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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.231.100894" ID-Pensoft-Pub="1314-2003-231-1" ID-Pensoft-UUID="0D762CF99C605B04AF072C867EEB1488" ModsDocID="1314-2003-231-1" checkinTime="1693383686345" checkinUser="pensoft" docAuthor="Knapp, Sandra, Saerkinen, Tiina &amp; Barboza, Gloria E." docDate="2023" docId="7CAF2565EC1F9EC7AA64562765242657" docLanguage="en" docName="PhytoKeys 231: 1-342" docOrigin="PhytoKeys 231" docPubDate="2023-08-29" docSource="http://dx.doi.org/10.3897/phytokeys.231.100894" docTitle="Solanum pallidum Rusby, Mem. Torrey Bot. Club 4: 228. 1895." docType="treatment" docVersion="1" id="0D762CF99C605B04AF072C867EEB1488" lastPageNumber="1" masterDocId="0D762CF99C605B04AF072C867EEB1488" masterDocTitle="A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)" masterLastPageNumber="342" masterPageNumber="1" pageNumber="1" updateTime="1693383686345" updateUser="pensoft">
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<mods:title id="7D6A827ACA9E90248718F6F1D26369F1">A revision of the South American species of the Morelloid clade (Solanum L., Solanaceae)</mods:title>
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<mods:namePart id="2F981407644A37F647E7A18A24629BA2">Knapp, Sandra</mods:namePart>
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<mods:affiliation id="DF74C4B1370641BB26B900DEB6B6C07D">Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="1506E783ECC28341A2BFED6C13A78CAD">Saerkinen, Tiina</mods:namePart>
<mods:nameIdentifier id="63F9ACE8D605EF7EEE7B79E80DD1FEF9" type="ORCID">https://orcid.org/0000-0002-6956-3093</mods:nameIdentifier>
<mods:affiliation id="0001843A94F39DA3C6C0EB37C9BB70D3">Royal Botanic Garden Edinburgh, 20 A Inverleith Row, Edinburgh EH 3 5 LR, UK</mods:affiliation>
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<mods:namePart id="A8C3F1E57F92B57DCBE9F614608C6750">Barboza, Gloria E.</mods:namePart>
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<mods:affiliation id="1735418DEDE580CB66BC6170627447D9">Instituto Multidisciplinario de Biologia Vegetal (CONICET-Universidad Nacional de Cordoba), Casilla de Correo 495, 5000 Cordoba, Argentina</mods:affiliation>
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<subSubSection id="BEE3DBEF361541BB68749A60D786FC5A" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="69A44C5638005DF9F96213C7C26EF155" pageId="0" pageNumber="1">
36.
<taxonomicName id="0000C0DD0D0DA6AA08AE95931266E28E" ID-CoL="4Y3ML" ID-ENA="1442336" authority="Rusby, Mem. Torrey Bot. Club 4: 228. 1895." authorityName="Rusby, Mem. Torrey Bot. Club 4: 228. 1895." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">Solanum pallidum Rusby, Mem. Torrey Bot. Club 4: 228. 1895.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="372D2BF77CBD471D818FE3CECA14B4D6" pageId="0" pageNumber="1" type="description">
<paragraph id="C7D3761AE8596401FB3CAC6D91BC498E" pageId="0" pageNumber="1">
<figureCitation id="8B97F691DA91B3B9D5AB7ADE47CB0340" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Representative habits and leaves of South American morelloids A prostrate annual herbs (S. weddellii) B large lax shrubs (S. aloysiifolium) C roots forming along a creeping stem (S. juninense) D spinose processes on stems of many species (S. huayavillense) E glandular trichomes found in some members of the clade (S. glandulosipilosum) F dendritic / branched trichomes found in S. pallidum G regularly 3 - lobed leaves in S. palitans H highly variable leaves along a single stem in S. salicifolium (A Saerkinen et al. 4038 B Barboza et al. 3505 C Saerkinen et al. 4754 D Barboza et al. 3536 E Barboza et al. 3520 F Saerkinen et al. 4010 G Atchison &amp; Gagnon 25 H Barboza et al. 3473). Photos by S. Knapp, G. Atchison, and T. Saerkinen." figureDoi="10.3897/phytokeys.231.100894.figure2" httpUri="https://binary.pensoft.net/fig/897463" pageId="0" pageNumber="1">Figs 2F</figureCitation>
