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<document id="08BE3E7BDB2DE3C9BD0A74E030A9600C" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A9A2FF462B9CC63FFEFAFAD0" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mathilda dania Hansen 2019, n. sp." docType="treatment" docUuid="CB14443B-85B2-41B9-8D0D-9D07AC7F3FE1" docUuidSource="ZooBank" docVersion="22" lastPageNumber="164" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="163" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="92037F1FA82F0A55F19CE914E232B938">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
</mods:titleInfo>
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<mods:namePart id="0FF62566947F362C3065F924FE2833AB">Hansen, Thomas</mods:namePart>
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<mods:date id="024DB585DED068140975E1128DB03031">2019</mods:date>
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<mods:number id="855D71FF55140195E900804C349B18E5">2019-08-12</mods:number>
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<mods:number id="A0143D765F04160CFD9B5E04481B897A">4654</mods:number>
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<mods:classification id="B4D22F36E90DCFD5B39E31D68A93F089">journal article</mods:classification>
<mods:identifier id="C85B2736CE24C75C8E7094F5D90481F1" type="CLB-Dataset">26049</mods:identifier>
<mods:identifier id="FE839B3953A64C250C0534F652FA33E7" type="DOI">10.11646/zootaxa.4654.1.1</mods:identifier>
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<treatment id="BF18F633A9A2FF462B9CC63FFEFAFAD0" ID-GBIF-Taxon="159074633" LSID="urn:lsid:zoobank.org:act:CB14443B-85B2-41B9-8D0D-9D07AC7F3FE1" httpUri="http://treatment.plazi.org/id/BF18F633A9A2FF462B9CC63FFEFAFAD0" lastPageId="163" lastPageNumber="164" pageId="162" pageNumber="163">
<subSubSection id="7FAB14AEA9A2FF472B9CC63FFE35F86E" box="[151,412,1906,1933]" pageId="162" pageNumber="163" type="nomenclature">
<paragraph id="370E4725A9A2FF472B9CC63FFE35F86E" blockId="162.[151,412,1906,1969]" box="[151,412,1906,1933]" pageId="162" pageNumber="163">
<heading id="6C46F049A9A2FF472B9CC63FFE35F86E" bold="true" box="[151,412,1906,1933]" fontSize="11" level="1" pageId="162" pageNumber="163" reason="1">
<emphasis id="05C59B37A9A2FF472B9CC63FFE35F86E" bold="true" box="[151,412,1906,1933]" pageId="162" pageNumber="163">
<taxonomicName id="F0B13CA6A9A2FF472B9CC63FFEF8F86E" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[151,337,1906,1933]" class="Gastropoda" family="Mathildidae" genus="Mathilda" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="162" pageNumber="163" phylum="Mollusca" rank="species" species="dania" status="sp. nov.">
<emphasis id="05C59B37A9A2FF472B9CC63FFEF8F86E" bold="true" box="[151,337,1906,1933]" italics="true" pageId="162" pageNumber="163">Mathilda dania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EF6264CA9A2FF472A52C63EFE35F86E" box="[345,412,1907,1933]" pageId="162" pageNumber="163" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA9A2FF472B9CC6DAFEBAF852" box="[151,275,1942,1969]" pageId="162" pageNumber="163" type="description">
<paragraph id="370E4725A9A2FF472B9CC6DAFEBAF852" blockId="162.[151,412,1906,1969]" box="[151,275,1942,1969]" pageId="162" pageNumber="163">
<figureCitation id="AF8A5BA0A9A2FF472B9CC6DAFF40F852" box="[151,233,1942,1969]" captionStart="FIGURE 35" captionStartId="167.[151,250,188,213]" captionTargetId="figure@166.