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<document ID-DOI="http://dx.doi.org/10.3897/afrinvertebr.64.113133" ID-Pensoft-Pub="2305-2562-3-303" ID-Pensoft-UUID="1B4311FDBAF2573E8131BD435D38D7EF" ID-ZooBank="000F15D70DD746F7982625F13C43AF4C" ModsDocID="2305-2562-64-3-303" checkinTime="1702376364544" checkinUser="pensoft" docAuthor="Muller, Burgert S., Swart, Vaughn R. &amp; Snyman, Louwrens P." docDate="2023" docId="9F00609B1F1C5AF08EA89E840930D6E6" docLanguage="en" docName="AfrInvert 64(3): 303-322" docOrigin="African Invertebrates 64 (3)" docPubDate="2023-12-11" docSource="http://dx.doi.org/10.3897/afrinvertebr.64.113133" docTitle="Atrichops intermedius Muller 2023, sp. nov." docType="treatment" docUuid="B3B59959-1502-472A-89C2-C21017DE04D1" docUuidSource="ZooBank" docVersion="1" id="1B4311FDBAF2573E8131BD435D38D7EF" lastPageNumber="303" masterDocId="1B4311FDBAF2573E8131BD435D38D7EF" masterDocTitle="Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species" masterLastPageNumber="322" masterPageNumber="303" pageNumber="303" updateTime="1702376364544" updateUser="pensoft">
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<mods:title id="57804C5ABFECFF3CC3DF2E436AEB77E7">Afrotropical Atrichops Verrall (Diptera, Athericidae) with description of a new species</mods:title>
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<mods:namePart id="C939ED0D1DAEF81BC4CEF1996142475E">Muller, Burgert S.</mods:namePart>
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<mods:affiliation id="33E9343BC3D81EA1D87261FD381D77BF">Department Terrestrial Invertebrates, National Museum, 36 Aliwal Street, Bloemfontein, 9301, South Africa &amp; Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
<mods:nameIdentifier id="ADCC41804CD63CADD1330432CC513816" type="email">burgert.muller@nasmus.co.za</mods:nameIdentifier>
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<mods:namePart id="5F13882CFCF7EC1EEE28906D58A71D51">Swart, Vaughn R.</mods:namePart>
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<mods:affiliation id="F4C2A463BA899B1E4538C32AECC485B9">Department Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein, 9300, South Africa</mods:affiliation>
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<mods:namePart id="314C47A14DF5A6C1D9E2B1C0B8CE12DF">Snyman, Louwrens P.</mods:namePart>
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<mods:affiliation id="8C051F6B0A2B9BF6E6D8E692CB5DAF9E">Invertebrate Zoology, Royal Alberta Museum, 9810 103 A Avenue NW, Edmonton, Alberta, T 5 J 0 G 2, Canada</mods:affiliation>
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<mods:title id="73DB8766B93E41B59CAC3317B7C3D976">African Invertebrates</mods:title>
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<mods:date id="6A9C62A1777852D125A498D1478715A5">2023</mods:date>
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<taxonomicName id="98F69CE577A8660DB7517C2754989737" LSID="https://zoobank.org/B3B59959-1502-472A-89C2-C21017DE04D1" authority="Muller" authorityName="Muller" authorityYear="2023" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius" status="sp. nov.">Atrichops intermedius Muller</taxonomicName>
<taxonomicNameLabel id="A48C620D4D9BEA81F9E31960FF2F8F24" pageId="0" pageNumber="303">sp. nov.</taxonomicNameLabel>
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<figureCitation id="F87C4D831B58B3C84385506FB2F40C2F" captionStart="Figures 16" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">Figs 2</figureCitation>
<figureCitation id="35FBA78F5AB58C613E13E391C4D1898A" captionStart="Figures 16" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">, 5</figureCitation>
<figureCitation id="EAC4DA9660CF2CDA7667DA47F39A5E7C" captionStart="Figures 712" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">, 8</figureCitation>
<figureCitation id="93FEEEB56FF41D0DA1D89EBAFAF6D3B2" captionStart="Figures 712" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">, 11</figureCitation>
<figureCitation id="377176B540B9A25F58F03B31C920AD46" captionStart="Figures 1318" captionStartId="F3" captionText="Figures 13 - 18. Atrichops spp. ♂ ♀ dorsal view 13 ♂ A. adamastor (BMSA (D) 38908) 14 ♂ A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 15 ♂ A. stuckenbergi (BMSA (D) 94158) 16 ♀ A. adamastor holotype (MZLU 00205685) 17 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 18 ♀ A. stuckenbergi holotype (CAS). (16 modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0; 18 modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303">, 14</figureCitation>
<figureCitation id="B1B1EEFE1F4A8C9D262E87518EC9E30A" captionStart="Figures 1318" captionStartId="F3" captionText="Figures 13 - 18. Atrichops spp. ♂ ♀ dorsal view 13 ♂ A. adamastor (BMSA (D) 38908) 14 ♂ A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 15 ♂ A. stuckenbergi (BMSA (D) 94158) 16 ♀ A. adamastor holotype (MZLU 00205685) 17 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 18 ♀ A. stuckenbergi holotype (CAS). (16 modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0; 18 modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303">, 17</figureCitation>
<figureCitation id="FD98451E635E213C51BA0944DA4B1EC8" captionStart="Figures 1924" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">, 20</figureCitation>
<figureCitation id="71A47198E202A5C2564A3B75E6046D84" captionStart="Figures 1924" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">, 23</figureCitation>
<figureCitation id="291FEDB0F39D849078B08BD5B0D06EFE" captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">, 27</figureCitation>
<figureCitation id="F0227757CBDB895CF0C855E48EFDB4FA" captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">, 28</figureCitation>
<figureCitation id="12CE05842C683D146FC34CE0A136F204" captionStart="Figures 3136" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">, 32</figureCitation>
<figureCitation id="61580789EEA1FAED950947D4063E0DC5" captionStart="Figures 3136" captionStartId="F6" captionText="Figures 31 - 36. Atrichops spp., 31 - 33 ♂ terminalia 31 A. adamastor (BMSA (D) 38904) 32 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 33 A. stuckenbergi (BMSA (D) 94160), 34 - 36 ♂ epandrium and cerci 34 A. adamastor (BMSA (D) 38904) 35 A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 36 A. stuckenbergi (BMSA (D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303">, 35</figureCitation>
