomicName>
labium;
<emphasis id="679EEAAAD4308B62BE66F978A20BAED5" bold="true" box="[525,542,1735,1759]" pageId="2" pageNumber="67">E</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62BE41F978A309AED5" authority="(Frey, 1976)" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[554,796,1734,1759]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4308B62BE41F978A229AED4" box="[554,572,1735,1758]" italics="true" pageId="2" pageNumber="67">O</emphasis>
.
<emphasis id="679EEAAAD4308B62BE22F977A28FAED5" box="[585,666,1736,1759]" italics="true" pageId="2" pageNumber="67">rosettae</emphasis>
(
<bibRefCitation id="317B4B49D4308B62BECDF978A300AED5" author="Frey, G." box="[678,789,1734,1759]" pageId="2" pageNumber="67" pagination="375 - 388" refId="ref5976" refString="Frey, G. (1976) Neue Sericinae und Macrodactylini aus Brasilien (Coleoptera Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 27, 375 - 388." type="journal article" year="1976">Frey, 1976</bibRefCitation>
)
</taxonomicName>
frons and antennae;
<emphasis id="679EEAAAD4308B62BF85F978A3E8AED5" bold="true" box="[1006,1021,1735,1759]" pageId="2" pageNumber="67">F</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62B861F978A46CAED5" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[1034,1145,1735,1759]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4308B62B861F978A409AED4" box="[1034,1052,1735,1758]" italics="true" pageId="2" pageNumber="67">O</emphasis>
.
<emphasis id="679EEAAAD4308B62B843F977A46CAED5" box="[1064,1145,1736,1759]" italics="true" pageId="2" pageNumber="67">rosettae</emphasis>
</taxonomicName>
left anterior tarsomeres with sparse bristles;
<emphasis id="679EEAAAD4308B62BD58F955A152AF09" bold="true" box="[307,327,1770,1795]" pageId="2" pageNumber="67">G</emphasis>
,
<taxonomicName id="92EA4D3BD4308B62BD3FF953A1D9AF09" authorityName="Martinez" authorityYear="1959" box="[340,460,1771,1795]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="birabeni">
<emphasis id="679EEAAAD4308B62BD3FF953A1D9AF09" box="[340,460,1771,1795]" italics="true" pageId="2" pageNumber="67">M. birabeni</emphasis>
</taxonomicName>
right anterior tarsomeres with pads.
</paragraph>
</caption>
<caption id="01956630D4318B63BCFCF892A209AFB8" ID-DOI="http://doi.org/10.5281/zenodo.3735992" ID-Zenodo-Dep="3735992" httpUri="https://zenodo.org/record/3735992/files/figure.png" pageId="3" pageNumber="68" startId="3.[151,250,1837,1862]" targetBox="[160,1411,196,1810]" targetPageId="3">
<paragraph id="555536B8D4318B63BCFCF892A209AFB8" blockId="3.[151,1437,1837,1970]" pageId="3" pageNumber="68">
<emphasis id="679EEAAAD4318B63BCFCF892A107AF4C" bold="true" box="[151,274,1837,1862]" pageId="3" pageNumber="68">FIGURE 2.</emphasis>
<taxonomicName id="92EA4D3BD4318B63BD73F892A1D4AF4F" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[280,449,1837,1861]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4318B63BD73F892A1D4AF4F" box="[280,449,1837,1861]" italics="true" pageId="3" pageNumber="68">Ovomanonychus</emphasis>
</taxonomicName>
diagnostic characters.
