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<document id="4BC38A139E8F336590D231CA7284ECB0" ID-CLB-Dataset="298099" ID-DOI="10.5070/P940561331" ID-GBIF-Dataset="59287004-b877-4428-8e25-e1bb8d48aa37" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913477" ID-ZooBank="1756B24A-813B-423F-896F-91B21FF58A79" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1712169731283" checkinUser="felipe" docAuthor="Hickman, Carole S." docDate="2023" docId="C23987DDFFE8292DFC04FD07EFE5B858" docLanguage="en" docName="PaleoBios.40.5.1-51.pdf" docOrigin="PaleoBios 40 (5)" docSource="http://dx.doi.org/10.5070/p940561331" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.1:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="1" docTitle="Glycymeris andersoni DICKERSON 1917" docType="treatment" docVersion="5" lastPageNumber="15" masterDocId="3E00FFA5FFF8293CFF9FFFA4EE04BD6F" masterDocTitle="Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs" masterLastPageNumber="51" masterPageNumber="1" pageNumber="15" updateTime="1717686085320" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="A86057DE3CA4B67A54D19B561AECB23A">Paleogene marine bivalves of the deep-water Keasey Formation in Oregon, Part II: The pteriomorphs</mods:title>
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<treatment id="C23987DDFFE8292DFC04FD07EFE5B858" ID-DOI="http://doi.org/10.5281/zenodo.11505103" ID-GBIF-Taxon="232155344" ID-Zenodo-Dep="11505103" LSID="urn:lsid:plazi:treatment:C23987DDFFE8292DFC04FD07EFE5B858" httpUri="http://treatment.plazi.org/id/C23987DDFFE8292DFC04FD07EFE5B858" lastPageId="17" lastPageNumber="15" pageId="16" pageNumber="15">
<subSubSection id="028A6540FFE8292CFC04FD07EB90BFD1" box="[923,1428,675,703]" pageId="16" pageNumber="15" type="nomenclature">
<paragraph id="4A2F36CBFFE8292CFC04FD07EB90BFD1" blockId="16.[923,1428,675,734]" box="[923,1428,675,703]" pageId="16" pageNumber="15">
<heading id="116781A7FFE8292CFC04FD07EB90BFD1" allCaps="true" box="[923,1428,675,703]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="15" reason="2">
<taxonomicName id="8D904D48FFE8292CFC04FD07EB90BFD1" authority="DICKERSON, 1917" authorityName="DICKERSON" authorityYear="1917" box="[923,1428,675,703]" class="Bivalvia" family="Glycymerididae" genus="Glycymeris" kingdom="Animalia" order="Arcida" pageId="16" pageNumber="15" phylum="Mollusca" rank="species" species="andersoni">
<emphasis id="78E4EAD9FFE8292CFC04FD07EACFBFD0" box="[923,1227,675,703]" italics="true" pageId="16" pageNumber="15">GLYCYMERIS ANDERSONI</emphasis>
<bibRefCitation id="2E014B3AFFE8292CFB4EFD02EB90BFD1" author="Dickerson, R. E." box="[1233,1428,678,702]" pageId="16" pageNumber="15" pagination="157 - 192" refId="ref29390" refString="Dickerson, R. E. 1917. Climate and its influence upon the Ologicene faunas of the Pacific Coast with descritpions of some new spe- cies from the Molopophorus lincolnensis Zone. Proceedings of the California Academy of Sciences 7 (6): 157 - 192, pls. 27 - 31." type="journal article" year="1917">DICKERSON, 1917</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFE8292CFBA6FD62EAF2BFB1" box="[1081,1270,710,734]" pageId="16" pageNumber="15" type="description">
<paragraph id="4A2F36CBFFE8292CFBA6FD62EAF2BFB1" blockId="16.[923,1428,675,734]" box="[1081,1270,710,734]" pageId="16" pageNumber="15">