<figureCitation id="93BE0A427BB2F2E3102F2581AB6C331B" captionStart="Figures 3739" captionStartId="F7" captionText="Figures 37 - 39. Atrichops spp., ♀ genital fork and spermatheca 37 A. adamastor (BMSA (D) 38899 38 A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 39 A. stuckenbergi (BMSA (D) 95855). Abbreviations: gen fk - genital fork; spmth - spermatheca. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures37-39" httpUri="https://binary.pensoft.net/fig/950655" pageId="0" pageNumber="303">, 38</figureCitation>
</paragraph>
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<paragraph id="091529487FD5629F395C8E6AE90BA159" pageId="0" pageNumber="303">Material examined.</paragraph>
<paragraph id="FA88C9E7868AE3081F050FCCCDD6B5BA" pageId="0" pageNumber="303">
<emphasis id="051B4B84646717014E493D5C5E5BFAEF" bold="true" pageId="0" pageNumber="303">
Type material:
<materialsCitation id="DB2D0A169C18C404EE85B5193C08B3F1" collectingDate="1971-02-23" collectorName="Stuckenberg, B. R" country="South Africa" county="Gladdespruit River" elevation="907" latitude="-25.511171" location="Transvaal" longLatPrecision="1" longitude="30.907625" municipality="Nelspruit" specimenCount="1" specimenCount-male="1" stateProvince="Mpumalanga" typeStatus="Holotype">
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<typeStatus id="BA94EBA752A02DBECB6CB78D02AF5BCA">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="B067A6B8690A04C6EFB5851987AE69CA" name="South Africa">South Africa</collectingCountry>
<specimenCount id="1370560260EEF49DF557E413CC10CD75" type="male">1♂</specimenCount>
;
<collectingRegion id="A2E0FA84D9B2F4838C4C5E8526185BD7" country="South Africa" name="Mpumalanga">Mpumalanga</collectingRegion>
,
<collectingCounty id="DB27FF07756856092CCF4955E76DBB58">Gladdespruit River</collectingCounty>
nr
<collectingMunicipality id="5E011D242406D5ED17CE5B441428F428">Nelspruit</collectingMunicipality>
[Mbombela] airfield,
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; [
<geoCoordinate id="895B6945752B55C337375C600BD92B98" degrees="25" direction="south" minutes="30.6703" orientation="latitude" precision="1" value="-25.511171">25°30.6703'S</geoCoordinate>
,
<geoCoordinate id="3989CE537DE9A05135088946E75A0E52" degrees="30" direction="east" minutes="54.4575" orientation="longitude" precision="1" value="30.907625">30°54.4575'E</geoCoordinate>
]; 2530
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;
<quantity id="D368FA06AF89710756D949E147552D12" metricMagnitude="2" metricUnit="m" metricValue="9.0678" unit="ft" value="2975.0">
<elevation id="602C14B422C8E63FE1F9AF3F29FC83C6" metricMagnitude="2" metricUnit="m" metricValue="9.0678" unit="ft" value="2975.0">2975 ft</elevation>
</quantity>
[
<elevation id="B24E363285061DA159EB418F7323DEC6" metricMagnitude="2" metricUnit="m" metricValue="9.07" unit="m" value="907.0">
<quantity id="56742CF4FEDC95EEDD274161542A7946" metricMagnitude="2" metricUnit="m" metricValue="9.07" unit="m" value="907.0">907 m</quantity>
asl
</elevation>
];
<collectingDate id="4FC795F47D3857D26DC0A10076F29049" value="1971-02-23">23 Feb. 1971</collectingDate>
; Stuckenberg [
<collectorName id="701E41E55EADB82B0C95D0488AA15963">Stuckenberg, B.R</collectorName>
] leg.; streamside bush; (NMSA-DIP 81808) (NMSA).
</materialsCitation>
</emphasis>
</paragraph>
<paragraph id="1D190BCF4B3CDEE079B7059D18FCA0FE" pageId="0" pageNumber="303">
<materialsCitation id="D27746675886DAE2374154A89D2FE82D" collectingDate="1971-02-23" collectorName="Stuckenberg, B. R" country="South Africa" county="Gladdespruit River" elevation="907" latitude="-25.511171" location="Transvaal" longLatPrecision="1" longitude="30.907625" municipality="Nelspruit" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Mpumalanga" typeStatus="Paratypes">
<emphasis id="EFB517ED70922717B80AC9BC174C76DD" bold="true" italics="true" pageId="0" pageNumber="303">
<typeStatus id="E2A49AC09B4C84CD9FA052F2BFC01331">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="AA3BCDDB68056758D385B33C5A4F60C3" name="South Africa">South Africa</collectingCountry>
<specimenCount id="380E731D2F77EE69D573200766F99877" type="male">2♂</specimenCount>
<specimenCount id="B1273AE06069EAB2BF1A63DD73BA4051" type="female">2♀</specimenCount>
;
<collectingRegion id="933CEF53559262EBE964032B49FC7522" country="South Africa" name="Mpumalanga">Mpumalanga</collectingRegion>
,
<collectingCounty id="F14BED3BD06B809BCAEF9574CB11F2C8">Gladdespruit River</collectingCounty>
nr
<collectingMunicipality id="1B40418711E2A6D4E72E2F283A9D198D">Nelspruit</collectingMunicipality>
[Mbombela] airfield,
<location id="34202C73E25CCCB69CCB5300052912BA" LSID="urn:lsid:plazi:treatment:9F00609B1F1C5AF08EA89E840930D6E6:34202C73E25CCCB69CCB5300052912BA" country="South Africa" county="Gladdespruit River" latitude="-25.511171" longLatPrecision="1" longitude="30.907625" municipality="Nelspruit" name="Transvaal" stateProvince="Mpumalanga">Transvaal</location>
; [
<geoCoordinate id="987024D6DCEE70F9013757E4D3503BD9" degrees="25" direction="south" minutes="30.6703" orientation="latitude" precision="1" value="-25.511171">25°30.6703'S</geoCoordinate>
,
<geoCoordinate id="BE400171937B67A8F2B1CB5C0ECE5FDE" degrees="30" direction="east" minutes="54.4575" orientation="longitude" precision="1" value="30.907625">30°54.4575'E</geoCoordinate>
]; 2530
<location id="D332050FAB7A5DF531EA6EB97434E9FC" LSID="urn:lsid:plazi:treatment:9F00609B1F1C5AF08EA89E840930D6E6:D332050FAB7A5DF531EA6EB97434E9FC" country="South Africa" county="Gladdespruit River" latitude="-25.511171" longLatPrecision="1" longitude="30.907625" municipality="Nelspruit" name="Db" stateProvince="Mpumalanga">Db</location>
;
<quantity id="556F97191CBB1C6C689BD3ECB58E2D39" metricMagnitude="2" metricUnit="m" metricValue="9.0678" unit="ft" value="2975.0">
<elevation id="BBDE6C9C120F9A5D22E68C6E7C9A6A78" metricMagnitude="2" metricUnit="m" metricValue="9.0678" unit="ft" value="2975.0">2975 ft</elevation>
</quantity>
[
<elevation id="807E4A37CCE603123AF44A92D316AECA" metricMagnitude="2" metricUnit="m" metricValue="9.07" unit="m" value="907.0">
<quantity id="D522C7CF63B8AB1AE5F5CEB5D4BCBF6E" metricMagnitude="2" metricUnit="m" metricValue="9.07" unit="m" value="907.0">907 m</quantity>
asl
</elevation>
];
<collectingDate id="10CF90F2A9263924ADC716350F1E7509" value="1971-02-23">23 Feb.1971</collectingDate>
; Stuckenberg [
<collectorName id="E6CE2B19788CACFFD47E121D213D51A0">Stuckenberg, B.R</collectorName>
] leg.; streamside bush; (
<specimenCount id="E79CBE44F52C6D4447BCBF0E089FE090" type="male">2♂</specimenCount>
NMSA-DIP 81811, NMSA-DIP 28122;
<specimenCount id="210717AED294C7FFBF22702B46082103" type="female">1♀</specimenCount>
NMSA-DIP 81809) (NMSA)
</materialsCitation>
.