<emphasis id="679EEAAAD4318B63BECDF892A2ADAF4C" bold="true" box="[678,696,1837,1862]" pageId="3" pageNumber="68">A</emphasis>
,
<taxonomicName id="92EA4D3BD4318B63BEAEF892A450AF4C" authority="(Frey, 1976)" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[709,1093,1837,1862]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4318B63BEAEF892A3D1AF4F" box="[709,964,1837,1861]" italics="true" pageId="3" pageNumber="68">Ovomanonychus rosettae</emphasis>
(
<bibRefCitation id="317B4B49D4318B63BFBBF892A42BAF4C" author="Frey, G." box="[976,1086,1837,1862]" pageId="3" pageNumber="68" pagination="375 - 388" refId="ref5976" refString="Frey, G. (1976) Neue Sericinae und Macrodactylini aus Brasilien (Coleoptera Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 27, 375 - 388." type="journal article" year="1976">Frey, 1976</bibRefCitation>
)
</taxonomicName>
mesotarsomere;
<emphasis id="679EEAAAD4318B63B89AF891A517AF4C" bold="true" box="[1265,1282,1838,1862]" pageId="3" pageNumber="68">B</emphasis>
,
<taxonomicName id="92EA4D3BD4318B63B965F891A57EAF4F" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[1294,1387,1838,1861]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="inajae" status="sp. nov.">
<emphasis id="679EEAAAD4318B63B965F891A57EAF4F" box="[1294,1387,1838,1861]" italics="true" pageId="3" pageNumber="68">O. inajae</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4318B63B91AF891A0F7AF60" bold="true" pageId="3" pageNumber="68">
<taxonomicNameLabel id="7CAD57D1D4318B63B91AF891A0F7AF60" pageId="3" pageNumber="68" rank="species">new species</taxonomicNameLabel>
</emphasis>
spurs set below and above the tarsal articulation;
<emphasis id="679EEAAAD4318B63BEB9F8EEA2F1AF60" bold="true" box="[722,740,1873,1898]" pageId="3" pageNumber="68">C</emphasis>
,
<taxonomicName id="92EA4D3BD4318B63BE84F8EDA34BAF63" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[751,862,1874,1897]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4318B63BE84F8EDA34BAF63" box="[751,862,1874,1897]" italics="true" pageId="3" pageNumber="68">O. rosettae</emphasis>
</taxonomicName>
metacoxae longer than ventrite II;
<emphasis id="679EEAAAD4318B63B8D0F8EDA4D8AF60" bold="true" box="[1211,1229,1874,1898]" pageId="3" pageNumber="68">D</emphasis>
,
<taxonomicName id="92EA4D3BD4318B63B8B2F8EEA15AAF84" authority="(Blanchard, 1851)" baseAuthorityName="Blanchard" baseAuthorityYear="1851" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="679EEAAAD4318B63B8B2F8EEA588AF63" box="[1241,1437,1873,1897]" italics="true" pageId="3" pageNumber="68">Manonychus ovalis</emphasis>
(Blanchard, 1851)
</taxonomicName>
metacoxae length equal to ventrite II;
<emphasis id="679EEAAAD4318B63BEB0F8C9A2F9AF84" bold="true" box="[731,748,1910,1934]" pageId="3" pageNumber="68">E</emphasis>
,
<taxonomicName id="92EA4D3BD4318B63BE92F8C9A371AF87" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[761,868,1910,1933]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="679EEAAAD4318B63BE92F8C9A371AF87" box="[761,868,1910,1933]" italics="true" pageId="3" pageNumber="68">O. striatus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4318B63BF00F8C9A3FDAF84" bold="true" box="[875,1000,1909,1934]" pageId="3" pageNumber="68">
<taxonomicNameLabel id="7CAD57D1D4318B63BF00F8C9A3FDAF84" box="[875,1000,1909,1934]" pageId="3" pageNumber="68" rank="species">new species</taxonomicNameLabel>
</emphasis>
metacoxae densely bristled;
<emphasis id="679EEAAAD4318B63B97BF8C9A50AAF84" bold="true" box="[1296,1311,1910,1934]" pageId="3" pageNumber="68">F</emphasis>
,
<taxonomicName id="92EA4D3BD4318B63B947F8C9A588AF87" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[1324,1437,1910,1933]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4318B63B947F8C9A588AF87" box="[1324,1437,1910,1933]" italics="true" pageId="3" pageNumber="68">O. rosettae</emphasis>
</taxonomicName>
apical region of phallobase excavated.
