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<document id="312E7CEE0C9930CCC1BE1872E2DFD53D" ID-CLB-Dataset="32205" ID-DOI="10.11646/zootaxa.4317.3.3" ID-GBIF-Dataset="5d5d90e2-75db-46bd-b979-7c2dfaab10dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="884738" ID-ZooBank="0D977C9B-2B9C-46E7-B154-E795B7Ff5B01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504596256472" checkinUser="plazi" docAuthor="Sureshan, Pavittu M." docDate="2017" docId="9518B56BC4418113FF52FB44BD89FD6E" docLanguage="en" docName="zootaxa.4317.3.3.pdf" docOrigin="Zootaxa 4317 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eumenes Latreille in de Saussure 1852" docType="key" docVersion="13" lastPageNumber="471" masterDocId="6921CD13C4408111FFC5FFFCB815FF99" masterDocTitle="A taxonomic review of the Indian species of the genus Eumenes Latreille (Hymenoptera: Vespidae: Eumeninae)" masterLastPageNumber="498" masterPageNumber="469" pageNumber="470" updateTime="1698467852617" updateUser="ExternalLinkService">
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<mods:title id="72E8051E5B4AACEF6F6B9F5BED8BAAF3">A taxonomic review of the Indian species of the genus Eumenes Latreille (Hymenoptera: Vespidae: Eumeninae)</mods:title>
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<treatment id="9518B56BC4418113FF52FB44BD89FD6E" ID-DOI="http://doi.org/10.5281/zenodo.4489815" ID-GBIF-Taxon="133183400" ID-Zenodo-Dep="4489815" LSID="urn:lsid:plazi:treatment:9518B56BC4418113FF52FB44BD89FD6E" httpUri="http://treatment.plazi.org/id/9518B56BC4418113FF52FB44BD89FD6E" lastPageId="2" lastPageNumber="471" pageId="1" pageNumber="470">
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Key to the Indian species of
<taxonomicName id="DAB17FFEC4418110FE33FB4BBACCFB4B" ID-CoL="4FY8" ID-ENA="76987" authority="Latreille" authorityName="Latreille in de Saussure" authorityYear="1852" box="[502,729,1207,1234]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="genus">
<emphasis id="2FC5D86FC4418110FE33FB4BBA73FB48" bold="true" box="[502,614,1207,1233]" italics="true" pageId="1" pageNumber="470">Eumenes</emphasis>
Latreille
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<keyStep id="A6451CD8C4418110FF52FB02BD8EFAF2" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52FB02BD8EFAA8" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52FB02BD8EFAA8" pageId="1" pageNumber="470">
1. T1 very long (4.5-5.0 mm), lateral margins of postpetiolus substraight and parallel (
<figureCitation id="858A18F8C4418110FBD3FB02BC9DFA8A" box="[1046,1160,1278,1299]" captionStart="FIGURES 25 30" captionStartId="9.[151,264,1959,1982]" captionTargetBox="[151,1436,444,1898]" captionTargetId="figure@9.[151,1436,444,1898]" captionTargetPageId="9" captionText="FIGURES 25 30. Eumenes belli Giordani Soika, ♀. 25, Body, lateral view. 26, Head, frontal view. 27, Antenna. 28, Head and mesosoma, dorsal view. 29, Petiole, dorsal view. 30, Metasoma except petiole, dorsal view." httpUri="https://zenodo.org/record/884748/files/figure.png" pageId="1" pageNumber="470">Figs 25, 29</figureCitation>
,
<figureCitation id="858A18F8C4418110FB51FB02BCBAFA8A" box="[1172,1199,1278,1299]" captionStart="FIGURES 73 78" captionStartId="23.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@23.[151,1436,193,1647]" captionTargetPageId="23" captionText="FIGURES 73 78. Eumenes sikkimensis Giordani Soika, ♀. 73. Body, lateral view. 74, Head, frontal view. 75, Antenna. 76, Head and mesosoma, dorsal view. 77, Petiole, dorsal view. 78, T 2, dorsal view." httpUri="https://zenodo.org/record/884764/files/figure.png" pageId="1" pageNumber="470">77</figureCitation>
); head much wider than long (
<figureCitation id="858A18F8C4418110FEDDFAE0B976FAA8" box="[280,355,1308,1329]" captionStart="FIGURES 25 30" captionStartId="9.[151,264,1959,1982]" captionTargetBox="[151,1436,444,1898]" captionTargetId="figure@9.[151,1436,444,1898]" captionTargetPageId="9" captionText="FIGURES 25 30. Eumenes belli Giordani Soika, ♀. 25, Body, lateral view. 26, Head, frontal view. 27, Antenna. 28, Head and mesosoma, dorsal view. 29, Petiole, dorsal view. 30, Metasoma except petiole, dorsal view." httpUri="https://zenodo.org/record/884748/files/figure.png" pageId="1" pageNumber="470">Figs 26</figureCitation>
,
<figureCitation id="858A18F8C4418110FEA9FAE0B992FAA8" box="[364,391,1308,1329]" captionStart="FIGURES 73 78" captionStartId="23.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@23.[151,1436,193,1647]" captionTargetPageId="23" captionText="FIGURES 73 78. Eumenes sikkimensis Giordani Soika, ♀. 73. Body, lateral view. 74, Head, frontal view. 75, Antenna. 76, Head and mesosoma, dorsal view. 77, Petiole, dorsal view. 78, T 2, dorsal view." httpUri="https://zenodo.org/record/884764/files/figure.png" pageId="1" pageNumber="470">74</figureCitation>