<materialsCitation id="B8A0821B5DA89AF91E3F16CE62AAA2E3" collectingDate="2000-03-06" collectorName="Picker, M." country="South Africa" latitude="-24.9258" location="Graskop" longLatPrecision="1" longitude="30.829988" specimenCount="1" specimenCount-female="1" stateProvince="Mpumalanga" typeStatus="paratype">
<specimenCount id="653ACF1458D1891DE5C935DC2A655AC6" type="female">1♀</specimenCount>
;
<collectingRegion id="56AE854B2703A22DD2535D0B1079F830" country="South Africa" name="Mpumalanga">Mpumalanga</collectingRegion>
,
<location id="3923008D999E41DF94BB2B877171395D" LSID="urn:lsid:plazi:treatment:9F00609B1F1C5AF08EA89E840930D6E6:3923008D999E41DF94BB2B877171395D" country="South Africa" latitude="-24.9258" longLatPrecision="1" longitude="30.829988" name="Graskop" stateProvince="Mpumalanga">Graskop</location>
, [
<geoCoordinate id="FEC756D10F821AA5BE8EA2F7D8C47221" degrees="24" direction="south" minutes="55.5480" orientation="latitude" precision="1" value="-24.9258">24°55.5480'S</geoCoordinate>
,
<geoCoordinate id="E6FD422917365CC6AF542E0346B0D057" degrees="30" direction="east" minutes="49.7993" orientation="longitude" precision="1" value="30.829988">30°49.7993'E</geoCoordinate>
];
<collectingDate id="A43AA8365F1AA9883C3FED9964589B04" value="2000-03-06">6 Mar. 2000</collectingDate>
;
<collectorName id="E7CCCA83DB20F0DB0A8A867C303DE08A">Picker, M.</collectorName>
leg.; (NMSA-DIP 212861) (NMSA)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5CD7EE63AB203A36A266725DFD827EDB" pageId="0" pageNumber="303" type="diagnosis">
<paragraph id="8A1FF260DCA269CC875317FCA22FE00C" pageId="0" pageNumber="303">Diagnosis.</paragraph>
<paragraph id="147E465AC6DF5B85806258F7D2414577" pageId="0" pageNumber="303">
This species is most similar to
<taxonomicName id="B3AE884E14B8F5B710F48190B01AF3F6" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="82394175A2561E5E2DA1BBF9E5E9CA31" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
. However, it differs from
<taxonomicName id="156B6CAA6E68939C79A6566ED15A8AD1" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="6F7580985558276C09BB49093D605468" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
in several ways: ♂ more narrowly dichoptic, ♀ with a darkened transverse line between the face and clypeus, ♂ legs nearly unicolorous yellowish compared to both males of
<taxonomicName id="CE547417A153C1F0ED85D80AFEF7FF2A" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="E61A3CB8822B504BDE4449E038DA8C03" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
and
<taxonomicName id="5BE2BDED4352DB2F266A69A1B75F3058" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="B10FA208CB6661610E821F703113F639" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
, which have yellow hind femora and all tibiae and tarsi darker; Additionally,
<taxonomicName id="87DF9650B8A7F7A2F9BADFD1AC09A6D1" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="E3C84A68A7E36135281E57A14803C726" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
</taxonomicName>
has wings with darker brown suffusions over base and apex of the discal cell that are similar to
<taxonomicName id="1878CD3C72A157C2B01CF6C400B502AB" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="491F4C365E28DC8E259F974970E34A0E" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
, in contrast to
<taxonomicName id="955CFE9CCB4BA79DC237C969F7C0D31B" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="47351A24FCD2E79575E4651B453A929C" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
which has a more uniformly suffused wing. Its gonocoxite is also more rounded and less tapering toward base compared to both
<taxonomicName id="A1D84B5BFE109B377A68C9A18B6656FC" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="B845B402CFFE57F848A0559826E9F728" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
and
<taxonomicName id="6BE90374BD6A63E7FB82B9747411A64D" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="3881A89AD3B6B80C9FCF18042F32D3F4" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="68EB687BA35572EEA633E4556AC31773" pageId="0" pageNumber="303" type="description">
<paragraph id="B060C108E58A7EC32C81253F1BA3C4B5" pageId="0" pageNumber="303">Description.</paragraph>
<paragraph id="E35B1D46D1A86FE51885421167811A5F" pageId="0" pageNumber="303">
<emphasis id="4F7D62461AAE04F969C6DE509D040993" bold="true" italics="true" pageId="0" pageNumber="303">Measurements</emphasis>
(♂ n = 2, ♀ n = 2): Wing span: ♂ 5.4-5.8 mm (avg. 5.6 mm); ♀ 5.9-6.5 mm (avg. 6.2 mm); body length: ♂ 5.5 mm-6.1 mm (avg. 5.8 mm); ♀ 4.8-5.4 mm (avg. 5.1 mm); wing span to body length ratio (avg.): ♂ 0.96; ♀ 1.2.
</paragraph>
<paragraph id="D3F3746BCC025BF60949DEEDFDBB1D03" pageId="0" pageNumber="303">
<emphasis id="C5163F787A6B11D39A90CFB6FD519D70" bold="true" pageId="0" pageNumber="303">
Male.