<figureCitation id="568B603BCF7239129791F6EC147E290E" captionStart="Figure 110" captionStartId="F110" captionText="Figure 110. Solanum pallidum A habit B habit with larger leaves C detail of adaxial leaf surface D detail of abaxial leaf surface E flower bud F dissected flower G fruit (A, C-G Knapp et al. 10445 B Knapp et al. 10444). Illustration by R. Wise." figureDoi="10.3897/phytokeys.231.100894.figure110" httpUri="https://binary.pensoft.net/fig/897571" pageId="0" pageNumber="1">, 110</figureCitation>
<figureCitation id="F5E91E47A18874A1A03D6FE860B973AD" captionStart="Figure 111" captionStartId="F111" captionText="Figure 111. Solanum pallidum A habit B flower buds with dense indumentum of dendritic trichomes C flowers at full anthesis D maturing fruits (A Saerkinen et al. 4010 B Knapp et al. 10433 C Saerkinen et al. 4042 D Saerkinen et al. 4010). Photos by S. Knapp and T. Saerkinen." figureDoi="10.3897/phytokeys.231.100894.figure111" httpUri="https://binary.pensoft.net/fig/897572" pageId="0" pageNumber="1">, 111</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="59B831C958385EAC8BC114F3717D22F4" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="58148ADEC18EF17E1AAF4CAAFF5B2445" pageId="0" pageNumber="1">
<taxonomicName id="9F5F15621736E8E98EC657F38E287E71" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum lechleri" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="lechleri">Solanum lechleri</taxonomicName>
Rusby, Bull. Torrey Bot. Club 26: 193. 1899. Type. Bolivia. La Paz: Prov. Larecaja, Guanai [Guanay], May 1886,
<emphasis id="181CBEFC1772D7271E3391C716CA5F74" italics="true" pageId="0" pageNumber="1">H.H. Rusby 790</emphasis>
(no herbaria cited; lectotype, designated here: NY [00172060]; isolectotypes: GH [00077701], NY [00172059, 00743694], PH [00030433], US [00027648, acc. # 1416231; 01014267, acc. # 32604]).
</paragraph>
<paragraph id="CA9141EB8AA820212B6D6F0272B55387" pageId="0" pageNumber="1">
<taxonomicName id="68B195F442186C606EBE7189F497B495" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum lilacinum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="lilacinum">Solanum lilacinum</taxonomicName>
Rusby, Bull. Torrey Bot. Club 26: 192. 1899. Type. Bolivia. La Paz: Prov. Nor Yungas, Unduavi, Oct 1885,
<emphasis id="B6EADF8116A0CD000DBF1CB7E09AA57C" italics="true" pageId="0" pageNumber="1">H.H. Rusby 779</emphasis>
(no herbaria cited; lectotype, designated here: NY [00172067]; isolectotypes: BM [BM000778229], NY [00172068, 00172069], US [00027653, acc. # 32597], WIS [v0256196WIS]).
</paragraph>
<paragraph id="33F7592CB8215FB81BFD6B9759BA1357" pageId="0" pageNumber="1">
<taxonomicName id="20E6FC8E7DBBD760BAC13490940AB395" authorityName="Rusby" authorityYear="1907" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum rosulatum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rosulatum">Solanum rosulatum</taxonomicName>
Rusby, Bull. New York Bot. Gard. 4: 418. 1907. Type. Bolivia. Sin. loc., [no date],
<emphasis id="305D555CD5140220A62CD62E32AD7331" italics="true" pageId="0" pageNumber="1">M. Bang 2518</emphasis>
(no herbaria cited; lectotype, designated here: NY [00172157]; isolectotype: US [00027779, acc. # 1324745]).