); temple without hairs.................................................................... 2
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4418110FF52FAC5BD8EFAF2" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52FAC5BD8EFAF2" pageId="1" pageNumber="470">
- T1 of normal length (3.0-3.5 mm), pyriform (pear-shaped) (
<figureCitation id="858A18F8C4418110FCE2FAC5BB72FAD7" box="[807,871,1337,1358]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1935,1958]" captionTargetBox="[151,1436,408,1862]" captionTargetId="figure@3.[151,1436,408,1862]" captionTargetPageId="3" captionText="FIGURES 1 6. Eumenes affinissimus de Saussure. 1 5 ♀. 1, Body, lateral view. 2, Head, frontal view. 3, Head and mesosoma, dorsal view. 4, Petiole, dorsal view. 5, T 2, dorsal view. 6, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884740/files/figure.png" pageId="1" pageNumber="470">Figs 4</figureCitation>
, 12, 20, 35, 40, 48, 56, 64); head slightly wider than long (
<figureCitation id="858A18F8C4418110FF22FAAAB933FAF2" box="[231,294,1366,1387]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1935,1958]" captionTargetBox="[151,1436,408,1862]" captionTargetId="figure@3.[151,1436,408,1862]" captionTargetPageId="3" captionText="FIGURES 1 6. Eumenes affinissimus de Saussure. 1 5 ♀. 1, Body, lateral view. 2, Head, frontal view. 3, Head and mesosoma, dorsal view. 4, Petiole, dorsal view. 5, T 2, dorsal view. 6, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884740/files/figure.png" pageId="1" pageNumber="470">Figs 2</figureCitation>
, 10, 18, 38, 46, 54, 62); temple with hairs............................................................ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4418110FF52FA88BD8EFA78" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52FA88BD8EFA3F" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52FA88BD8EFA3F" pageId="1" pageNumber="470">
2. T1, in profile view, distinctly swollen in apical half; antero-lateral angles of pronotum more prominent; scape of male antennae very broad...........................................................................
<taxonomicName id="DAB17FFEC4418110FB11FA6DBD8EFA3F" ID-CoL="3CDD7" authority="Bingham" authorityName="Bingham" authorityYear="1898" box="[1236,1435,1425,1446]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="2FC5D86FC4418110FB11FA6DBD28FA3F" box="[1236,1341,1425,1446]" italics="true" pageId="1" pageNumber="470">antennatus</emphasis>
Bingham
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4418110FF52FA52BD8EFA78" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52FA52BD8EFA78" pageId="1" pageNumber="470">
- T1, in profile view, not swollen in apical half (
<figureCitation id="858A18F8C4418110FD6CFA52BAE0FA5A" box="[681,757,1454,1475]" captionStart="FIGURES 25 30" captionStartId="9.[151,264,1959,1982]" captionTargetBox="[151,1436,444,1898]" captionTargetId="figure@9.[151,1436,444,1898]" captionTargetPageId="9" captionText="FIGURES 25 30. Eumenes belli Giordani Soika, ♀. 25, Body, lateral view. 26, Head, frontal view. 27, Antenna. 28, Head and mesosoma, dorsal view. 29, Petiole, dorsal view. 30, Metasoma except petiole, dorsal view." httpUri="https://zenodo.org/record/884748/files/figure.png" pageId="1" pageNumber="470">Figs 25</figureCitation>
,
<figureCitation id="858A18F8C4418110FCC5FA52BB0EFA5A" box="[768,795,1454,1475]" captionStart="FIGURES 73 78" captionStartId="23.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@23.[151,1436,193,1647]" captionTargetPageId="23" captionText="FIGURES 73 78. Eumenes sikkimensis Giordani Soika, ♀. 73. Body, lateral view. 74, Head, frontal view. 75, Antenna. 76, Head and mesosoma, dorsal view. 77, Petiole, dorsal view. 78, T 2, dorsal view." httpUri="https://zenodo.org/record/884764/files/figure.png" pageId="1" pageNumber="470">73</figureCitation>
); antero-lateral angles of pronotum less prominent; scape of male antennae not very broad, normal.......................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4418110FF52FA15BD89F9EA" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52FA15BD89F9A0" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52FA15BD89F9A0" pageId="1" pageNumber="470">
3. Clypeus yellow; scape ventrally yellow; pronotum mostly yellow, black near tegulae; scutellum with yellow marks; ventral side of petiole with two long yellow stripes which occupy the entire apex and merge with apical band on dorsal side; S2 yellow except at base............................................................................