</paragraph>
</caption>
<caption id="01956630D4368B64BCFCFBECA43FAC9A" ID-DOI="http://doi.org/10.5281/zenodo.3735994" ID-Zenodo-Dep="3735994" httpUri="https://zenodo.org/record/3735994/files/figure.png" pageId="4" pageNumber="69" startId="4.[151,250,1107,1132]" targetBox="[157,1429,188,1074]" targetPageId="4">
<paragraph id="555536B8D4368B64BCFCFBECA43FAC9A" blockId="4.[151,1436,1107,1168]" pageId="4" pageNumber="69">
<emphasis id="679EEAAAD4368B64BCFCFBECA106AC66" bold="true" box="[151,275,1107,1132]" pageId="4" pageNumber="69">FIGURE 3.</emphasis>
Distribution map of
<taxonomicName id="92EA4D3BD4368B64BD81FBECA286AC61" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[490,659,1107,1131]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="679EEAAAD4368B64BD81FBECA286AC61" box="[490,659,1107,1131]" italics="true" pageId="4" pageNumber="69">Ovomanonychus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4368B64BEF1FBECA31FAC61" bold="true" box="[666,778,1107,1131]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="7CAD57D1D4368B64BEF1FBECA31FAC61" box="[666,778,1107,1131]" pageId="4" pageNumber="69" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
:
<taxonomicName id="92EA4D3BD4368B64BF73FBECA412AC61" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[792,1031,1107,1131]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="inajae" status="sp. nov.">
<emphasis id="679EEAAAD4368B64BF73FBECA412AC61" box="[792,1031,1107,1131]" italics="true" pageId="4" pageNumber="69">Ovomanonychus inajae</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4368B64B865FBECA499AC66" bold="true" box="[1038,1164,1107,1132]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="7CAD57D1D4368B64B865FBECA499AC66" box="[1038,1164,1107,1132]" pageId="4" pageNumber="69" rank="species">new species</taxonomicNameLabel>
</emphasis>
(red square),
<taxonomicName id="92EA4D3BD4368B64B972FBEBA10DAC85" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="679EEAAAD4368B64B972FBEBA10DAC85" italics="true" pageId="4" pageNumber="69">Ovomanonychus striatus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4368B64BD75FBC8A18EAC9A" bold="true" box="[286,411,1143,1168]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="7CAD57D1D4368B64BD75FBC8A18EAC9A" box="[286,411,1143,1168]" pageId="4" pageNumber="69" rank="species">new species</taxonomicNameLabel>
</emphasis>
(yellow circles), and
<taxonomicName id="92EA4D3BD4368B64BE1CFBC8A362AC85" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[631,887,1143,1167]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="rosettae">
<emphasis id="679EEAAAD4368B64BE1CFBC8A362AC85" box="[631,887,1143,1167]" italics="true" pageId="4" pageNumber="69">Ovomanonychus rosettae</emphasis>
</taxonomicName>
(blue diamonds).
</paragraph>
</caption>
<paragraph id="555536B8D4368B64BCACF89EA2ECAF55" blockId="4.[151,1437,1213,2032]" pageId="4" pageNumber="69">
<emphasis id="679EEAAAD4368B64BCACF89EA1A3AF36" bold="true" box="[199,438,1825,1852]" pageId="4" pageNumber="69">Sexual dimorphism.</emphasis>
Males and females are quite similar, except in the protibial tooth I stronger in females and pygidium slightly convex in females and flat in males.
</paragraph>
</subSubSection>
<subSubSection id="1DF06533D4368B65BCACF8D5A57DA8D2" lastPageId="5" lastPageNumber="70" pageId="4" pageNumber="69" type="etymology">
<paragraph id="555536B8D4368B64BCACF8D5A4B6AFAD" blockId="4.[151,1437,1213,2032]" pageId="4" pageNumber="69">
<emphasis id="679EEAAAD4368B64BCACF8D5A15BAF8E" bold="true" box="[199,334,1898,1924]" pageId="4" pageNumber="69">Etymology.</emphasis>
Adjective in the nominative singular. From the Latin
<emphasis id="679EEAAAD4368B64BFC6F8D5A3FFAF89" box="[941,1002,1898,1923]" italics="true" pageId="4" pageNumber="69">ovum</emphasis>
(“egg”) +
<taxonomicName id="92EA4D3BD4368B64B808F8D5A4E7AF8E" authorityName="Moser" authorityYear="1919" box="[1123,1266,1898,1924]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4368B64B808F8D5A4E7AF8E" box="[1123,1266,1898,1924]" italics="true" pageId="4" pageNumber="69">Manonychus</emphasis>
</taxonomicName>
in reference to the body shape and original placement of the
<typeStatus id="8A51881AD4368B64BEFEF831A2D0AFA2" box="[661,709,1934,1960]" pageId="4" pageNumber="69">type</typeStatus>
species. The name is masculine in gender.