<emphasis id="3645A01D13226D6E43BD5230FCD05C9C" italics="true" pageId="0" pageNumber="303">Head</emphasis>
</emphasis>
(Fig.
<figureCitation id="BA849D1F37E3A14B216060ABF2CA2FCD" captionStart="Figures 16" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">5</figureCitation>
): Colour dark brown, with some yellowish pruinosity on face; eye densely covered in ommatrichia; narrowly dichoptic; ommatidia on lower
<normalizedToken id="A60408E29B317595283C34099CEBE278" originalValue="½">1/2</normalizedToken>
of eye smaller than upper
<normalizedToken id="2E2B76917279626456D33EA39BC21D82" originalValue="½">1/2</normalizedToken>
; lateral edge of eye with indentation (absent in ♀); ocellar tubercle base level with frons, ocellar tubercle with short dark setulae; vertex dark brown, almost appearing black, with markedly longer setulae than ocellar tubercle, similar to that of frons; ocelli similar in size; ocellar tubercle in front of dorsal margin of eye, margin less indented than in ♀; vertex narrower than in ♀; dorsal inner edge of eye, surrounding ocellar tubercle without discernible paired dark markings; occiput same dark brown as frons, when viewed at angle appearing shiny blackish; upper occiput with short dark setulae on dorsal margin and on rest of upper surface, lower occiput with erect, long and pale setulae, these continue ventrally on head to before mouthparts bearing dark ventral setulae; frons dark brown, almost black, narrow area above antenna lighter brown; frons at narrowest slightly narrower than anterior ocellus, widening towards antennal base; frons with well-developed dark setulae on surface except for narrow area above antennae, these similar to ocellar and vertical setulae; frons less setulose than ♀; face lateral edges bare; gena bare; face light brown, clypeus darker brown; clypeus bare; face separated from clypeus by prominent, deep, dark transverse suture; clypeus less prominent than in ♀; face much wider than in ♀, with prominent longitudinal emarginations on sides of clypeus, giving appearance that face bulges laterally; face and clypeus not visible in profile; antennal bases close together, almost touching; scape light-brown, dorsally infuscate around setulae; pedicel comparatively darker than scape, dorsal surface infuscate; scape and pedicel setulae dark; 1st flagellomere reniform; appearing almost 2
<normalizedToken id="CBA63C3CE221FC53154000F73BB84D71" originalValue="×">x</normalizedToken>
the height of pedicel, same colour as pedicel, basal margins lighter brown; 2nd flagellomere arista-like, dark brown; scape and pedicel setulae similar in size, palpus brown, well-developed, ca 0.5
<normalizedToken id="0D87664080B75FA888F338C5B26E27FE" originalValue="×">x</normalizedToken>
length of proboscis; proboscis comparatively smaller than ♀ proboscis in relation to head; proboscis dark brown with long dark setulae.
</paragraph>
<paragraph id="915FE139AC6E5F2DD85EFA21F506E5B3" pageId="0" pageNumber="303">
<emphasis id="ED8A08079D266F3DAEB9B1826F638688" bold="true" italics="true" pageId="0" pageNumber="303">Thorax</emphasis>
(Figs
<figureCitation id="F575B34790EEFA1D87D6579F6C12A8CA" captionStart="Figures 1318" captionStartId="F3" captionText="Figures 13 - 18. Atrichops spp. ♂ ♀ dorsal view 13 ♂ A. adamastor (BMSA (D) 38908) 14 ♂ A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 15 ♂ A. stuckenbergi (BMSA (D) 94158) 16 ♀ A. adamastor holotype (MZLU 00205685) 17 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 18 ♀ A. stuckenbergi holotype (CAS). (16 modified from original provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0; 18 modified from original provided by Rikio Matsumoto, OMNH). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures13-18" httpUri="https://binary.pensoft.net/fig/950650" pageId="0" pageNumber="303">14</figureCitation>
,
<figureCitation id="BC7B0C6AB222B8F2C59FC96EF582EE6E" captionStart="Figures 1924" captionStartId="F4" captionText="Figures 19 - 24. Atrichops spp. ♂ ♀ proepimeral process, lateral view 19 ♂ A. adamastor (BMSA (D) 38904) 20 ♂ A. intermedius sp. nov. paratype (NMSA-Dip. 81811) 21 ♂ A. stuckenbergi (BMSA (D) 94158) 22 ♀ A. adamastor (BMSA (D) 38899) 23 ♀ A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 24 ♀ A. stuckenbergi holotype (CAS). (24 modified from original provided by Rikio Matsumoto, OMNH). Abbreviations: prepm - proepimeral process." figureDoi="10.3897/afrinvertebr.64.113133.figures19-24" httpUri="https://binary.pensoft.net/fig/950652" pageId="0" pageNumber="303">20</figureCitation>
): Scutum with short dark setulae, postsutural setulae longer than presutural setulae; scutellum with well-developed dark setulae; postpronotal lobe lighter brown than scutum, with fine dark setulae; scutum and scutellum uniformly dark brown when viewed dorsally, scutellum similar in colour as anepimeron, anatergite and katatergite when viewed laterally; pleura generally brown in colour, anepimeron, katatergite and anatergite lighter brown than rest of pleura (more pronounced in ♀); proepimeron with reduced knoblike process near anterior edge; notopleuron with well-developed dark setulae; area surrounding posterior spiracle dark brown, postspiracular scale dark brown almost black; proepisternum brown and pronotum yellow; anterior spiracle bare posteriorly; proepimeron and proepisternum bare, anepisternum with short dark setulae; katatergite with long pale setulae; rest of pleura bare; postmetacoxal bridge narrow.
</paragraph>
<paragraph id="8D3B6E60E3F333781AA6FB2085DD3535" pageId="0" pageNumber="303">
<emphasis id="5B4CFED6612800FD4628A8F96FDF45AA" bold="true" italics="true" pageId="0" pageNumber="303">Legs</emphasis>
(Fig.