<taxonomicName id="DAB17FFEC4418110FACEF9D8BD89F9A0" ID-CoL="6GX29" authority="Dover" authorityName="Dover" authorityYear="1925" box="[1291,1436,1572,1593]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="comberi">
<emphasis id="2FC5D86FC4418110FACEF9D8BD4FF9A0" box="[1291,1370,1572,1593]" italics="true" pageId="1" pageNumber="470">comberi</emphasis>
Dover
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4418110FF52F9BDBD89F9EA" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F9BDBD89F9EA" pageId="1" pageNumber="470">- Clypeus black; scape entirely black; pronotum mostly black except narrow yellow transverse band on anterior and posterior apex of dorsal face; scutellum without yellow marks; ventral side of petiole without yellow stripes; S2 entirely black...... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4418110FF52F980BD8EF8BA" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52F980BD8EF952" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F980BD8EF952" pageId="1" pageNumber="470">
4. Punctures on T2 at apex, before apical lamella, very thick and dense (
<figureCitation id="858A18F8C4418110FC51F980BBCBF908" box="[916,990,1660,1681]" captionStart="FIGURES 25 30" captionStartId="9.[151,264,1959,1982]" captionTargetBox="[151,1436,444,1898]" captionTargetId="figure@9.[151,1436,444,1898]" captionTargetPageId="9" captionText="FIGURES 25 30. Eumenes belli Giordani Soika, ♀. 25, Body, lateral view. 26, Head, frontal view. 27, Antenna. 28, Head and mesosoma, dorsal view. 29, Petiole, dorsal view. 30, Metasoma except petiole, dorsal view." httpUri="https://zenodo.org/record/884748/files/figure.png" pageId="1" pageNumber="470">Fig. 30</figureCitation>
); apex of clypeus narrower and more weakly emarginate, with rounded apical teeth (
<figureCitation id="858A18F8C4418110FD92F965BA8AF937" box="[599,671,1689,1710]" captionStart="FIGURES 25 30" captionStartId="9.[151,264,1959,1982]" captionTargetBox="[151,1436,444,1898]" captionTargetId="figure@9.[151,1436,444,1898]" captionTargetPageId="9" captionText="FIGURES 25 30. Eumenes belli Giordani Soika, ♀. 25, Body, lateral view. 26, Head, frontal view. 27, Antenna. 28, Head and mesosoma, dorsal view. 29, Petiole, dorsal view. 30, Metasoma except petiole, dorsal view." httpUri="https://zenodo.org/record/884748/files/figure.png" pageId="1" pageNumber="470">Fig. 26</figureCitation>
); mesepisternum, tegula, metanotum and propodeum with yellow markings........................................................................................
<taxonomicName id="DAB17FFEC4418110FB10F94ABD8EF952" ID-CoL="3CDDX" authority="Giordani Soika" authorityName="Giordani Soika" authorityYear="1973" box="[1237,1435,1718,1739]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="belli">
<emphasis id="2FC5D86FC4418110FB10F94ABD11F952" box="[1237,1284,1718,1739]" italics="true" pageId="1" pageNumber="470">belli</emphasis>
Giordani Soika
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4418110FF52F928BD8EF8BA" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F928BD8EF8BA" pageId="1" pageNumber="470">
- Punctures on T2 at apex, before apical lamella, sparse, integument almost smooth in middle (
<figureCitation id="858A18F8C4418110FB98F928BCB0F970" box="[1117,1189,1748,1769]" captionStart="FIGURES 73 78" captionStartId="23.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@23.[151,1436,193,1647]" captionTargetPageId="23" captionText="FIGURES 73 78. Eumenes sikkimensis Giordani Soika, ♀. 73. Body, lateral view. 74, Head, frontal view. 75, Antenna. 76, Head and mesosoma, dorsal view. 77, Petiole, dorsal view. 78, T 2, dorsal view." httpUri="https://zenodo.org/record/884764/files/figure.png" pageId="1" pageNumber="470">Fig. 78</figureCitation>
); apex of clypeus broader and less weakly emarginate, with sharper apical teeth (
<figureCitation id="858A18F8C4418110FD3EF90DBB50F89F" box="[763,837,1777,1798]" captionStart="FIGURES 73 78" captionStartId="23.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@23.[151,1436,193,1647]" captionTargetPageId="23" captionText="FIGURES 73 78. Eumenes sikkimensis Giordani Soika, ♀. 73. Body, lateral view. 74, Head, frontal view. 75, Antenna. 76, Head and mesosoma, dorsal view. 77, Petiole, dorsal view. 78, T 2, dorsal view." httpUri="https://zenodo.org/record/884764/files/figure.png" pageId="1" pageNumber="470">Fig. 74</figureCitation>
); mesepisternum, tegula, metanotum and propodeum without yellow markings.................................................................