[151,1435,179,1918]" captionTargetPageId="166" captionText="FIGURE 35. A–B. Pseudomalaxis sp., Silicone cast of MGUH 33306. Width 4.3 mm. C–D. Climacopoma? sp., Silicone cast of MGUH 33307. Width 13.1 and 9.9 mm respectively. E–I. Mathilda dania n. sp. E. Silicone cast of holotype MGUH 33308. Width 1.9 mm. F. Silicone cast of paratype MGUH 33309. Width 2.0 mm. G. Silicone cast of paratype MGUH 33310. Width 0.8 mm. H. Silicone cast of paratype MGUH 33311. Width 0.7 mm. I. Silicone cast of paratype MGUH 33312. Width 1.7 mm. J–K. Clathrobaculus sp., Silicone cast of MGUH 33313. Width 0.5 mm. L–P. Ringicula sp. L. Silicone cast of MGUH 33314. Width 7.0 mm. M–N. Silicone cast of MGUH 33315 and a partly hidden second specimen. Width 7.3 and 6.6 mm respectively. O–P. Silicone cast of MGUH 33316. Width 5.6, 5.3 mm respectively. Q–T. Cylichna stevniensis n. sp. Q. Silicone cast of holotype MGUH 33317. Width 1.4 mm. R–S. Internal mould of paratype MGUH 33318. Width 1.5 mm. T. Internal mould of paratype MGUH 33319. Width 2.8 mm. U–V. Cylichna? sp. A. Cast of MGUH 33320. Width 2.3, 2.4 mm respectively. W–Y. Cylichna? sp. B. W. Wax cast of MGUH 33321. Width 2.4 mm. X–Y. Inner mould of MGUH 33322. Width 1.3 mm. Z–AE. Roxania sp. Cast of MGUH 33323. Width 3.2, 3.3 mm respectively." figureDoi="http://doi.org/10.5281/zenodo.3442126" httpUri="https://zenodo.org/record/3442126/files/figure.png" pageId="162" pageNumber="163">Figs 35</figureCitation>
E–I
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA9A2FF462B9CC693FEFAFAD0" lastPageId="163" lastPageNumber="164" pageId="162" pageNumber="163" type="materials_examined">
<paragraph id="370E4725A9A2FF462B9CC693FCCAFF34" blockId="162.[151,1436,2014,2041]" lastBlockId="163.[151,1437,152,1331]" lastPageId="163" lastPageNumber="164" pageId="162" pageNumber="163">
<emphasis id="05C59B37A9A2FF472B9CC693FEBBF81A" bold="true" box="[151,274,2014,2041]" pageId="162" pageNumber="163">Diagnosis.</emphasis>
Protoconch smooth, nearly perpendicular to teleoconch axis. Teleoconch moderately slender with ap- proximately 50 thin transverse ribs and three to four coarser, slightly less densely spaced spiral ribs adapical of keel. Keel situated one quarter the whorl height from abapical suture.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC1ADFC60FF18" blockId="163.[151,1437,152,1331]" box="[199,969,224,251]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC1ADFE1DFF18" bold="true" box="[199,436,224,251]" pageId="163" pageNumber="164">Derivation of name.</emphasis>
Refers to the Danian age and its Danish origin.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC048FE56FE4C" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC048FEDEFEFC" bold="true" box="[199,375,260,287]" pageId="163" pageNumber="164">Type material.</emphasis>
<materialsCitation id="87D94D78A9A3FF462A8BC048FB96FEFC" ID-GBIF-Occurrence="2331574455" box="[384,1087,261,287]" collectionCode="MGUH" pageId="163" pageNumber="164" specimenCode="MGUH 33308" specimenCount="1" typeStatus="holotype">
<typeStatus id="E80AF987A9A3FF462A8BC048FE41FEFC" box="[384,488,261,287]" pageId="163" pageNumber="164" type="holotype">Holotype</typeStatus>
<specimenCode id="6717EF5EA9A3FF462AF9C048FD3EFEFC" box="[498,663,261,287]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="163" pageNumber="164">MGUH 33308</specimenCode>
is a nearly complete external mould
</materialsCitation>
.
<materialsCitation id="87D94D78A9A3FF462F47C048FB8BFEA0" ID-GBIF-Occurrence="2331574342" collectionCode="MGUH" pageId="163" pageNumber="164" specimenCode="MGUH 33309" specimenCount="1" typeStatus="paratype">
<typeStatus id="E80AF987A9A3FF462F47C048FB07FEFC" box="[1100,1198,261,287]" pageId="163" pageNumber="164" type="paratype">Paratype</typeStatus>
<specimenCode id="6717EF5EA9A3FF462FB3C048FAF5FEFC" box="[1208,1372,261,287]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="163" pageNumber="164">MGUH 33309</specimenCode>
is an external mould and comes from the
<taxonomicName id="F0B13CA6A9A3FF462926C064FC2BFEA0" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[557,898,297,323]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="163" pageNumber="164" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Skeldervig
</materialsCitation>
.