<figureCitation id="9A561B5D1ECF991161FD5C85B3CE568C" captionStart="Figures 16" captionStartId="F1" captionText="Figures 1 - 6. Atrichops spp. ♂ 1 - 3 lateral habitus 4 - 6 head anterior view 1 A. adamastor (BMSA (D) 38908) 2 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 3 A. stuckenbergi (BMSA (D) 95848) 4 A. adamastor (BMSA (D) 38904) 5 A. intermedius sp. nov. holotype (NMSA-Dip. 81808) 6 A. stuckenbergi (BMSA (D) 95848). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures1-6" httpUri="https://binary.pensoft.net/fig/950648" pageId="0" pageNumber="303">2</figureCitation>
): Coxae brown; fore coxa with short black setulae on surface, more pronounced apically, mid coxa with long black setulae on surface, hind coxa with black setulae on anterior and lateral apical edges, with well-developed anterior apical point; trochanters brownish yellow with some scattered short dark setulae; hind trochanter somewhat lighter than rest; apical margins of trochanters with darker markings; fore and mid femora, and basal
<normalizedToken id="C362BD2F8B5557AD017126317E29DD2F" originalValue="¼">1/4</normalizedToken>
of hind femur yellow, apical
<normalizedToken id="F6D437EB3CD4F7EE7A310BFBD47275CB" originalValue="¾">3/4</normalizedToken>
of hind femur dark brown; fore, mid and hind femora with small anterior apical dark mark; fore, mid and hind tibiae yellow; tarsi dark brown; fore tarsal claws symmetrical, empodium at least 2
<normalizedToken id="90662D42966DB73969944E128E03E7C7" originalValue="×">x</normalizedToken>
size of pulvilli on fore legs; fore femur covered with dark setulae on all surfaces, posteroventral setulae forming row, mid femur with row of longer anteroventral setae and hind femur with similar dark setulae on ventral and dorsal surfaces; hind leg stouter than fore and mid legs; fore tarsi densely covered with long setulae along dorsal and ventral surfaces, at least as long as width of segment, similar setulae on fore tibia, although much sparser; hind tarsal segments 0.9-1.1 (♂) as long as hind tibia.
</paragraph>
<paragraph id="862D72C7807B9A9F6FCA30110B5FB1BD" pageId="0" pageNumber="303">
<emphasis id="04F0AC806107CC87AFAEB9148771C040" bold="true" italics="true" pageId="0" pageNumber="303">Wing</emphasis>
(Fig.
<figureCitation id="4E5023F40E3039F7883AFF6CB972AC97" captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">27</figureCitation>
): Suffused brown on entire surface, with much darker pterostigma over area of veins
<emphasis id="A2491EADFAE4C72D0D5EB71B608C7AE0" italics="true" pageId="0" pageNumber="303">R1</emphasis>
and
<emphasis id="5318D356B4CE6471E0B9A7231E9A61C8" italics="true" pageId="0" pageNumber="303">R2+3</emphasis>
and cell
<emphasis id="8B0B089EE57CA7D03E034C57AA070448" italics="true" pageId="0" pageNumber="303">r1</emphasis>
; veins light brown; cell
<emphasis id="DC366D9B56B24518269F3AA86625F6CB" italics="true" pageId="0" pageNumber="303">br</emphasis>
and discal cell darker suffused apically; costa with distinct downward flexure over pterostigma; cell
<emphasis id="FEAE15FDBDB890A5A756EEBB9C925C9A" italics="true" pageId="0" pageNumber="303">cua</emphasis>
closed short distance from wing margin, cell
<emphasis id="201E0A468C94BE27EB8C8D53D9B81B5B" italics="true" pageId="0" pageNumber="303">m3</emphasis>
open, veins
<emphasis id="BA48DAAD89606947932A9632C531EEBF" italics="true" pageId="0" pageNumber="303">M1</emphasis>
,
<emphasis id="3F56A364072210506DB8D4785E1D249C" italics="true" pageId="0" pageNumber="303">M2</emphasis>
,
<emphasis id="2C10396FDED44B32D264629BBAB8CFE7" italics="true" pageId="0" pageNumber="303">M3</emphasis>
present; halter with stalk yellow on basal half, gradually darkening apically with dark brown knob.
</paragraph>
<caption id="774EF8E897ADA49F249F764A1AEF1B07" doi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303" start="Figures 2530" startId="F5">
<paragraph id="39504748B283B38C49EA6C7F49A9CF65" pageId="0" pageNumber="303">
<emphasis id="16A1771101513443E40C5300E5576932" bold="true" pageId="0" pageNumber="303">Figures 25-30.</emphasis>
<taxonomicName id="45866AEF2400F5DE534E9EE2AD9A50AE" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="8411D9DEA54C084614E8A1FF54E5637E" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
</taxonomicName>
spp. ♂♀ wings dorsal view,
<taxonomicName id="3398FC954C58D041FCF1373B6B2BAD9D" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="65C0352D31C3FBBFDEDB7F80A7DFFEF1" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
:
<emphasis id="EB2DCA17E4572C20DFD16A816F1CD3F8" bold="true" pageId="0" pageNumber="303">25</emphasis>
♂ (BMSA(D)38904)
<emphasis id="7A3230A2833F5510BB364021B85146C3" bold="true" pageId="0" pageNumber="303">26</emphasis>
♀ (BMSA(D)38904);
<taxonomicName id="B934F00F4D943A0F1C9640C11F1147CD" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="4CBA99FA01673B4AE2131C4F7B59D0D8" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
</taxonomicName>
sp. nov.:
<emphasis id="969091B6E0C2BD95991470F611115C12" bold="true" pageId="0" pageNumber="303">27</emphasis>
♂ paratype (NMSA-Dip. 81811)
<emphasis id="D78D3396EF0F5825C2A13B1AADC64938" bold="true" pageId="0" pageNumber="303">28</emphasis>
♀ paratype (NMSA-Dip. 212861)
<taxonomicName id="1706B0C0CC762B6DEF67FF695E21F303" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="509D7E7CA32463144E80C2DC7ED0E10A" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
:
<emphasis id="80A1F190B3DC42926F50E9884274468F" bold="true" pageId="0" pageNumber="303">29</emphasis>
♂ (BMSA(D)94161)
<emphasis id="69A939A556D2164AACCAFCF580BA9141" bold="true" pageId="0" pageNumber="303">30</emphasis>
♀ (BMSA(D)95861). Abbreviations:
<emphasis id="458E5FD21CD9EE94A8C70753842C8BFD" italics="true" pageId="0" pageNumber="303">br</emphasis>
- basal radial cell;
<emphasis id="E6D626CBB2EA3180A51DA48EBBA3AB9F" italics="true" pageId="0" pageNumber="303">bm</emphasis>
- basal medial cell;
<emphasis id="462903C8C40A3C92FAEFD0BDE6B9FE4F" italics="true" pageId="0" pageNumber="303">cua</emphasis>