<taxonomicName id="DAB17FFEC4418110FB52F8F2BD8EF8BA" ID-CoL="3CDP5" authority="Giordani Soika" authorityName="Giordani Soika" authorityYear="1986" box="[1175,1435,1806,1827]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="sikkimensis">
<emphasis id="2FC5D86FC4418110FB52F8F2BD11F8BA" box="[1175,1284,1806,1827]" italics="true" pageId="1" pageNumber="470">sikkimensis</emphasis>
Giordani Soika
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4418110FF52F8D0BD8EF8C7" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52F8D0BD8EF8D8" blockId="1.[151,1436,1278,2033]" box="[151,1435,1836,1857]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F8D0BD8EF8D8" box="[151,1435,1836,1857]" pageId="1" pageNumber="470">
5. T2 with long erect hairs, at least at base, about as long as those of T1 (
<figureCitation id="858A18F8C4418110FC47F8D0BBCCF8D8" box="[898,985,1836,1857]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1935,1958]" captionTargetBox="[151,1436,408,1862]" captionTargetId="figure@3.[151,1436,408,1862]" captionTargetPageId="3" captionText="FIGURES 1 6. Eumenes affinissimus de Saussure. 1 5 ♀. 1, Body, lateral view. 2, Head, frontal view. 3, Head and mesosoma, dorsal view. 4, Petiole, dorsal view. 5, T 2, dorsal view. 6, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884740/files/figure.png" pageId="1" pageNumber="470">Figs 1, 6</figureCitation>
, 45, 50).............................. 6
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4418110FF52F8B5BD8EF8C7" blockId="1.[151,1436,1278,2033]" box="[151,1435,1865,1886]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F8B5BD8EF8C7" box="[151,1435,1865,1886]" pageId="1" pageNumber="470">- T2 with hairs very short, much shorter than those of T1 (Figs 9, 14, 18, 31, 37, 53, 61, 71)............................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4418110FF52F89ABD8EF84A" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52F89ABD8EF800" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F89ABD8EF800" pageId="1" pageNumber="470">
6. S2 with many whitish hairs, about as long as those of T2 (
<figureCitation id="858A18F8C4418110FCD7F89ABB7CF8E2" box="[786,873,1894,1915]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1935,1958]" captionTargetBox="[151,1436,408,1862]" captionTargetId="figure@3.[151,1436,408,1862]" captionTargetPageId="3" captionText="FIGURES 1 6. Eumenes affinissimus de Saussure. 1 5 ♀. 1, Body, lateral view. 2, Head, frontal view. 3, Head and mesosoma, dorsal view. 4, Petiole, dorsal view. 5, T 2, dorsal view. 6, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884740/files/figure.png" pageId="1" pageNumber="470">Figs 1, 6</figureCitation>
); T2 with long glossy hairs (
<figureCitation id="858A18F8C4418110FB45F89ABCAFF8E2" box="[1152,1210,1894,1915]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1935,1958]" captionTargetBox="[151,1436,408,1862]" captionTargetId="figure@3.[151,1436,408,1862]" captionTargetPageId="3" captionText="FIGURES 1 6. Eumenes affinissimus de Saussure. 1 5 ♀. 1, Body, lateral view. 2, Head, frontal view. 3, Head and mesosoma, dorsal view. 4, Petiole, dorsal view. 5, T 2, dorsal view. 6, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884740/files/figure.png" pageId="1" pageNumber="470">Fig. 5</figureCitation>
); punctures on T2 very small and sparse, interspaces much larger than diameter of punctures (
<figureCitation id="858A18F8C4418110FC44F878BBAFF800" box="[897,954,1924,1945]" captionStart="FIGURES 1 6" captionStartId="3.[151,264,1935,1958]" captionTargetBox="[151,1436,408,1862]" captionTargetId="figure@3.[151,1436,408,1862]" captionTargetPageId="3" captionText="FIGURES 1 6. Eumenes affinissimus de Saussure. 1 5 ♀. 1, Body, lateral view. 2, Head, frontal view. 3, Head and mesosoma, dorsal view. 4, Petiole, dorsal view. 5, T 2, dorsal view. 6, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884740/files/figure.png" pageId="1" pageNumber="470">Fig. 5</figureCitation>
)...................