<figureCitation id="D2AB2A4EFFE8292CFBA6FD62EAB5BFB1" box="[1081,1201,710,734]" captionStart="Figure 8" captionStartId="17.[112,187,1111,1135]" captionTargetBox="[112,1519,192,1084]" captionTargetId="figure-290@17.[112,1519,189,1087]" captionTargetPageId="17" captionText="Figure 8. Glycymeris andersoni Dickerson (1917), Gries Ranch Formation. A. Interior of right valve, hypotype UCMP 110730. B, C. Exterior and interior of right valve, hypotype UCMP 110729. D, E. Ventral and do2rsal of articulated valves, hypotype UCMP 110731. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913493" httpUri="https://zenodo.org/record/10913493/files/figure.png" pageId="16" pageNumber="15">FIGS. 8A–E</figureCitation>
,
<figureCitation id="D2AB2A4EFFE8292CFB24FD62EAF2BFB1" box="[1211,1270,710,734]" captionStart="Figure 9" captionStartId="18.[112,187,764,788]" captionTargetBox="[112,803,192,740]" captionTargetId="figure-581@18.[112,803,192,740]" captionTargetPageId="18" captionText="Figure 9. Comparison of dorsal slope angles in Juvenile and adult shells of Glycymeris andersoni Dickerson (1917) and Glycymeris winlockensis Effinger (1938), Gries Ranch Formation. A. Adult G. andersoni hypotype UCMP 110732, scale bar=1 cm. B. Adult G. winlockensis hypotype UCMP 110734, scale bar=1 cm. C. Juvenile G. andersoni hypotype UCMP 110733, scale bar=5 mm. D. Juvenile G. winlockensis hypotype UCMP 110735, scale bar=5 mm." figureDoi="http://doi.org/10.5281/zenodo.10913497" httpUri="https://zenodo.org/record/10913497/files/figure.png" pageId="16" pageNumber="15">9A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFE8292CFCA0FD5EEA61BB52" pageId="16" pageNumber="15" type="reference_group">
<paragraph id="4A2F36CBFFE8292CFCA0FD5EEA31BE5E" blockId="16.[831,1507,760,961]" pageId="16" pageNumber="15">
<taxonomicName id="8D904D48FFE8292CFCA0FD5EEB23BE7B" authority="Dickerson (1917)" authorityName="DICKERSON" authorityYear="1917" box="[831,1319,760,790]" class="Bivalvia" family="Glycymerididae" genus="Glycymeris" kingdom="Animalia" order="Arcida" pageId="16" pageNumber="15" phylum="Mollusca" rank="species" species="andersoni">
<emphasis id="78E4EAD9FFE8292CFCA0FD5EEA40BE7B" box="[831,1092,760,790]" italics="true" pageId="16" pageNumber="15">Glycymeris andersoni</emphasis>
<bibRefCitation id="2E014B3AFFE8292CFBD5FD5EEB23BE7B" author="Dickerson, R. E." box="[1098,1319,762,788]" pageId="16" pageNumber="15" pagination="157 - 192" refId="ref29390" refString="Dickerson, R. E. 1917. Climate and its influence upon the Ologicene faunas of the Pacific Coast with descritpions of some new spe- cies from the Molopophorus lincolnensis Zone. Proceedings of the California Academy of Sciences 7 (6): 157 - 192, pls. 27 - 31." type="journal article" year="1917">Dickerson (1917)</bibRefCitation>
</taxonomicName>
. pp. 166–167; pl. 27, figs. 3a–c.
</paragraph>
<paragraph id="4A2F36CBFFE8292CFCA0FCE6EDA5BE16" blockId="16.[831,1507,760,961]" pageId="16" pageNumber="15">
<taxonomicName id="8D904D48FFE8292CFCA0FCE6EB9CBE33" authority="Dickerson, Effinger (1938)" authorityName="Dickerson, Effinger" authorityYear="1938" box="[831,1432,832,862]" class="Bivalvia" family="Glycymerididae" genus="Glycymeris" kingdom="Animalia" order="Arcida" pageId="16" pageNumber="15" phylum="Mollusca" rank="species" species="andersoni">
<emphasis id="78E4EAD9FFE8292CFCA0FCE6EA40BE33" box="[831,1092,832,862]" italics="true" pageId="16" pageNumber="15">Glycymeris andersoni</emphasis>
Dickerson,
<bibRefCitation id="2E014B3AFFE8292CFB4AFCE6EB9CBE33" author="Effinger, W. L." box="[1237,1432,834,860]" pageId="16" pageNumber="15" pagination="355 - 390" refId="ref29720" refString="Effinger, W. L. 1938. The Gries Ranch fauna (Oligocene) of Western Washington. Journal of Paleontology 12 (4): 355 - 390." type="journal article" year="1938">Effinger (1938)</bibRefCitation>
</taxonomicName>
. p. 366.