<materialsCitation id="87D94D78A9A3FF462F26C064FB69FE84" ID-GBIF-Occurrence="2331574406" collectionCode="MGUH" pageId="163" pageNumber="164" specimenCode="MGUH 33310" specimenCount="2" typeStatus="paratype">
<typeStatus id="E80AF987A9A3FF462F26C064FB26FEA0" box="[1069,1167,297,323]" pageId="163" pageNumber="164" type="paratype">Paratype</typeStatus>
<specimenCode id="6717EF5EA9A3FF462F9DC064FA90FEA0" box="[1174,1337,297,323]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="163" pageNumber="164">MGUH 33310</specimenCode>
is a protoconch from
<quantity id="F049EAC0A9A3FF462A3FC000FE13FE84" box="[308,442,332,359]" metricMagnitude="-1" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="163" pageNumber="164" unit="cm" value="22.5" valueMax="25.0" valueMin="20.0">20 to 25 cm</quantity>
above the K-Pg boundary at Højerup, while the protoconch
<typeStatus id="E80AF987A9A3FF462F6BC000FB69FE84" box="[1120,1216,333,359]" pageId="163" pageNumber="164" type="paratype">paratype</typeStatus>
</materialsCitation>
<materialsCitation id="87D94D78A9A3FF462FC3C000FCA5FE68" ID-GBIF-Occurrence="2331574422" collectionCode="MGUH" pageId="163" pageNumber="164" specimenCode="MGUH 33311" specimenCount="1" typeStatus="paratype">
<specimenCode id="6717EF5EA9A3FF462FC3C000FAC0FE84" box="[1224,1385,333,359]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="163" pageNumber="164">MGUH 33311</specimenCode>
was found
<quantity id="F049EAC0A9A3FF462BEBC03CFEC1FE68" box="[224,360,368,395]" metricMagnitude="-1" metricUnit="m" metricValue="4.7" metricValueMax="5.2" metricValueMin="4.2" pageId="163" pageNumber="164" unit="cm" value="47.0" valueMax="52.0" valueMin="42.0">42 to 52 cm</quantity>
above the K-Pg boundary at Rødvig
</materialsCitation>
.
<materialsCitation id="87D94D78A9A3FF462813C03CFE52FE4C" ID-GBIF-Occurrence="2331574429" collectionCode="MGUH" location="Vokslev" municipality="Northern Jutland" pageId="163" pageNumber="164" specimenCode="MGUH 33312" specimenCount="1" typeStatus="paratype">
<typeStatus id="E80AF987A9A3FF462813C03CFCD3FE68" box="[792,890,369,395]" pageId="163" pageNumber="164" type="paratype">Paratype</typeStatus>
<specimenCode id="6717EF5EA9A3FF462889C03CFB8CFE68" box="[898,1061,369,395]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="163" pageNumber="164">MGUH 33312</specimenCode>
is an external mould of the spire from
<location id="326E11FEA9A3FF462BDFC0D8FE86FE4C" LSID="urn:lsid:plazi:treatment:BF18F633A9A2FF462B9CC63FFEFAFAD0:326E11FEA9A3FF462BDFC0D8FE86FE4C" box="[212,303,405,431]" municipality="Northern Jutland" name="Vokslev" pageId="163" pageNumber="164">Vokslev</location>
,
<collectingMunicipality id="D76ADD5FA9A3FF462A31C0D8FE52FE4C" box="[314,507,405,431]" pageId="163" pageNumber="164">Northern Jutland</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC0F5FEAEFDDC" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC0F5FE16FE30" bold="true" box="[199,447,440,467]" pageId="163" pageNumber="164">Additional material.</emphasis>
<materialsCitation id="87D94D78A9A3FF462AC1C0F5FEAAFDDC" ID-GBIF-Occurrence="2331574463" collectionCode="OSM" country="Denmark" location="Natural History Museum" pageId="163" pageNumber="164" specimenCount="10">
<collectionCode id="51A0DFE0A9A3FF462AC1C0F5FDA3FE30" box="[458,522,440,467]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="163" pageNumber="164">ØSM</collectionCode>
.10042-89-c–d,
<specimenCount id="21B78CACA9A3FF4629B5C0F4FCF3FE30" box="[702,858,441,467]" pageId="163" pageNumber="164" type="generic">10 specimens</specimenCount>
with the informal sample numbers SH.294.A–C, SH.398, SH.446, SH.499.A–B, SR.476.A, SR.624, SR.631, SR.641, SR.668.B and SO.189, and 3 more or less fragmentary moulds without numbers from the uncatalogued old collections at the
<location id="326E11FEA9A3FF462F57C34CFAD2FDF8" LSID="urn:lsid:plazi:treatment:BF18F633A9A2FF462B9CC63FFEFAFAD0:326E11FEA9A3FF462F57C34CFAD2FDF8" box="[1116,1403,513,539]" country="Denmark" name="Natural History Museum" pageId="163" pageNumber="164">Natural History Museum</location>
of
<collectingCountry id="4FA607B5A9A3FF462B9CC368FEAAFDDC" box="[151,259,549,575]" name="Denmark" pageId="163" pageNumber="164">Denmark</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC304FF71FD64" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC304FDEAFD80" bold="true" box="[199,579,584,611]" pageId="163" pageNumber="164">
<typeStatus id="E80AF987A9A3FF462BCCC304FEABFD80" box="[199,258,585,611]" pageId="163" pageNumber="164">Type</typeStatus>
stratum and
<typeStatus id="E80AF987A9A3FF462AA1C304FE74FD80" box="[426,477,585,611]" pageId="163" pageNumber="164">type</typeStatus>
locality.