<taxonomicName id="DAB17FFEC4418110FB74F878BD8EF800" ID-CoL="3CDCV" authority="de Saussure" authorityName="de Saussure" box="[1201,1435,1924,1945]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="affinissimus">
<emphasis id="2FC5D86FC4418110FB74F878BD31F800" box="[1201,1316,1924,1945]" italics="true" pageId="1" pageNumber="470">affinissimus</emphasis>
de Saussure
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4418110FF52F85DBD8EF84A" blockId="1.[151,1436,1278,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F85DBD8EF84A" pageId="1" pageNumber="470">
- S2 with very short hairs, erect long hairs absent (Fig. 45); T2 at base with medium length hairs, rest with much shorter hairs; punctures on T2 larger and denser (
<figureCitation id="858A18F8C4418110FDF1F842BA6FF84A" box="[564,634,1982,2003]" captionStart="FIGURES 49 52" captionStartId="16.[151,264,1632,1655]" captionTargetBox="[151,1436,157,1611]" captionTargetId="figure@16.[151,1436,157,1611]" captionTargetPageId="16" captionText="FIGURES 49 52. Eumenes persimilis Giordani Soika. 49 ♀, T 2, dorsal view. 50 52 ♂. 50, Body, lateral view. 51, Apical antennal articles. 52, Metasoma dorsal view. FIGURES 53 54. Eumenes placens Nurse, ♀. 53, Body, lateral view. 54, Head, frontal view." httpUri="https://zenodo.org/record/884756/files/figure.png" pageId="1" pageNumber="470">Fig. 49</figureCitation>
).............................................
<taxonomicName id="DAB17FFEC4418110FB61F842BD8EF84A" ID-CoL="3CDMK" authority="Giordani Soika" authorityName="Giordani Soika" box="[1188,1435,1982,2003]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="2FC5D86FC4418110FB61F842BD17F84A" box="[1188,1282,1982,2003]" italics="true" pageId="1" pageNumber="470">persimilis</emphasis>
Giordani Soika
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4418113FF52F820BD8EFF7E" lastPageId="2" lastPageNumber="471" pageId="1" pageNumber="470">
<paragraph id="1D0E047DC4418110FF52F820BD89F868" blockId="1.[151,1436,1278,2033]" box="[151,1436,2012,2033]" pageId="1" pageNumber="470">
<keyLead id="A640A748C4418110FF52F820BD89F868" box="[151,1436,2012,2033]" pageId="1" pageNumber="470">
7. Lateral margin of tegula with a distinct emargination in anterior third (
<figureCitation id="858A18F8C4418110FC4EF820BBC6F868" box="[907,979,2012,2033]" captionStart="FIGURES 55 60" captionStartId="20.[151,264,1920,1943]" captionTargetBox="[151,1436,445,1899]" captionTargetId="figure@20.[151,1436,445,1899]" captionTargetPageId="20" captionText="FIGURES 55 60. Eumenes placens Nurse. 55 57 ♀. 55. Head and mesosoma, dorsal view. 56, Petiole, dorsal view. 57, T 2, dorsal view. 58 60 ♂. 58, Body, lateral view. 59, Head, frontal view. 60, Clypeus, frontal view." httpUri="https://zenodo.org/record/884758/files/figure.png" pageId="1" pageNumber="470">Fig. 55</figureCitation>
); dorsal surface of pronotum almost flat; meta-
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF24FF6BBD8EFF7E" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
soma extensively ferruginous (Fig. 53)..........................................................
<taxonomicName id="DAB17FFEC4428113FAD7FF6BBD8EFF35" ID-CoL="3CDMR" authority="Nurse" authorityName="Nurse" box="[1298,1435,151,172]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="placens">
<emphasis id="2FC5D86FC4428113FAD7FF6BBD4EFF35" box="[1298,1371,151,172]" italics="true" pageId="2" pageNumber="471">placens</emphasis>
Nurse
</taxonomicName>
<keyLead id="A640A748C4428113FF52FF49BD8EFF7E" pageId="2" pageNumber="471">- Tegula normal, lateral margin without emargination in anterior third; dorsal surface of pronotum convex as usual; metasoma either little ferruginous or different........................................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4428113FF52FF13BD8EFEC5" pageId="2" pageNumber="471">
<paragraph id="1D0E047DC4428113FF52FF13BD8EFEBB" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FF13BD8EFEBB" pageId="2" pageNumber="471">
8. T2, in profile view, strongly gibbous in middle, strongly impressed preapically, its apical margin strongly reflected (Fig. 9); body black with dark red markings (with an orange-yellow spot on interantennal space)..........