</paragraph>
<paragraph id="555536B8D4368B64BCACF80EA261AFFA" blockId="4.[151,1437,1213,2032]" pageId="4" pageNumber="69">
<emphasis id="679EEAAAD4368B64BCACF80EA172AFC6" bold="true" box="[199,359,1969,1996]" pageId="4" pageNumber="69">Composition.</emphasis>
<taxonomicName id="92EA4D3BD4368B64BD05F80DA30FAFC6" authority="(Frey, 1976)" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" baseAuthorityName="Frey" baseAuthorityYear="1976" box="[366,794,1969,1996]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="rosettae" status="comb. nov.">
<emphasis id="679EEAAAD4368B64BD05F80DA29CAFC1" box="[366,649,1970,1996]" italics="true" pageId="4" pageNumber="69">Ovomanonychus rosettae</emphasis>
(
<bibRefCitation id="317B4B49D4368B64BEF3F80DA307AFC6" author="Frey, G." box="[664,786,1969,1996]" pageId="4" pageNumber="69" pagination="375 - 388" refId="ref5976" refString="Frey, G. (1976) Neue Sericinae und Macrodactylini aus Brasilien (Coleoptera Melolonthidae). Entomologische Arbeiten aus dem Museum G. Frey, 27, 375 - 388." type="journal article" year="1976">Frey, 1976</bibRefCitation>
)
</taxonomicName>
<emphasis id="679EEAAAD4368B64BF4AF80DA3FBAFC6" bold="true" box="[801,1006,1969,1996]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="7CAD57D1D4368B64BF4AF80DA3FBAFC6" box="[801,1006,1969,1996]" pageId="4" pageNumber="69" rank="species">new combination</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="92EA4D3BD4368B64BF97F80DA511AFC1" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[1020,1284,1970,1996]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="inajae" status="sp. nov.">
<emphasis id="679EEAAAD4368B64BF97F80DA511AFC1" box="[1020,1284,1970,1996]" italics="true" pageId="4" pageNumber="69">Ovomanonychus inajae</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4368B64B960F80DA580AFC6" bold="true" box="[1291,1429,1969,1996]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="7CAD57D1D4368B64B960F80DA580AFC6" box="[1291,1429,1969,1996]" pageId="4" pageNumber="69" rank="species">new species</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="92EA4D3BD4368B64BCACF869A1C9AFE5" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[199,476,2006,2032]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
<emphasis id="679EEAAAD4368B64BCACF869A1C9AFE5" box="[199,476,2006,2032]" italics="true" pageId="4" pageNumber="69">Ovomanonychus striatus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4368B64BD88F869A279AFFA" bold="true" box="[483,620,2005,2032]" pageId="4" pageNumber="69">
<taxonomicNameLabel id="7CAD57D1D4368B64BD88F869A279AFFA" box="[483,620,2005,2032]" pageId="4" pageNumber="69" rank="species">new species</taxonomicNameLabel>
</emphasis>
.
</paragraph>
<paragraph id="555536B8D4378B65BCACFF27A57DA8D2" blockId="5.[151,1437,152,611]" pageId="5" pageNumber="70">
<emphasis id="679EEAAAD4378B65BCACFF27A21CA8B9" bold="true" box="[199,521,152,179]" pageId="5" pageNumber="70">Geographical distribution.</emphasis>
Endemic to the Midwest (
<collectingRegion id="972EF85AD4378B65BF2CFF26A3F6A8B9" box="[839,995,153,179]" country="Brazil" name="Mato Grosso" pageId="5" pageNumber="70">Mato Grosso</collectingRegion>
), Southeastern (
<collectingRegion id="972EF85AD4378B65B8F5FF26A552A8B9" box="[1182,1351,153,179]" country="Brazil" name="Espirito Santo" pageId="5" pageNumber="70">Espírito Santo</collectingRegion>
,
<collectingRegion id="972EF85AD4378B65B93DFF26A0F6A8DD" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="70">Minas Gerais</collectingRegion>
,
<collectingRegion id="972EF85AD4378B65BC85FF02A186A8DD" box="[238,403,189,215]" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="70">Rio de Janeiro</collectingRegion>
, and
<collectingRegion id="972EF85AD4378B65BDA6FF02A257A8D2" box="[461,578,189,216]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="70">São Paulo</collectingRegion>
), Northeastern (
<collectingRegion id="972EF85AD4378B65BE9EFF02A329A8D2" box="[757,828,189,216]" country="Brazil" name="Bahia" pageId="5" pageNumber="70">Bahia</collectingRegion>
), and Southern (