</paragraph>
<paragraph id="4A2F36CBFFE8292CFCA0FC2EEA3CBEAE" blockId="16.[831,1507,760,961]" pageId="16" pageNumber="15">
<taxonomicName id="8D904D48FFE8292CFCA0FC2EEB95BECB" authority="Dickerson, Weaver (1943)" authorityName="Dickerson, Weaver" authorityYear="1943" box="[831,1425,904,934]" class="Bivalvia" family="Glycymerididae" genus="Glycymeris" kingdom="Animalia" order="Arcida" pageId="16" pageNumber="15" phylum="Mollusca" rank="species" species="andersoni">
<emphasis id="78E4EAD9FFE8292CFCA0FC2EEA40BECB" box="[831,1092,904,934]" italics="true" pageId="16" pageNumber="15">Glycymeris andersoni</emphasis>
Dickerson, Weaver (1943)
</taxonomicName>
. p. 59; pl. 10, figs. 5, 11.
</paragraph>
<paragraph id="4A2F36CBFFE8292CFCC0FC76EA19B8BA" blockId="16.[831,1520,978,1944]" pageId="16" pageNumber="15">
<emphasis id="78E4EAD9FFE8292CFCC0FC76EDF0BE83" bold="true" box="[863,1012,978,1004]" pageId="16" pageNumber="15">Discussion</emphasis>
—
<taxonomicName id="8D904D48FFE8292CFB8DFC76EB25BE82" authorityName="Dickerson, Weaver" authorityYear="1943" box="[1042,1313,977,1006]" class="Bivalvia" family="Glycymerididae" genus="Glycymeris" kingdom="Animalia" order="Arcida" pageId="16" pageNumber="15" phylum="Mollusca" rank="species" species="andersoni">
<emphasis id="78E4EAD9FFE8292CFB8DFC76EB25BE82" box="[1042,1313,977,1006]" italics="true" pageId="16" pageNumber="15">Glycymeris andersoni</emphasis>
</taxonomicName>
is easily recognized by its ovate, moderately inflated, nearly equilateral shell with broad flat-topped radial ribs and narrow interspaces accentuated by chalky preservation (
<figureCitation id="D2AB2A4EFFE8292CFAFAFB9FEBDBB93A" box="[1381,1503,1083,1109]" captionStart="Figure 8" captionStartId="17.[112,187,1111,1135]" captionTargetBox="[112,1519,192,1084]" captionTargetId="figure-290@17.[112,1519,189,1087]" captionTargetPageId="17" captionText="Figure 8. Glycymeris andersoni Dickerson (1917), Gries Ranch Formation. A. Interior of right valve, hypotype UCMP 110730. B, C. Exterior and interior of right valve, hypotype UCMP 110729. D, E. Ventral and do2rsal of articulated valves, hypotype UCMP 110731. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913493" httpUri="https://zenodo.org/record/10913493/files/figure.png" pageId="16" pageNumber="15">Fig. 8B, D</figureCitation>
). The commissural margin is strongly crenulate (
<figureCitation id="D2AB2A4EFFE8292CFA14FBFAEBE2B917" box="[1419,1510,1118,1144]" captionStart="Figure 8" captionStartId="17.[112,187,1111,1135]" captionTargetBox="[112,1519,192,1084]" captionTargetId="figure-290@17.[112,1519,189,1087]" captionTargetPageId="17" captionText="Figure 8. Glycymeris andersoni Dickerson (1917), Gries Ranch Formation. A. Interior of right valve, hypotype UCMP 110730. B, C. Exterior and interior of right valve, hypotype UCMP 110729. D, E. Ventral and do2rsal of articulated valves, hypotype UCMP 110731. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913493" httpUri="https://zenodo.org/record/10913493/files/figure.png" pageId="16" pageNumber="15">Fig. 8D</figureCitation>
) and the interlocking of the valves is further reinforced by interior crenulation (