- anterior cubital cell;
<emphasis id="3DDEEF2F8BF1A4E0A84C9463F22AC954" italics="true" pageId="0" pageNumber="303">CuA</emphasis>
+
<emphasis id="7615979B40DECB6CFE9090B29C8B994E" italics="true" pageId="0" pageNumber="303">CuP</emphasis>
- anterior branch of cubital vein + posterior branch of cubital vein;
<emphasis id="1F056DB6FCC61B9C0EB67996A42B56D1" italics="true" pageId="0" pageNumber="303">d</emphasis>
- discal cell;
<emphasis id="70567FC8E94A63F01E22CF70CE206FB0" italics="true" pageId="0" pageNumber="303">M1</emphasis>
- first branch of media;
<emphasis id="259F640FB516DB5C838AB6B74DD62B6E" italics="true" pageId="0" pageNumber="303">m1</emphasis>
- first medial cell;
<emphasis id="5968811DC4978FE1C90F9FE7F39FCF9C" italics="true" pageId="0" pageNumber="303">M2</emphasis>
- second branch of media;
<emphasis id="313CDBF2233A2D54D5AA8E8410124F2A" italics="true" pageId="0" pageNumber="303">m2</emphasis>
- second medial cell;
<emphasis id="C6B8337AF59145622D7C320D2941DDDF" italics="true" pageId="0" pageNumber="303">M3</emphasis>
- third branch of media;
<emphasis id="9AA299A6100A84470DA6B0207615E78E" italics="true" pageId="0" pageNumber="303">m3</emphasis>
- third medial cell;
<emphasis id="A867EA59613587CA45B3F33E935698B8" italics="true" pageId="0" pageNumber="303">R1</emphasis>
- anterior branch of radius;
<emphasis id="E1A078C8119F606935C88DD90D2B5CAB" italics="true" pageId="0" pageNumber="303">r1</emphasis>
- first radial cell;
<emphasis id="72680C1996D21ED74615C5A627147352" italics="true" pageId="0" pageNumber="303">R2+3</emphasis>
- second branch of radius;
<emphasis id="B0B3CC587A0C3226813B276BC3B80748" italics="true" pageId="0" pageNumber="303">r2+3</emphasis>
- second radial cell;
<emphasis id="01EB110D3E9CC549B8AD17A36D05FADE" italics="true" pageId="0" pageNumber="303">R4</emphasis>
- upper branch of third branch of radius;
<emphasis id="537D1B0BA712BCCCBF3B30CB4469F929" italics="true" pageId="0" pageNumber="303">R5</emphasis>
- lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="FD12D0DDD7B8BA1B4760BE75C298B083" doi="10.3897/afrinvertebr.64.113133.figures31-36" httpUri="https://binary.pensoft.net/fig/950654" pageId="0" pageNumber="303" start="Figures 3136" startId="F6">
<paragraph id="FC23A34B93B06FDED348E42F123C364C" pageId="0" pageNumber="303">
<emphasis id="A35B9848AE928DB0D52DE5565D2FB1B5" bold="true" pageId="0" pageNumber="303">Figures 31-36.</emphasis>
<taxonomicName id="5F62BD2EA51F6302796681A5DB4F665A" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="7B1CBEEBECE7BFB9CBFFE03BECE9C78D" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
</taxonomicName>
spp., 31-33 ♂ terminalia
<emphasis id="7648AEFCC4E8DF5FCA993DA1DC50F054" bold="true" pageId="0" pageNumber="303">31</emphasis>
<taxonomicName id="B275376B6D149031E579BA469024C2DC" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="84A02CC8CA0DFBDF3A60B6A40BFBD418" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
(BMSA(D)38904)
<emphasis id="AFDC7102B379727EEC726310822F1DA3" bold="true" pageId="0" pageNumber="303">32</emphasis>
<taxonomicName id="5885892B3579F457E76B10043D997F11" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="E39EE94B0A2E981C01F761C240173A82" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
</taxonomicName>
sp. nov. paratype (NMSA-Dip. 81811)
<emphasis id="0B29F4C4C85D1899107D136ECD5A83CE" bold="true" pageId="0" pageNumber="303">33</emphasis>
<taxonomicName id="B7CBDDA4BA528B61036885DDF8963F67" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="A8C184EB7031A86E740E3D8E130B1867" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
(BMSA(D) 94160), 34-36 ♂ epandrium and cerci
<emphasis id="CE7BCF35671921A61FC95FB5A7123831" bold="true" pageId="0" pageNumber="303">34</emphasis>
<taxonomicName id="76926BE9C217481FA74030DD0B9CD220" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="57B6F3587ECF2D22D1C6B1339B956EEE" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
(BMSA(D)38904)
<emphasis id="E97C617AE7134BE79A3C0A44C538E16C" bold="true" pageId="0" pageNumber="303">35</emphasis>
<taxonomicName id="B9DDE72B12AC350FBC169986EC4B1EC6" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="D7C5EF24FD3C00480E1DE5CB6521B420" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
</taxonomicName>
sp. nov. paratype (NMSA-Dip. 81811)
<emphasis id="CE1E0A622C58396CA76D1D452790BE5A" bold="true" pageId="0" pageNumber="303">36</emphasis>
<taxonomicName id="D76F329B10AFE526F4C5052B65E874FF" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="C85A4FD8565D6A7C8E868CD6B23E3E17" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
(BMSA(D) 94160). Abbreviations: aed tn - aedeagal tine; cerc - cercus; ej apod - ejaculatory apodeme; epand - epandrium; goncx - gonocoxite; goncx apod - gonocoxal apodeme; gonst - gonostylus; pm - paramere; pm apod - parameral apodeme; pm sh - parameral sheath.Scale bars: 0.15 mm.
</paragraph>
</caption>
<paragraph id="B4ED17AFF3F8C23CD8E3C26785E604C7" pageId="0" pageNumber="303">
<emphasis id="00DD127C868F6CB1318022C900A5B384" bold="true" italics="true" pageId="0" pageNumber="303">Abdomen</emphasis>
: Brown, with anterodorsal margins of tergites 2-4 brownish yellow; tergites and sternites similar in colour with short black setulae, except sternite 1 lighter in colour; tergites with longer black setulae laterally; long pale setulae on sternites 1-3; tergite 1 without median suture.
</paragraph>
<paragraph id="B882322ADA9FCB12949779D341079E45" pageId="0" pageNumber="303">
<emphasis id="E8457F3F1D544D0E1BFF0F7E3D7252F9" bold="true" italics="true" pageId="0" pageNumber="303">Terminalia</emphasis>
: Epandrium and cerci dark brown, hypoproct dark brown; epandrium, hypandrium and cercus with dark setulae; gonostylus finger-like, narrowing slightly towards apex, gonocoxite widening from middle towards base, apically rounded with long setulae; parameral apodeme short, not reaching base of gonocoxite in ventral view; gonocoxal apodeme markedly shorter than gonocoxite.