<taxonomicName id="DAB17FFEC4428113FB67FEF1BD8EFEBB" ID-CoL="3CDDL" authority="Meade-Waldo" authorityName="Meade-Waldo" box="[1186,1435,269,290]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="assamensis">
<emphasis id="2FC5D86FC4428113FB67FEF1BD1BFEBB" box="[1186,1294,269,290]" italics="true" pageId="2" pageNumber="471">assamensis</emphasis>
Meade-Waldo
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF52FED6BD8EFEC5" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FED6BD8EFEC5" pageId="2" pageNumber="471">- T2, in profile view, not or little swollen in middle, weakly impressed preapically, its apical margin slightly reflected (Figs 18, 31, 37, 61, 71); body colour pattern different................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4428113FF52FE99BD8EFE4B" pageId="2" pageNumber="471">
<paragraph id="1D0E047DC4428113FF52FE99BD89FE0E" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FE99BD89FE0E" pageId="2" pageNumber="471">
9. Punctation of T2 fine and dense (
<figureCitation id="858A18F8C4428113FDE2FE99BA65FEE3" box="[551,624,357,378]" captionStart="FIGURES 31 36" captionStartId="12.[151,264,1949,1972]" captionTargetBox="[151,1436,451,1905]" captionTargetId="figure@12.[151,1436,451,1905]" captionTargetPageId="12" captionText="FIGURES 31 36. Eumenes kangrae Dover, ♂. 31, Body, lateral view. 32, Head, frontal view. 33, Clypeus. 34, Antenna. 35, Head, mesosoma and petiole, dorsal view. 36, T 2, dorsal view." httpUri="https://zenodo.org/record/884750/files/figure.png" pageId="2" pageNumber="471">Fig. 36</figureCitation>
); propodeum and T1 extensively ferruginous; clypeus of ♂ more elongate (
<figureCitation id="858A18F8C4428113FA8AFE99B8EFFE0E" captionStart="FIGURES 31 36" captionStartId="12.[151,264,1949,1972]" captionTargetBox="[151,1436,451,1905]" captionTargetId="figure@12.[151,1436,451,1905]" captionTargetPageId="12" captionText="FIGURES 31 36. Eumenes kangrae Dover, ♂. 31, Body, lateral view. 32, Head, frontal view. 33, Clypeus. 34, Antenna. 35, Head, mesosoma and petiole, dorsal view. 36, T 2, dorsal view." httpUri="https://zenodo.org/record/884750/files/figure.png" pageId="2" pageNumber="471">Fig. 32, 33</figureCitation>
)......................................................................................
<taxonomicName id="DAB17FFEC4428113FACEFE7EBD89FE0E" ID-CoL="3CDJZ" authority="Dover" authorityName="Dover" authorityYear="1925" box="[1291,1436,386,407]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="kangrae">
<emphasis id="2FC5D86FC4428113FACEFE7EBD4FFE0E" box="[1291,1370,386,407]" italics="true" pageId="2" pageNumber="471">kangrae</emphasis>
Dover
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF52FE63BD8EFE4B" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FE63BD8EFE4B" pageId="2" pageNumber="471">
- Punctation of T2 coarser and less dense (
<figureCitation id="858A18F8C4428113FDAAFE63BAAEFE2D" box="[623,699,415,436]" captionStart="FIGURES 19 24" captionStartId="8.[151,264,1670,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@8.[151,1436,195,1649]" captionTargetPageId="8" captionText="FIGURES 19 24. Eumenes atrophicus (Fabricius). 19 21 ♀. 19, Head and mesosoma, dorsal view. 20, Petiole, dorsal view. 21, T 2, dorsal view. 22 24 ♂. 22, Body, lateral view. 23, Head, frontal view. 24. Clypeus, frontal view." httpUri="https://zenodo.org/record/884746/files/figure.png" pageId="2" pageNumber="471">Figs 21</figureCitation>
,
<figureCitation id="858A18F8C4428113FD00FE63BACAFE2D" box="[709,735,415,436]" captionStart="FIGURES 37 42" captionStartId="13.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@13.[151,1436,193,1647]" captionTargetPageId="13" captionText="FIGURES 37 42. Eumenes macrops de Saussure. 37 41 ♀. 37, Body, lateral view. 38, Head, frontal view. 39, Head and mesosoma, dorsal view. 40, Petiole. 41, T 2, dorsal view. 42 ♂, Body, lateral view." httpUri="https://zenodo.org/record/884752/files/figure.png" pageId="2" pageNumber="471">41</figureCitation>
,
<figureCitation id="858A18F8C4428113FD2FFE63BB11FE2D" box="[746,772,415,436]" captionStart="FIGURES 61 66" captionStartId="21.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@21.[151,1436,193,1647]" captionTargetPageId="21" captionText="FIGURES 61 66. Eumenes pomiformis (Fabricius). 61 65 ♀. 61. Body, latera view. 62, Head, frontal view. 63, Head and mesosoma, dorsal view. 64, Petiole, dorsal view. 65, T 2, dorsal view. 66, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884760/files/figure.png" pageId="2" pageNumber="471">66</figureCitation>