<figureCitation id="D2AB2A4EFFE8292CFBF4FB07EAE1B9D1" box="[1131,1253,1187,1214]" captionStart="Figure 8" captionStartId="17.[112,187,1111,1135]" captionTargetBox="[112,1519,192,1084]" captionTargetId="figure-290@17.[112,1519,189,1087]" captionTargetPageId="17" captionText="Figure 8. Glycymeris andersoni Dickerson (1917), Gries Ranch Formation. A. Interior of right valve, hypotype UCMP 110730. B, C. Exterior and interior of right valve, hypotype UCMP 110729. D, E. Ventral and do2rsal of articulated valves, hypotype UCMP 110731. Scale bars=1 cm." figureDoi="http://doi.org/10.5281/zenodo.10913493" httpUri="https://zenodo.org/record/10913493/files/figure.png" pageId="16" pageNumber="15">Fig. 8A, C</figureCitation>
). Maximum shell size is large (length and height&gt;
<quantity id="8D689B2EFFE8292CFB3CFB62EAF9B98E" box="[1187,1277,1222,1249]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="15" unit="mm" value="25.0">25 mm</quantity>
) relative to species in the older tropical Eocene faunas of the Pacific Northwest and
<collectingRegion id="8854F829FFE8292CFC22FAA8EA3EB849" box="[957,1082,1292,1318]" country="United States of America" name="California" pageId="16" pageNumber="15">California</collectingRegion>
. It is abundant in the Gries Ranch Formation and represented by both juvenile and adult specimens of equal linear dimension, consistent with isometric growth. The dorsal slope angle does not differ between large and small shells (
<figureCitation id="D2AB2A4EFFE8292CFB44FA3CEB5FB8DC" box="[1243,1371,1432,1459]" captionStart="Figure 9" captionStartId="18.[112,187,764,788]" captionTargetBox="[112,803,192,740]" captionTargetId="figure-581@18.[112,803,192,740]" captionTargetPageId="18" captionText="Figure 9. Comparison of dorsal slope angles in Juvenile and adult shells of Glycymeris andersoni Dickerson (1917) and Glycymeris winlockensis Effinger (1938), Gries Ranch Formation. A. Adult G. andersoni hypotype UCMP 110732, scale bar=1 cm. B. Adult G. winlockensis hypotype UCMP 110734, scale bar=1 cm. C. Juvenile G. andersoni hypotype UCMP 110733, scale bar=5 mm. D. Juvenile G. winlockensis hypotype UCMP 110735, scale bar=5 mm." figureDoi="http://doi.org/10.5281/zenodo.10913497" httpUri="https://zenodo.org/record/10913497/files/figure.png" pageId="16" pageNumber="15">Fig. 9A, B</figureCitation>
), indicating isometric growth.
</paragraph>
<paragraph id="4A2F36CBFFE8292CFCC0FA7AEA61BB52" blockId="16.[831,1520,978,1944]" pageId="16" pageNumber="15">
A second Gries Ranch glycymeridid species, described subsequently by
<bibRefCitation id="2E014B3AFFE8292CFB8BF9A4EAD3BB74" author="Effinger, W. L." box="[1044,1239,1536,1563]" pageId="16" pageNumber="15" pagination="355 - 390" refId="ref29720" refString="Effinger, W. L. 1938. The Gries Ranch fauna (Oligocene) of Western Washington. Journal of Paleontology 12 (4): 355 - 390." type="journal article" year="1938">Effinger (1938)</bibRefCitation>
, is treated below, and figured for comparison.
</paragraph>
</subSubSection>
<subSubSection id="028A6540FFE8292DFCC0F9E1EFE5B858" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15" type="materials_examined">
<paragraph id="4A2F36CBFFE8292CFCC0F9E1EDA5BBEC" blockId="16.[831,1520,978,1944]" pageId="16" pageNumber="15">
<emphasis id="78E4EAD9FFE8292CFCC0F9E1EA5FBB30" bold="true" box="[863,1115,1605,1631]" pageId="16" pageNumber="15">Material examined</emphasis>
—
<materialsCitation id="FAF83C96FFE8292CFBE4F9E2ED99BBEC" ID-GBIF-Occurrence="4873742308" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="IP2289" specimenCount="39">
<specimenCount id="5C96FD42FFE8292CFBE4F9E2EB26BB0F" box="[1147,1314,1606,1632]" count="39" pageId="16" pageNumber="15" type="generic">39 specimens</specimenCount>
from Loc.