</emphasis>
<taxonomicName id="F0B13CA6A9A3FF462946C304FC0FFD80" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[589,934,585,611]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="163" pageNumber="164" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
of the Rødvig Formation at Holtug, Stevns Klint.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC3DDFC92FCF4" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC3DDFEF3FD48" bold="true" box="[199,346,656,683]" pageId="163" pageNumber="164">Occurrence.</emphasis>
Middle to upper part of the
<taxonomicName id="F0B13CA6A9A3FF4629ACC3DCFBA8FD48" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[679,1025,657,683]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="163" pageNumber="164" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
of the Rødvig Formation and basal Korsnaeb Member of the Stevns Klint Formation. This species has been found at Rødvig, Skeldervig, Højerup and at Holtug, all localities at Stevns Klint. Additionally it occurs in material collected by C. Heinberg from the contemporaneous ‘dead layer’ in Vokslev Quarry, Northern Jutland.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC26DFEC1FCBC" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC26DFEF0FCD8" bold="true" box="[199,345,800,827]" pageId="163" pageNumber="164">Description.</emphasis>
Protoconch hyperstrophic, diverging about 125˚ from teleoconch axis, smooth with sharp transi- tion to teleoconch.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC224FF60FB9C" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">Teleoconch slender, conical; whorls with strong keel ¼ the whorl height from rather indistinct abapical suture; whorl sides flattened to slightly concave adapically and abapically to keel. Last whorl taking up 40 % of shell height. Base with weaker keel, abapically flattened. Aperture rounded, sub-rectangular with slightly thickened lip. Teleoconch sculpture on first whorl consisting of densely spaced weak transverse ribs and three to four spiral ribs, the one on the keel being the strongest. Number of spiral and transverse ribs increasing abapically on shell, the spiral ribs reaching a number of five to six moderately coarse ribs. Transverse ribs numbering around 50 on fourth whorl. Base, except for strong spiral rib on abapical keel, nearly smooth or carrying weak spiral ribs and faint transverse ribs.</paragraph>
<paragraph id="370E4725A9A3FF462BCCC5C4FAFDFB40" blockId="163.[151,1437,152,1331]" box="[199,1364,1160,1187]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC5C4FED7FB40" bold="true" box="[199,382,1161,1187]" pageId="163" pageNumber="164">Measurements.</emphasis>
<typeStatus id="E80AF987A9A3FF462A8FC5C4FE45FB40" box="[388,492,1161,1187]" pageId="163" pageNumber="164" type="holotype">Holotype</typeStatus>
<collectionCode id="51A0DFE0A9A3FF462AFFC5C4FDE3FB40" box="[500,586,1161,1187]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="163" pageNumber="164">MGUH</collectionCode>
33308 is
<quantity id="F049EAC0A9A3FF4629BCC5C4FCA4FB40" box="[695,781,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="163" pageNumber="164" unit="mm" value="4.3">4.3 mm</quantity>
high and
<quantity id="F049EAC0A9A3FF462877C5C4FC7BFB40" box="[892,978,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="163" pageNumber="164" unit="mm" value="1.7">1.7 mm</quantity>
wide with 5 ½ teleoconch whorls.