, 72); propodeum and T1 not ferruginous; clypeus of ♂ shorter (
<figureCitation id="858A18F8C4428113FAB4FE63B8EEFE4B" captionStart="FIGURES 19 24" captionStartId="8.[151,264,1670,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@8.[151,1436,195,1649]" captionTargetPageId="8" captionText="FIGURES 19 24. Eumenes atrophicus (Fabricius). 19 21 ♀. 19, Head and mesosoma, dorsal view. 20, Petiole, dorsal view. 21, T 2, dorsal view. 22 24 ♂. 22, Body, lateral view. 23, Head, frontal view. 24. Clypeus, frontal view." httpUri="https://zenodo.org/record/884746/files/figure.png" pageId="2" pageNumber="471">Figs 24</figureCitation>
,
<figureCitation id="858A18F8C4428113FEC0FE41B90AFE4B" box="[261,287,445,466]" captionStart="FIGURES 43 44" captionStartId="15.[151,264,1929,1952]" captionTargetBox="[151,1436,387,1841]" captionTargetId="figure@15.[151,1436,387,1841]" captionTargetPageId="15" captionText="FIGURES 43 44. Eumenes macrops de Saussure, ♂. 43, Head, frontal view. 44, Apical antennal articles. FIGURES 45 48. Eumenes persimilis Giordani Soika, ♀. 45, Body, lateral view. 46, Head, frontal view. 47, Head and mesosoma, dorsal view. 48, Petiole." httpUri="https://zenodo.org/record/884754/files/figure.png" pageId="2" pageNumber="471">43</figureCitation>
,
<figureCitation id="858A18F8C4428113FEECFE41B956FE4B" box="[297,323,445,466]" captionStart="FIGURES 67 68" captionStartId="22.[151,264,1638,1661]" captionTargetBox="[157,1421,188,1611]" captionTargetId="figure@22.[157,1421,188,1618]" captionTargetPageId="22" captionText="FIGURES 67 68. Eumenes pomiformis (Fabricius), ♂. 67. Head, frontal view. 68. Apical antennal articles. FIGURES 69 72. Eumenes punctatus de Saussure, ♂. 69, Head, frontal view. 70, Head and mesosoma, dorsal view. 71, Metasoma, lateral view. 72, T 2, dorsal view." httpUri="https://zenodo.org/record/884762/files/figure.png" pageId="2" pageNumber="471">67</figureCitation>
, 69)....................................................................................... 10
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4428113FF52FE26BD8EFDB3" pageId="2" pageNumber="471">
<paragraph id="1D0E047DC4428113FF52FE26BD8EFD95" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FE26BD8EFD95" pageId="2" pageNumber="471">
10. Legs almost entirely ferruginous (except coxae and trochanter, black), rarely with few yellow markings on fore and mid leg......................................................................................
<taxonomicName id="DAB17FFEC4428113FB17FE0BBD8EFD95" ID-CoL="6GX28" authority="de Saussure" authorityName="de Saussure" box="[1234,1435,503,524]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="macrops">
<emphasis id="2FC5D86FC4428113FB17FE0BBD31FD95" box="[1234,1316,503,524]" italics="true" pageId="2" pageNumber="471">macrops</emphasis>
de Saussure
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF52FDE9BD8EFDB3" blockId="2.[151,1436,151,759]" box="[151,1435,533,554]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FDE9BD8EFDB3" box="[151,1435,533,554]" pageId="2" pageNumber="471">- Legs black with yellow marks........................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4428113FF52FDCEBD8EFDFD" pageId="2" pageNumber="471">
<paragraph id="1D0E047DC4428113FF52FDCEBD8EFDDE" blockId="2.[151,1436,151,759]" box="[151,1435,562,583]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FDCEBD8EFDDE" box="[151,1435,562,583]" pageId="2" pageNumber="471">
11. Apex of T1 with a wide yellow band, and a pair of brownish marks in the yellow band (
<figureCitation id="858A18F8C4428113FBEFFDCEBC65FDDE" box="[1066,1136,562,583]" captionStart="FIGURES 61 66" captionStartId="21.[151,264,1668,1691]" captionTargetBox="[151,1436,193,1647]" captionTargetId="figure@21.[151,1436,193,1647]" captionTargetPageId="21" captionText="FIGURES 61 66. Eumenes pomiformis (Fabricius). 61 65 ♀. 61. Body, latera view. 62, Head, frontal view. 63, Head and mesosoma, dorsal view. 64, Petiole, dorsal view. 65, T 2, dorsal view. 66, ♂, Body, lateral view." httpUri="https://zenodo.org/record/884760/files/figure.png" pageId="2" pageNumber="471">Fig. 64</figureCitation>
).....