<collectionCode id="2C81AE0EFFE8292CFA3CF9E2EBEBBB0F" box="[1443,1519,1606,1632]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP</collectionCode>
<specimenCode id="1A369EB0FFE8292CFCA0F9CDED99BBEC" box="[831,925,1641,1667]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">IP2289</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="4A2F36CBFFE8292CFCC0F92FEB23BA3C" blockId="16.[831,1520,978,1944]" pageId="16" pageNumber="15">
<emphasis id="78E4EAD9FFE8292CFCC0F92FEA4EBBCA" bold="true" box="[863,1098,1675,1701]" pageId="16" pageNumber="15">
<typeStatus id="952B8869FFE8292CFCC0F92FEDA6BBCA" box="[863,930,1675,1701]" pageId="16" pageNumber="15">Type</typeStatus>
information
</emphasis>
—Figured hypotypes, all from Loc.
<materialsCitation id="FAF83C96FFE8292CFCA0F90AEDF0BBA7" ID-GBIF-Occurrence="4873742323" box="[831,1012,1710,1736]" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="IP2289" specimenCount="1" typeStatus="holotype">
<collectionCode id="2C81AE0EFFE8292CFCA0F90AED8FBBA7" box="[831,907,1710,1736]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP</collectionCode>
<specimenCode id="1A369EB0FFE8292CFC0AF90AEDF0BBA7" box="[917,1012,1710,1736]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">IP2289</specimenCode>
</materialsCitation>
:
<materialsCitation id="FAF83C96FFE8292CFB9EF90AEABDBBA7" ID-GBIF-Occurrence="4873742331" box="[1025,1209,1710,1736]" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="UCMP 110729" specimenCount="1" typeStatus="holotype">
<specimenCode id="1A369EB0FFE8292CFB9EF90AEABDBBA7" box="[1025,1209,1710,1736]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP 110729</specimenCode>
</materialsCitation>
, length 19.5 mm, height 19.2 mm;
<materialsCitation id="FAF83C96FFE8292CFC5EF975EA7EBB84" ID-GBIF-Occurrence="4873742342" box="[961,1146,1745,1771]" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="UCMP 110730" specimenCount="1" typeStatus="holotype">
<specimenCode id="1A369EB0FFE8292CFC5EF975EA7EBB84" box="[961,1146,1745,1771]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP 110730</specimenCode>
</materialsCitation>
, length 22.1 mm, height 21.6 mm;
<materialsCitation id="FAF83C96FFE8292CFC1EF957EA3DBA62" ID-GBIF-Occurrence="4873742325" box="[897,1081,1779,1805]" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="UCMP 110731" specimenCount="1" typeStatus="holotype">
<specimenCode id="1A369EB0FFE8292CFC1EF957EA3DBA62" box="[897,1081,1779,1805]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP 110731</specimenCode>
</materialsCitation>
, length 29.0 mm, height 27.7 mm;
<materialsCitation id="FAF83C96FFE8292CFCA0F8B2EDF1BA5F" ID-GBIF-Occurrence="4873742332" box="[831,1013,1814,1840]" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="UCMP 110732" specimenCount="1" typeStatus="holotype">
<specimenCode id="1A369EB0FFE8292CFCA0F8B2EDF1BA5F" box="[831,1013,1814,1840]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP 110732</specimenCode>
</materialsCitation>
, length 19.0 mm, height 17.1 mm;
<materialsCitation id="FAF83C96FFE8292CFA3CF8B2EDA6BA3C" ID-GBIF-Occurrence="4873742309" collectionCode="UCMP" pageId="16" pageNumber="15" specimenCode="UCMP 110733" specimenCount="1" typeStatus="holotype">
<specimenCode id="1A369EB0FFE8292CFA3CF8B2EDA6BA3C" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="16" pageNumber="15" type="University or college">UCMP 110733</specimenCode>
</materialsCitation>
, length 8.2 mm, height 7.6 mm.
</paragraph>
<paragraph id="4A2F36CBFFE8292DFCC0F8FFEFE5B858" blockId="16.[831,1520,978,1944]" lastBlockId="17.[112,800,1243,1336]" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15">
<emphasis id="78E4EAD9FFE8292CFCC0F8FFEAB2BA1A" bold="true" box="[863,1206,1883,1909]" pageId="16" pageNumber="15">Stratigraphic occurrence</emphasis>
—This species is known only from the Gries Ranch Formation. It is not present in the tropical Eocene fauna of the underlying Cowlitz Formation or in the recovery fauna of the overlying Pittsburg Bluff and Eugene Formations.
</paragraph>
</subSubSection>
</treatment>
</document>
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