</paragraph>
<paragraph id="EE3680FD312349124F15AF560E880CB9" pageId="0" pageNumber="1">
<taxonomicName id="9FBDA590647316AD143EEBF3B58BF7FF" authorityName="Rusby" authorityYear="1907" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum symmetrifolium" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="symmetrifolium">Solanum symmetrifolium</taxonomicName>
Rusby, Bull. New York Bot. Gard. 4: 418. 1907. Type. Bolivia. Sin loc., [no date],
<emphasis id="60E1BD9639B6EF3D7AD8BD68F2358ED9" italics="true" pageId="0" pageNumber="1">M. Bang 2870</emphasis>
(no herbaria cited; lectotype, designated here: NY [00172200]; isolectotypes: K [K000585654, K000585655], NY [00172201]).
</paragraph>
<paragraph id="F0B72E5F8605F5BA34D3329D2FD63C62" pageId="0" pageNumber="1">
<taxonomicName id="053236531386C96C4404347AA25C8D43" authorityName="Rusby" authorityYear="1907" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum sarachioides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sarachioides">Solanum sarachioides</taxonomicName>
Rusby, Bull. New York Bot. Gard. 4: 420. 1907, nom illeg., non
<taxonomicName id="617B068ADA04B28E590FEBC745573E87" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum sarrachoides" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sarrachoides">Solanum sarrachoides</taxonomicName>
Sendtn. (1846). Type. Bolivia. Sin. loc., [no date],
<emphasis id="9BCD620E7085C8D778E27C392B223143" italics="true" pageId="0" pageNumber="1">M. Bang 2517</emphasis>
(no herbaria cited; lectotype, designated here: NY [00172168]; isolectotype US [00027789, acc. # 1416169]).
</paragraph>
<paragraph id="B0257E6DD9BED80E1A609195120740B6" pageId="0" pageNumber="1">
<taxonomicName id="92CF6F18F0C45ACC03211241AED067DE" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum buchtienii" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="buchtienii">Solanum buchtienii</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 10: 558. 1912. Type. Bolivia. La Paz: Prov. Nor Yungas, Unduavi, 12 Feb 1907,
<emphasis id="39C0014CF08B9A110A63BFA9A284AB74" italics="true" pageId="0" pageNumber="1">O. Buchtien 765</emphasis>
(no herbaria cited; lectotype, designated here: HBG [HBG-511410]).
</paragraph>
<paragraph id="69BC2B6383C9D4EA8C4B527DC6DDCFA4" pageId="0" pageNumber="1">
<taxonomicName id="781810B711047B6236C72F6A7A851A72" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum subauriferum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="subauriferum">Solanum subauriferum</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 10: 559. 1912. Type. Bolivia. La Paz: Prov. Sud Yungas, Sirupaya prope Yanacachi, 22 Nov 1906,
<emphasis id="814845DC2989C2AD7A42E8D7950A276E" italics="true" pageId="0" pageNumber="1">O. Buchtien 332</emphasis>
(lectotype, designated here: US [00027813, acc. # 1175818, as
<normalizedToken id="64E909058D73B9003FDBA0331EFDCA68" originalValue="“322”">&quot;322&quot;</normalizedToken>
]; isolectotypes: NY [00824366], WRSL).
</paragraph>
<paragraph id="B282FC635FB2B6D7033277D9F4174134" pageId="0" pageNumber="1">
<taxonomicName id="8AA5667EED448112DE6CA8E92EEBF5B9" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum scotinonectarium" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="scotinonectarium">Solanum scotinonectarium</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 10: 560. 1912. Type. Bolivia. La Paz: Prov. Sud Yungas, &quot;Sirupay bei Yanacachi&quot;, 22 Nov 1906,
<emphasis id="8D9EF1EF70B9F0234ECC1CCAADE2473D" italics="true" pageId="0" pageNumber="1">O. Buchtien 332</emphasis>
(lectotype, designated here: US [00027813, acc. # 1175818]; isolectotype: NY [00824366], WRSL [not seen]).