</paragraph>
<paragraph id="FC1EA37FADA58102CE8501475C812925" pageId="0" pageNumber="303">
<emphasis id="862DD4024C9A552DBFCCB9BFD04D251F" bold="true" pageId="0" pageNumber="303">Female.</emphasis>
Similar characters as ♂ except for the following:
</paragraph>
<paragraph id="7FD6A542323F8383EFEA9E5F26EE782F" pageId="0" pageNumber="303">
<emphasis id="EB98D93C0131F3F3B9C854906E9526DF" bold="true" italics="true" pageId="0" pageNumber="303">Head</emphasis>
(Fig.
<figureCitation id="98F3892346013EB96021E8CD7011A206" captionStart="Figures 712" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">11</figureCitation>
): Widely dichoptic, ommatidia of similar size, comparatively larger than in ♂; lateral edge of eye without indentation (present in ♂); ocellar tubercle base with sunken appearance; ocellar tubercle with slightly longer setulae than in ♂; dorsal margin of eye more indented than in ♂; vertex wider than in ♂; dorsal inner edge of eye, surrounding ocellar tubercle with paired dark markings; frons at least 2
<normalizedToken id="22976FDC814ED077EB6AF48143FB8508" originalValue="×">x</normalizedToken>
width of ocellar tubercle, narrowing only slightly towards antennal base; frons more setulose than ♂; face separated from clypeus by transverse emargination with darkened transverse line; clypeus more prominent than in ♂; face and clypeus visible in profile; 1st flagellomere smaller in comparison to pedicel compared to ♂; 2nd flagellomere ca 0.75
<normalizedToken id="172C4788F3AA4FCAEAC264D309B9EEE8" originalValue="×">x</normalizedToken>
length of proboscis; proboscis comparatively larger than ♂ proboscis in relation to head.
</paragraph>
<paragraph id="47ED5FB3592E026445AE0686418413CF" pageId="0" pageNumber="303">
<emphasis id="2D37C136CE37CF3762CA450BFE889A2B" bold="true" italics="true" pageId="0" pageNumber="303">Legs</emphasis>
(Fig.
<figureCitation id="3D77E0129AD07D01ACAC21880A7246FB" captionStart="Figures 712" captionStartId="F2" captionText="Figures 7 - 12. Atrichops spp. ♀ 7 - 9 lateral habitus 10 - 12 head anterior view 7 A. adamastor holotype and specimen labels (MZLU 00205685) 8 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 9 A. stuckenbergi (CAS) 10 A. adamastor holotype (MZLU 00205685) 11 A. intermedius sp. nov. paratype (NMSA-Dip. 81809) 12 A. stuckenbergi (BMSA (D) 95856). (7, 10 modified from originals provided by Rune Bygebjerg, MZLU, CC BY-NC 2.0 9, 12 modified from originals provided by Rikio Matsumoto, Osaka Museum of Natural History). Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures7-12" httpUri="https://binary.pensoft.net/fig/950649" pageId="0" pageNumber="303">8</figureCitation>
): Trochanters yellow; apical margins of trochanters with darker markings, less pronounced than in ♂; all femora almost entirely yellow, except for small anterior apical dark marking on each femur; tarsi yellow; fore tarsi without prominent long setulae on dorsal and ventral surfaces; hind tarsal segments 0.8-1.0 (♀) times as long as hind tibia.
</paragraph>
<paragraph id="CACB05DFB4E1200D0E0765D3E3CD5EA6" pageId="0" pageNumber="303">
<emphasis id="39AE26EF260437434EF792EA01A63851" bold="true" italics="true" pageId="0" pageNumber="303">Wing</emphasis>
(Fig.
<figureCitation id="706FB3617B9DFA56248AD570CE3DF06E" captionStart="Figures 2530" captionStartId="F5" captionText="Figures 25 - 30. Atrichops spp. ♂ ♀ wings dorsal view, A. adamastor: 25 ♂ (BMSA (D) 38904) 26 ♀ (BMSA (D) 38904); A. intermedius sp. nov.: 27 ♂ paratype (NMSA-Dip. 81811) 28 ♀ paratype (NMSA-Dip. 212861) A. stuckenbergi: 29 ♂ (BMSA (D) 94161) 30 ♀ (BMSA (D) 95861). Abbreviations: br - basal radial cell; bm - basal medial cell; cua - anterior cubital cell; CuA + CuP - anterior branch of cubital vein + posterior branch of cubital vein; d - discal cell; M 1 - first branch of media; m 1 - first medial cell; M 2 - second branch of media; m 2 - second medial cell; M 3 - third branch of media; m 3 - third medial cell; R 1 - anterior branch of radius; r 1 - first radial cell; R 2 + 3 - second branch of radius; r 2 + 3 - second radial cell; R 4 - upper branch of third branch of radius; R 5 - lower branch of third branch of radius; Sc - subcostal vein; sc - subcostal cell. Scale bars: 1 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures25-30" httpUri="https://binary.pensoft.net/fig/950653" pageId="0" pageNumber="303">28</figureCitation>
): Some specimens with cell
<emphasis id="F4C3AB76B70DDAD6C51CFABB4F185D66" italics="true" pageId="0" pageNumber="303">br</emphasis>
and majority of discal cell surface hyaline.
</paragraph>
<paragraph id="8E5CAA5A634FA2485A6D99B292A7C560" pageId="0" pageNumber="303">
<emphasis id="7B60598BCDA5B59E7F745F15B17BF0C5" bold="true" italics="true" pageId="0" pageNumber="303">Abdomen</emphasis>
: Darker overall colour, similar anterodorsal margin and setation colour, setulae shorter overall.
</paragraph>
<paragraph id="CD4E4C9D100DA072633BBF82AFB57C2C" pageId="0" pageNumber="303">
<emphasis id="9057F38996E2B09654668EA4235A4319" bold="true" italics="true" pageId="0" pageNumber="303">Terminalia</emphasis>
: Cercus dark brown with dark setulae; genital fork (Fig.