</paragraph>
<paragraph id="370E4725A9A3FF462BCCC5E0FEFAFAD0" blockId="163.[151,1437,152,1331]" pageId="163" pageNumber="164">
<emphasis id="05C59B37A9A3FF462BCCC5E0FE92FB24" bold="true" box="[199,315,1197,1223]" pageId="163" pageNumber="164">Remarks.</emphasis>
<taxonomicName id="F0B13CA6A9A3FF462A49C5E0FE59FB24" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[322,496,1197,1223]" class="Gastropoda" family="Mathildidae" genus="Mathilda" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="163" pageNumber="164" phylum="Mollusca" rank="species" species="dania" status="sp. nov.">
<emphasis id="05C59B37A9A3FF462A49C5E0FE59FB24" box="[322,496,1197,1223]" italics="true" pageId="163" pageNumber="164">Mathilda dania</emphasis>
</taxonomicName>
<emphasis id="05C59B37A9A3FF462AFCC5E0FD9FFB24" bold="true" box="[503,566,1197,1223]" pageId="163" pageNumber="164">
<taxonomicNameLabel id="1EF6264CA9A3FF462AFCC5E0FD9FFB24" box="[503,566,1197,1223]" pageId="163" pageNumber="164" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resembles the Selandian
<taxonomicName id="F0B13CA6A9A3FF46285EC5E0FB14FB24" authority="(Ravn, 1939)" baseAuthorityName="Ravn" baseAuthorityYear="1939" box="[853,1213,1197,1223]" class="Gastropoda" family="Mathildidae" genus="Mathilda" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="163" pageNumber="164" phylum="Mollusca" rank="species" species="carinata">
<emphasis id="05C59B37A9A3FF46285EC5E0FB8BFB24" box="[853,1058,1197,1223]" italics="true" pageId="163" pageNumber="164">Mathilda carinata</emphasis>
(
<bibRefCitation id="53203AD4A9A3FF462F3AC5E0FB1CFB24" author="Ravn, J. P. J." box="[1073,1205,1197,1223]" pageId="163" pageNumber="164" pagination="1 - 106" refId="ref138409" refString="Ravn, J. P. J. (1939) Etudes sur les mollusques du Paleocene de Copenhague. Det Kongelige Danske Videnskabernes Selskab. Biologiske Skrifter, 1 (1), 1 - 106 pp., pls. 1 - 4." type="book chapter" year="1939">Ravn, 1939</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="4FA607B5A9A3FF462E09C5E0FAC4FB24" box="[1282,1389,1197,1223]" name="Denmark" pageId="163" pageNumber="164">Denmark</collectingCountry>
, but has distinctly less convex whorls with a stronger keel and fewer strong spiral ribs abapical to the keel. It differs from the Danian
<taxonomicName id="F0B13CA6A9A3FF462A5EC5B8FD49FAEC" authority="(Ravn, 1933)" baseAuthorityName="Ravn" baseAuthorityYear="1933" box="[341,736,1269,1295]" class="Gastropoda" family="Mathildidae" genus="Mathilda" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="163" pageNumber="164" phylum="Mollusca" rank="species" species="unicarinata">
<emphasis id="05C59B37A9A3FF462A5EC5B8FDEFFAEC" box="[341,582,1269,1295]" italics="true" pageId="163" pageNumber="164">Mathilda unicarinata</emphasis>
(
<bibRefCitation id="53203AD4A9A3FF46295FC5B8FD7EFAEC" author="Ravn, J. P. J." box="[596,727,1269,1295]" pageId="163" pageNumber="164" pagination="1 - 74" refId="ref138358" refString="Ravn, J. P. J. (1933) Etudes sur les Pelecypodes et Gastropodes Daniens du Calcaire de Faxe. Det Kongelige Danske Videnskabernes Selskabs Skrifter, 9. raekke, Naturvidenskabelig og mathematisk Afdeling, 2, 1 - 74, pls. 1 - 7." type="journal article" year="1933">Ravn, 1933</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="4FA607B5A9A3FF46282FC5B8FC25FAEC" box="[804,908,1269,1295]" name="Denmark" pageId="163" pageNumber="164">Denmark</collectingCountry>
by having only two spirals above the carina on the early whorls.
</paragraph>
</subSubSection>
</treatment>
</document>
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