</paragraph>
<paragraph id="8CCB7C3C056AA1C1E55BFB7E6917A246" pageId="0" pageNumber="1">
<taxonomicName id="BFA99BA09DFAF088B25D8F5DBEA3718B" authorityName="Bitter" authorityYear="1912" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum planifurcum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="planifurcum">Solanum planifurcum</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 11: 2. 1912. Type. Peru. Puno: Prov. Sandia, sin. loc., 2,100-2,500 m, 6 Apr 1902,
<emphasis id="B25E9DD11250ACF46D64DFC5E2BC3B1B" italics="true" pageId="0" pageNumber="1">A. Weberbauer 685</emphasis>
(holotype: B, destroyed [F neg. 2631]; lectotype, designated here: F [v0076175F, acc. # 647966, fragment of holotype).
</paragraph>
<paragraph id="FE8771CC0D1265FEA7EEFE8DB170F955" pageId="0" pageNumber="1">
<taxonomicName id="9377ABEFADC6928FFC11D80284B64EA6" authorityName="Bitter" authorityYear="1913" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum sandianum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sandianum">Solanum sandianum</taxonomicName>
Bitter, Bot. Jahrb. Syst. 50, Beibl. 111: 62. 1913. Type. Peru. Puno: Prov. Sandia, supra Cuyocuyo, 3,800 m,
<emphasis id="7BFD096D5276478AEC207264BEDE58AB" italics="true" pageId="0" pageNumber="1">A. Weberbauer 930</emphasis>
(holotype B, destroyed [F. neg. 2636]; no duplicates found); Peru. Puno: Prov. Sandia, km 137 on road from Cuyocuyo to Quiscupunco, 3641 m, 21 Mar 2012,
<emphasis id="72E6413CB3429E99BD078A6BAE4063C9" italics="true" pageId="0" pageNumber="1">
T.
<normalizedToken id="B5E67989E924B97DD74D538B3ABDAC00" originalValue="Särkinen">Saerkinen</normalizedToken>
, A. Mathews &amp; P.
<normalizedToken id="2B9CC59CEF8B7B9792A88FAA9E59F2AE" originalValue="Gonzáles">Gonzales</normalizedToken>
4055
</emphasis>
(neotype, designated here: USM [acc. # 00265491]; isoneotypes: BM [BM001120017, BM001120240, BM001120241]).
</paragraph>
</subSubSection>
<subSubSection id="0E758969D4FD93E9A2F3C398AD445ABA" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="D80BB95E066487F16B6D297FFEC7BF96" pageId="0" pageNumber="1">
<typeStatus id="5A31D26A919E0592CA33ECD670F01D0B">Type</typeStatus>
.
</paragraph>
<paragraph id="2FC8D6BA308EF3A672CE7490BF76192E" pageId="0" pageNumber="1">
<materialsCitation id="8EE8C32EDDBBBF232334C52D10D993C6" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="M. Bang" country="Bolivia" location="La Paz" specimenCount="1" stateProvince="La Paz" typeStatus="lectotype">
<collectingCountry id="1DFFF0062F7A7E589EDB63F4509DCC52" name="Bolivia">Bolivia</collectingCountry>
. Vic.
<collectingRegion id="0EE1DEBE896B8B144B84BF41516FCFC3" country="Bolivia" name="La Paz">La Paz</collectingRegion>
,
<emphasis id="29FB25EA88C8A34EEECC5B7D4AB40FC1" italics="true" pageId="0" pageNumber="1">
<collectorName id="B3254E294FA28C83135D268DF5A7272A">M. Bang</collectorName>
64
</emphasis>
[a] (no herbaria cited;
<typeStatus id="D064FDFD0FEC41306F125E283F122EB6">lectotype</typeStatus>
, designated here: NY [00172111]; isolectotypes: GH, LE, M [M-0165963], MO [MO-503708, acc. # 1815483], NY [00172112],
</materialsCitation>
<collectingCountry id="11B58457DFAE672AE96A5CD8D4CBB4AC" name="Philippines">PH</collectingCountry>
[00030385],
<collectingCountry id="12798B7EBD46BBD8FE32146B306CC728" name="United States of America">US</collectingCountry>
[00027725, acc. # 58341).