<collectingRegion id="972EF85AD4378B65BF99FF02A457A8DD" box="[1010,1090,189,215]" country="Brazil" name="Parana" pageId="5" pageNumber="70">Paraná</collectingRegion>
) regions of
<collectingCountry id="2DFD7628D4378B65B8A3FF02A519A8DD" box="[1224,1292,189,215]" name="Brazil" pageId="5" pageNumber="70">Brazil</collectingCountry>
(
<figureCitation id="CDD12A3DD4378B65B976FF02A54EA8D2" box="[1309,1371,189,216]" captionStart="FIGURE 3" captionStartId="4.[151,250,1107,1132]" captionTargetBox="[157,1429,188,1074]" captionTargetId="figure@4.[151,1436,181,1081]" captionTargetPageId="4" captionText="FIGURE 3. Distribution map of Ovomanonychus new genus: Ovomanonychus inajae new species (red square), Ovomanonychus striatus new species (yellow circles), and Ovomanonychus rosettae (blue diamonds)." figureDoi="http://doi.org/10.5281/zenodo.3735994" httpUri="https://zenodo.org/record/3735994/files/figure.png" pageId="5" pageNumber="70">Fig.3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1DF06533D4378B65BCACFF5EA36EAA69" pageId="5" pageNumber="70" type="discussion">
<paragraph id="555536B8D4378B65BCACFF5EA16EA9A5" blockId="5.[151,1437,152,611]" pageId="5" pageNumber="70">
<emphasis id="679EEAAAD4378B65BCACFF5EA12EA8F1" bold="true" box="[199,315,225,251]" pageId="5" pageNumber="70">Remarks.</emphasis>
<taxonomicName id="92EA4D3BD4378B65BD29FF5EA1E8A8F1" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[322,509,225,251]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="679EEAAAD4378B65BD29FF5EA1E8A8F1" box="[322,509,225,251]" italics="true" pageId="5" pageNumber="70">Ovomanonychus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4378B65BE6FFF5EA295A8F1" bold="true" box="[516,640,225,251]" pageId="5" pageNumber="70">
<taxonomicNameLabel id="7CAD57D1D4378B65BE6FFF5EA295A8F1" box="[516,640,225,251]" pageId="5" pageNumber="70" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="92EA4D3BD4378B65BF6BFF5EA39AA8F1" authorityName="Moser" authorityYear="1919" box="[768,911,225,251]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4378B65BF6BFF5EA39AA8F1" box="[768,911,225,251]" italics="true" pageId="5" pageNumber="70">Manonychus</emphasis>
</taxonomicName>
in the labrum hidden by the clypeus (in dorsal view), with superior margin inflated; and labium quadrate.
<taxonomicName id="92EA4D3BD4378B65BF4EFEBAA3F5A915" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[805,992,261,287]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4378B65BF4EFEBAA3F5A915" box="[805,992,261,287]" italics="true" pageId="5" pageNumber="70">Ovomanonychus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="92EA4D3BD4378B65B81AFEBAA515A915" authorityName="Moser" authorityYear="1919" box="[1137,1280,261,287]" class="Insecta" family="Melolonthidae" genus="Manonychus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus">
<emphasis id="679EEAAAD4378B65B81AFEBAA515A915" box="[1137,1280,261,287]" italics="true" pageId="5" pageNumber="70">Manonychus</emphasis>
</taxonomicName>
(characters in parenthesis), in the galea with outer margin curved (
<figureCitation id="CDD12A3DD4378B65BE8FFE96A32FA949" box="[740,826,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="5" pageNumber="70">Fig. 1B</figureCitation>
) (galea with outer margin lobed;
<figureCitation id="CDD12A3DD4378B65B8DDFE96A519A949" box="[1206,1292,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="5" pageNumber="70">Fig. 1C</figureCitation>
); elytra with costae (
<figureCitation id="CDD12A3DD4378B65BC80FEF2A154A96D" box="[235,321,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="5" pageNumber="70">Fig. 1A</figureCitation>
) (elytra almost smooth); tarsomeres in all legs with sparse bristles ventrally (
<figureCitation id="CDD12A3DD4378B65B8FFFEF2A4F0A96D" box="[1172,1253,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="5" pageNumber="70">Fig. 1F</figureCitation>
) (protarsomeres and mesotarsomeres with pads;