<figureCitation id="E8FE9D6739E07C67A4B1222A27DB9F3E" captionStart="Figures 3739" captionStartId="F7" captionText="Figures 37 - 39. Atrichops spp., ♀ genital fork and spermatheca 37 A. adamastor (BMSA (D) 38899 38 A. intermedius sp. nov. paratype (NMSA-Dip. 212861) 39 A. stuckenbergi (BMSA (D) 95855). Abbreviations: gen fk - genital fork; spmth - spermatheca. Scale bars: 0.15 mm." figureDoi="10.3897/afrinvertebr.64.113133.figures37-39" httpUri="https://binary.pensoft.net/fig/950655" pageId="0" pageNumber="303">38</figureCitation>
) with distal apodeme slender, median lobe with shallow apical emargination, paired apical lobes with angular appearance, arms each having angular projection; 3 sclerotised, elongated spermathecae.
</paragraph>
<caption id="C3C92004BF823738123431320936C3B1" doi="10.3897/afrinvertebr.64.113133.figures37-39" httpUri="https://binary.pensoft.net/fig/950655" pageId="0" pageNumber="303" start="Figures 3739" startId="F7">
<paragraph id="C52A6B02BB846E51329386630A3D1300" pageId="0" pageNumber="303">
<emphasis id="ADBFDDEA6EDE25F65697667A207963B0" bold="true" pageId="0" pageNumber="303">Figures 37-39.</emphasis>
<taxonomicName id="19719BEBAA0ACB0109EC16CA95DE5358" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="F7FD5CC4281FEA23839B9C3D64FCBDF7" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
</taxonomicName>
spp., ♀ genital fork and spermatheca
<emphasis id="33A749371E1934D50BB78724BF0A68F0" bold="true" pageId="0" pageNumber="303">37</emphasis>
<taxonomicName id="CBB6874A87D818926C64CB8103A51E97" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="A0F1066B4DC090320B1016E0ED38EB66" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
(BMSA(D)38899
<emphasis id="1AA9BE5E2874630F02714A81B24DEB03" bold="true" pageId="0" pageNumber="303">38</emphasis>
<taxonomicName id="D8DBD6151F8F21FEADA6D31F5A58497D" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="AF1BD7CF206EBFF70E2631E432C84595" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
</taxonomicName>
sp. nov. paratype (NMSA-Dip. 212861)
<emphasis id="0A6E9BF9DF43B5683E272A13D1EEB11A" bold="true" pageId="0" pageNumber="303">39</emphasis>
<taxonomicName id="A8E218AF3CDA51047CBA921915477BF3" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="92648E56A500E765F9F0C9E7C3349D3A" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
(BMSA(D) 95855). Abbreviations: gen fk - genital fork; spmth - spermatheca. Scale bars: 0.15 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F383378C5E3DEFA0862A69EE480F8C2F" pageId="0" pageNumber="303" type="etymology">
<paragraph id="C20711FB156D112F2CAF68146213ADEE" pageId="0" pageNumber="303">Etymology.</paragraph>
<paragraph id="91306D29E267138D75F5689617E324DA" pageId="0" pageNumber="303">
From the Latin &quot;
<emphasis id="660709B06C339CF747F536576AD9C2AE" italics="true" pageId="0" pageNumber="303">intermedius</emphasis>
&quot;, meaning in the middle, referring to the species known occurrence being roughly between the distributions (Fig.
<figureCitation id="5D2A5FB1CC32A378F2B36121F38FCD34" captionStart="Figure 40" captionStartId="F8" captionText="Figure 40. Distribution of Atrichops spp. in the Afrotropical Region. Map generated using Africa Terrestrial Ecosystems data (Sayre 2023)." figureDoi="10.3897/afrinvertebr.64.113133.figure40" httpUri="https://binary.pensoft.net/fig/950656" pageId="0" pageNumber="303">40</figureCitation>
) of
<taxonomicName id="8347C6991DD82C0D30930D0FDF7FA253" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. adamastor" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="adamastor">
<emphasis id="D1F0640230321A9BF923B912BD9D52FD" italics="true" pageId="0" pageNumber="303">A. adamastor</emphasis>
</taxonomicName>
and
<taxonomicName id="F0CD62FC0D15EFE00EC230E48D0ACA2C" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. stuckenbergi" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="stuckenbergi">
<emphasis id="0701C977F75C4B65588E8DDA8775C5AD" italics="true" pageId="0" pageNumber="303">A. stuckenbergi</emphasis>
</taxonomicName>
, but also in reference to
<taxonomicName id="1961F953FA689F7BC954786B8347429B" class="Hexapoda" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="A. intermedius" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="1899D40E86EE360951D8173821487A26" italics="true" pageId="0" pageNumber="303">A. intermedius</emphasis>
</taxonomicName>
exhibiting a mixture of characteristics found in both the other two Afrotropical species.
</paragraph>
<caption id="34EB9C7E5BAA5FF4F4D71447301FAD5E" doi="10.3897/afrinvertebr.64.113133.figure40" httpUri="https://binary.pensoft.net/fig/950656" pageId="0" pageNumber="303" start="Figure 40" startId="F8">
<paragraph id="4CB5AB0BFCE4FF45019083AED7AF9831" pageId="0" pageNumber="303">
<emphasis id="E7AF7921B3D4C3E2D63F338C86E6E92F" bold="true" pageId="0" pageNumber="303">Figure 40.</emphasis>
Distribution of
<taxonomicName id="52857ADC2660D4F544259628F7F5BE36" authorityName="Verrall" authorityYear="1909" class="Insecta" family="Athericidae" genus="Atrichops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Atrichops" order="Diptera" pageId="0" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="CE16E633FF7283DA8B03DA3946E22711" italics="true" pageId="0" pageNumber="303">Atrichops</emphasis>
</taxonomicName>
spp. in the Afrotropical Region. Map generated using Africa Terrestrial Ecosystems data (
<bibRefCitation id="1C7B4B8F9C577AE456E90B1466B83EF4" DOI="https://doi.org/10.5066/P9BHBKA2" author="Sayre, R" journalOrPublisher="Struik, Cape Town" pageId="0" pageNumber="303" refId="B12" refString="Sayre, R, 2023. Africa Terrestrial Ecosystems: U.S. Geological Survey data release. https://doi.org/10.5066/P9BHBKA2" title="Africa Terrestrial Ecosystems: U. S. Geological Survey data release." url="https://doi.org/10.5066/P9BHBKA2" year="2023">Sayre 2023</bibRefCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0F890C34F16A0598DA0E3877B6CD85FD" pageId="0" pageNumber="303" type="distribution">
<paragraph id="720515A009D17DFE05217DAC92E26D3F" pageId="0" pageNumber="303">Distribution.</paragraph>
<paragraph id="EA0BD0141A957EC44D810BAB882EDB06" pageId="0" pageNumber="303">South Africa (Mpumalanga).</paragraph>
</subSubSection>
</treatment>
</document>