</paragraph>
<caption id="615B89E4B96DC90127697E5D8B9641E6" doi="10.3897/phytokeys.231.100894.figure110" httpUri="https://binary.pensoft.net/fig/897571" pageId="0" pageNumber="1" start="Figure 110" startId="F110">
<paragraph id="06E3F059A612E49AA6F8F58543DB74A3" pageId="0" pageNumber="1">
<emphasis id="6A45FF53A36C176B0712709DD86D7D49" bold="true" pageId="0" pageNumber="1">Figure 110.</emphasis>
<taxonomicName id="B9150465B1F983EE78C2F060C3A09245" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="A7D57FE9C4A70A9BD077D7EEC0BA4FDF" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
<emphasis id="9263A0D26A7321AAC08EBDF9448B0689" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis id="BBC037FB6374E15164F9568B39221E60" bold="true" pageId="0" pageNumber="1">B</emphasis>
habit with larger leaves
<emphasis id="FD2126D87E7B4D0CDA4CC13176FD5313" bold="true" pageId="0" pageNumber="1">C</emphasis>
detail of adaxial leaf surface
<emphasis id="7799D5558F18D4A71E693DFC925CEFFB" bold="true" pageId="0" pageNumber="1">D</emphasis>
detail of abaxial leaf surface
<emphasis id="97B58533CCF585890A03DFCCAE764FFD" bold="true" pageId="0" pageNumber="1">E</emphasis>
flower bud
<emphasis id="6D5AA5FA3D7B911E2FB7E7219BB50325" bold="true" pageId="0" pageNumber="1">F</emphasis>
dissected flower
<emphasis id="B3529E7973CAA18A0194C25FD5182FDC" bold="true" pageId="0" pageNumber="1">G</emphasis>
fruit (
<emphasis id="3B47E8A2585DC2D1257F4B6F7AABF5F5" bold="true" pageId="0" pageNumber="1">A, C-G</emphasis>
<emphasis id="745E64E7AD0E88CAB62E26FA5E6F6015" italics="true" pageId="0" pageNumber="1">Knapp et al. 10445</emphasis>
<emphasis id="ABBA9A4A2D6104886BC1BDA8D838BE38" bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="778D30D47E440E1C4C1F24145C421A51" italics="true" pageId="0" pageNumber="1">Knapp et al. 10444</emphasis>
). Illustration by R. Wise.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="768654299BAB61CD90A36A6CBF9CD64F" pageId="0" pageNumber="1" type="description">
<paragraph id="2EC9D46215A81222E3C338F2AC986905" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="60AECB693A02DA75F6C152C308E19DB0" pageId="0" pageNumber="1">
Scandent or lax shrub 1-3 high, with elongate branches. Stems terete, densely pubescent with transparent eglandular dendritic uniseriate trichomes 0.5-1 mm long; new growth densely pubescent with transparent eglandular dendritic uniseriate trichomes 0.5-1 mm long, these drying white or yellowish white in herbarium specimens; bark of older stems greenish brown, somewhat glabrescent. Sympodial units difoliate to plurifoliate, the leaves not geminate. Leaves simple, the blades 2.5-10 cm long, 1-5.2 cm wide, elliptic to ovate-elliptic, widest at the middle or in the lower third, membranous, discolorous; adaxial surfaces moderately and evenly pubescent with transparent dendritic uniseriate trichomes to 0.5 mm long; abaxial surfaces sparsely to densely pubescent with transparent eglandular dendritic uniseriate trichomes to 0.5 mm long, thinner and more delicate than the trichomes of the adaxial surfaces; principal veins 6-8 pairs, often drying yellowish tan; base cuneate to acute; margins entire; apex acute to acuminate; petioles 0.8-1 cm long, adaxially pubescent like the upper leaf surfaces. Inflorescences internodal or terminal at branch tips, forked to several times