<taxonomicName id="DAB17FFEC4428113FB7AFDCEBD8EFDDE" ID-CoL="6GWPM" authority="(Fabricius)" authorityName="" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[1215,1435,562,583]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="pomiformis">
<emphasis id="2FC5D86FC4428113FB7AFDCEBD39FDDE" box="[1215,1324,562,583]" italics="true" pageId="2" pageNumber="471">pomiformis</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF52FDB3BD8EFDFD" blockId="2.[151,1436,151,759]" box="[151,1435,591,612]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FDB3BD8EFDFD" box="[151,1435,591,612]" pageId="2" pageNumber="471">
- Apex of T1 just with a narrow yellow band (
<figureCitation id="858A18F8C4428113FD4EFDB3BAC3FDFD" box="[651,726,591,612]" captionStart="FIGURES 19 24" captionStartId="8.[151,264,1670,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@8.[151,1436,195,1649]" captionTargetPageId="8" captionText="FIGURES 19 24. Eumenes atrophicus (Fabricius). 19 21 ♀. 19, Head and mesosoma, dorsal view. 20, Petiole, dorsal view. 21, T 2, dorsal view. 22 24 ♂. 22, Body, lateral view. 23, Head, frontal view. 24. Clypeus, frontal view." httpUri="https://zenodo.org/record/884746/files/figure.png" pageId="2" pageNumber="471">Figs 20</figureCitation>
, 72)..................................................... 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4428113FF52FD91BD8EFD25" pageId="2" pageNumber="471">
<paragraph id="1D0E047DC4428113FF52FD91BD80FD06" blockId="2.[151,1436,151,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FD91BD80FD06" pageId="2" pageNumber="471">
12. Three well separated narrow yellow transverse marks on pronotum (
<figureCitation id="858A18F8C4428113FCAAFD91BBA3FD1B" box="[879,950,621,642]" captionStart="FIGURES 19 24" captionStartId="8.[151,264,1670,1693]" captionTargetBox="[151,1436,195,1649]" captionTargetId="figure@8.[151,1436,195,1649]" captionTargetPageId="8" captionText="FIGURES 19 24. Eumenes atrophicus (Fabricius). 19 21 ♀. 19, Head and mesosoma, dorsal view. 20, Petiole, dorsal view. 21, T 2, dorsal view. 22 24 ♂. 22, Body, lateral view. 23, Head, frontal view. 24. Clypeus, frontal view." httpUri="https://zenodo.org/record/884746/files/figure.png" pageId="2" pageNumber="471">Fig. 19</figureCitation>
); scape entirely black (Fig. 18)..................................................................................................
<taxonomicName id="DAB17FFEC4428113FB04FD76BD80FD06" ID-CoL="3CDDN" authority="(Fabricius)" authorityName="" baseAuthorityName="Fabricius" baseAuthorityYear="1798" box="[1217,1429,650,671]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="atrophicus">
<emphasis id="2FC5D86FC4428113FB04FD76BD33FD06" box="[1217,1318,650,671]" italics="true" pageId="2" pageNumber="471">atrophicus</emphasis>
(Fabricius)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF52FD5BBD8EFD25" blockId="2.[151,1436,151,759]" box="[151,1435,679,700]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FD5BBD8EFD25" box="[151,1435,679,700]" pageId="2" pageNumber="471">- Uninterrupted broad yellow transverse band on pronotum (Fig. 70); ventral side of scape yellow (Fig. 69).............. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="A6451CD8C4428113FF52FD39BD89FD6E" pageId="2" pageNumber="471">
<paragraph id="1D0E047DC4428113FF52FD39BD80FD43" blockId="2.[151,1436,151,759]" box="[151,1429,709,730]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FD39BD80FD43" box="[151,1429,709,730]" pageId="2" pageNumber="471">
13. Apical margins of T3 &amp; T4 with yellow marks.............................................
<taxonomicName id="DAB17FFEC4428113FB04FD39BD80FD43" ID-CoL="3CDN2" authority="de Saussure" authorityName="de Saussure" authorityYear="1852" box="[1217,1429,709,730]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="2FC5D86FC4428113FB04FD39BD0BFD43" box="[1217,1310,709,730]" italics="true" pageId="2" pageNumber="471">punctatus</emphasis>
de Saussure
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D0E047DC4428113FF52FD1EBD89FD6E" blockId="2.[151,1436,151,759]" box="[151,1436,738,759]" pageId="2" pageNumber="471">
<keyLead id="A640A748C4428113FF52FD1EBD89FD6E" box="[151,1436,738,759]" pageId="2" pageNumber="471">
- Apical margins of T3 &amp; T4 without yellow marks...............................................
<taxonomicName id="DAB17FFEC4428113FB33FD1EBD89FD6E" ID-CoL="6H925" authority="Cameron" authorityName="Cameron" authorityYear="1898" box="[1270,1436,738,759]" class="Insecta" family="Vespidae" genus="Eumenes" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="buddha">
<emphasis id="2FC5D86FC4428113FB33FD1EBD2BFD6E" box="[1270,1342,738,759]" italics="true" pageId="2" pageNumber="471">buddha</emphasis>
Cameron
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>