<figureCitation id="CDD12A3DD4378B65BE6FFECEA248A981" box="[516,605,369,395]" captionStart="FIGURE 1" captionStartId="2.[151,250,1662,1687]" captionTargetBox="[170,1408,192,1623]" captionTargetId="figure@2.[151,1436,179,1634]" captionTargetPageId="2" captionText="FIGURE 1. Ovomanonychus diagnostic characters. A,Ovomanonychus striatus new species male paratype dorsal habitus; B, O. striatus right maxilla, galea with outer margin curved; C, Manonychus birabeni Martínez, 1959 right maxilla, galea with outer margin lobed; D, O. striatus labium; E, O. rosettae (Frey, 1976) frons and antennae; F, O. rosettae left anterior tarsomeres with sparse bristles; G, M. birabeni right anterior tarsomeres with pads." figureDoi="http://doi.org/10.5281/zenodo.3735988" httpUri="https://zenodo.org/record/3735988/files/figure.png" pageId="5" pageNumber="70">Fig. 1G</figureCitation>
); metacoxae longer than ventrite II (
<figureCitation id="CDD12A3DD4378B65B86BFECEA442A981" box="[1024,1111,369,395]" captionStart="FIGURE 2" captionStartId="3.[151,250,1837,1862]" captionTargetBox="[160,1411,196,1810]" captionTargetId="figure@3.[151,1436,181,1811]" captionTargetPageId="3" captionText="FIGURE 2. Ovomanonychus diagnostic characters. A, Ovomanonychus rosettae (Frey, 1976) mesotarsomere; B, O. inajae new species spurs set below and above the tarsal articulation; C, O. rosettae metacoxae longer than ventrite II; D, Manonychus ovalis (Blanchard, 1851) metacoxae length equal to ventrite II; E, O. striatus new species metacoxae densely bristled; F, O. rosettae apical region of phallobase excavated." figureDoi="http://doi.org/10.5281/zenodo.3735992" httpUri="https://zenodo.org/record/3735992/files/figure.png" pageId="5" pageNumber="70">Fig. 2C</figureCitation>
) (metacoxae length equal to ventrite II;
<figureCitation id="CDD12A3DD4378B65BD7EFE2AA17AA9A5" box="[277,367,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1837,1862]" captionTargetBox="[160,1411,196,1810]" captionTargetId="figure@3.[151,1436,181,1811]" captionTargetPageId="3" captionText="FIGURE 2. Ovomanonychus diagnostic characters. A, Ovomanonychus rosettae (Frey, 1976) mesotarsomere; B, O. inajae new species spurs set below and above the tarsal articulation; C, O. rosettae metacoxae longer than ventrite II; D, Manonychus ovalis (Blanchard, 1851) metacoxae length equal to ventrite II; E, O. striatus new species metacoxae densely bristled; F, O. rosettae apical region of phallobase excavated." figureDoi="http://doi.org/10.5281/zenodo.3735992" httpUri="https://zenodo.org/record/3735992/files/figure.png" pageId="5" pageNumber="70">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="555536B8D4378B65BCACFE06A36EAA69" blockId="5.[151,1437,152,611]" pageId="5" pageNumber="70">
We place
<taxonomicName id="92EA4D3BD4378B65BD5FFE06A1FAA9D9" authority="Costa &amp; Cherman &amp; Iannuzzi, 2020" authorityName="Costa &amp; Cherman &amp; Iannuzzi" authorityYear="2020" box="[308,495,441,467]" class="Insecta" family="Scarabaeidae" genus="Ovomanonychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="70" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="679EEAAAD4378B65BD5FFE06A1FAA9D9" box="[308,495,441,467]" italics="true" pageId="5" pageNumber="70">Ovomanonychus</emphasis>
</taxonomicName>
<emphasis id="679EEAAAD4378B65BD9DFE06A264A9D9" bold="true" box="[502,625,441,467]" pageId="5" pageNumber="70">
<taxonomicNameLabel id="7CAD57D1D4378B65BD9DFE06A264A9D9" box="[502,625,441,467]" pageId="5" pageNumber="70" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
in Sericoidini based on the following combination of characters: labrum hidden by the clypeus (in dorsal view); clypeus and labrum separate by suture; ligula and labium fused; metatibial apically with two spurs, spurs set below and above the tarsal articulation; lateral keel along the abdominal ventrites; abdomen with six ventral sternites (basal sternite partially hidden by metacoxae), sternites approximately equal in length, evenly convex, separated by distinct sutures (
<bibRefCitation id="317B4B49D4378B65BE8CFDF6A37AAA69" author="Smith, A. B. T." box="[743,879,585,611]" pageId="5" pageNumber="70" pagination="1 - 28" refId="ref6386" refString="Smith, A. B. T. (2008) South American Melolonthinae (Coleoptera: Scarabaeidae) classification and nomenclature: some problems and solutions. Insecta Mundi, 60, 1 - 28." type="journal article" year="2008">Smith 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>