branched, 3-9(12) cm long, with 20-40 flowers clustered at the branch tips, moderately to densely pubescent with transparent eglandular dendritic uniseriate trichomes to 0.5 mm long like those of the stems; peduncle 1.5-4(6) cm long; pedicels 1-1.5 cm long, ca. 0.75 mm in diameter at the base, ca. 1.5 mm in diameter at the apex, tapering, spreading at anthesis, moderate to densely pubescent with transparent eglandular dendritic uniseriate trichomes to 0.5 mm long, articulated at the base; pedicel scars tightly packed at the ends of the inflorescence branches, 1-1.5 mm apart. Buds ellipsoid, the corolla strongly exserted from the calyx before anthesis, usually darker than the corolla in flower. Flowers 5-merous, cosexual (hermaphroditic). Calyx tube 1.5-2 mm long, conical, the lobes 1.5-2.5 mm long, ca. 1 mm wide, triangular, sometimes somewhat reflexed at anthesis, moderately pubescent with transparent eglandular dendritic trichomes to 0.5 mm long, like those of the pedicels. Corolla 2-2.3 cm in diameter, pale to dark purple with a green eye, stellate, lobed ca. halfway to the base, the lobes 6-9 mm long, 4-6 mm wide, deltate, spreading at anthesis, adaxially glabrous except for the papillate lobe tips, abaxially puberulent with white eglandular simple and dendritic trichomes where exposed in bud, these denser on the midveins and tips, the interpetalar tissue more glabrous. Stamens equal; filament tube minute to 0.1 mm long; free portion of the filaments 1-1.5 mm long, very densely pubescent adaxially with tangled simple uniseriate trichomes. Ovary conical, glabrous; style 6-8 mm long, straight, exserted beyond the anther cone, densely pubescent in the lower part with transparent simple uniseriate trichomes to 0.5 mm long; stigma small-capitate, the surfaces minutely papillate. Fruit a globose berry, 0.8-1 cm in diameter, green when immature, ripening to blackish purple, the pericarp thin, matte, opaque, glabrous; fruiting pedicels 1.3-1.4 cm long, ca. 1.2 mm in diameter at the base and apex, not markedly woody, deflexed, not persistent; fruiting calyx not markedly accrescent, the lobes to 4 mm long and slightly reflexed at the tips, the tube appressed to the berry. Seeds ca. 20 per berry, ca. 2 mm long, ca. 1.5 mm wide, flattened and teardrop shaped, reddish gold, the surfaces minutely pitted, the testal cells sinuate in outline. Stone cells 6 per berry, scattered through the mesocarp, ca. 1 mm in diameter, cream-coloured, two of the inclusions slightly smaller. Chromosome number: 2n = 24 (
<bibRefCitation id="92FB457C3A5CEFD8B47F94A2F5F5B02D" DOI="https://doi.org/10.12705/665.29" author="Chiarini, FE" editor="Marhold, K" journalOrPublisher="Taxon 66 (5)" pageId="0" pageNumber="1" pagination="1246 - 1247" refId="B40" refString="Chiarini, FE, Lipari, D, Barboza, GE, Knapp, S, 2017. Solanaceae. In: Marhold, K, Bresitwiser, I, Eds., IAPT/IOPB chromosome data 25. Taxon 66(5): 1246 - 1247, DOI: https://doi.org/10.12705/665.29" title="Solanaceae." url="https://doi.org/10.12705/665.29" volumeTitle="IAPT / IOPB chromosome data 25." year="2017">Chiarini et al. 2017</bibRefCitation>
, voucher
<emphasis id="F76CEB9D7A600880041849B5A40D1E0D" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="FC8F950A4098E1909EC851915E686DEA" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4014
</emphasis>
).
</paragraph>
<caption id="6395C5020FEF49641A81BDE25A0BA61C" doi="10.3897/phytokeys.231.100894.figure111" httpUri="https://binary.pensoft.net/fig/897572" pageId="0" pageNumber="1" start="Figure 111" startId="F111">
<paragraph id="0796F70A749B741022D33C6966C70B1F" pageId="0" pageNumber="1">
<emphasis id="8D073A7AD2CBE0CBBE7EFAC06D08108A" bold="true" pageId="0" pageNumber="1">Figure 111.</emphasis>
<taxonomicName id="9EF042807098BD7B55BD3FD2DB15CD95" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="1249C7821C860BBBE6BF5817D845EB08" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
<emphasis id="32CA694538743D9AAACCD7979238BF74" bold="true" pageId="0" pageNumber="1">A</emphasis>
habit
<emphasis id="CEEC52FBFC170357972FC0D0EF65203B" bold="true" pageId="0" pageNumber="1">B</emphasis>
flower buds with dense indumentum of dendritic trichomes
<emphasis id="F13340885A5AEC8FBA62DF2881A4D762" bold="true" pageId="0" pageNumber="1">C</emphasis>
flowers at full anthesis
<emphasis id="8604C5DC625D313E2E61272658B00C95" bold="true" pageId="0" pageNumber="1">D</emphasis>
maturing fruits (
<emphasis id="020F5048B780C5E5FB8571A492D73E60" bold="true" pageId="0" pageNumber="1">A</emphasis>
<emphasis id="BAE0509425392346A520D4A3C5F0664A" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="B7FB256F9A08C049D27EA208378C199F" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4010
</emphasis>
<emphasis id="62EE70C067EC9AB317BCBB7D8C87EDBA" bold="true" pageId="0" pageNumber="1">B</emphasis>
<emphasis id="C2B51809AF0238E8D22E584C941971A4" italics="true" pageId="0" pageNumber="1">Knapp et al. 10433</emphasis>
<emphasis id="A1C27FF8F7A2D3744FFBCC4F98575138" bold="true" pageId="0" pageNumber="1">C</emphasis>
<emphasis id="93BC659B22BB7F3850C5719F6ECD8A1E" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="AA4B4D19017BBA64C0C126944960607B" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4042
</emphasis>
<emphasis id="06D9F22A9442587295BF0498A5637306" bold="true" pageId="0" pageNumber="1">D</emphasis>
<emphasis id="823DCEBF3CD43763135E07A73D9C7B79" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="6F7418A06F4B498DF9481475FC3D54A6" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4010
</emphasis>
). Photos by S. Knapp and T.
<normalizedToken id="C6C83AA2CCA3E88716150DD358ACEB9B" originalValue="Särkinen">Saerkinen</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7C3776DD2A23A69062C1A075BD7B3647" pageId="0" pageNumber="1" type="distribution">
<paragraph id="9169C5E71DD9320F738E599D267C99FF" pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph id="2F356BD335DCF1801C37A86062109142" pageId="0" pageNumber="1">
<emphasis id="F50E6DA9641800992BEDD20A5A72AAE6" bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation id="298216AC4F4474540EABD0F645DDF83B" captionStart="Figure 112" captionStartId="F112" captionText="Figure 112. Distribution map of Solanum pallidum." figureDoi="10.3897/phytokeys.231.100894.figure112" httpUri="https://binary.pensoft.net/fig/897573" pageId="0" pageNumber="1">112</figureCitation>
).
</emphasis>
<taxonomicName id="5E6FDE5DB2E751794F6B12BBF83F69EB" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="A11BC72B393B3D41E920256BA170EDDB" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
occurs from south-central Peru (Depts.
<normalizedToken id="F0825CD1DB82405805A39754E7D8A3AF" originalValue="Huánuco">Huanuco</normalizedToken>
, Ayacucho, Cusco, Puno) to northern Bolivia (Depts. Cochabamba, La Paz) on the eastern Andean slopes.
</paragraph>
<caption id="7BBE6595186B9CBD34593638DD4D9B06" doi="10.3897/phytokeys.231.100894.figure112" httpUri="https://binary.pensoft.net/fig/897573" pageId="0" pageNumber="1" start="Figure 112" startId="F112">
<paragraph id="48B38A4454E12F7B0E51F140E49A560E" pageId="0" pageNumber="1">
<emphasis id="BE02A2841D3BE3CA29EA6D7EA6FA98AC" bold="true" pageId="0" pageNumber="1">Figure 112.</emphasis>
Distribution map of
<taxonomicName id="3F15B58D0F07C43E451D2A2C2AF1D322" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="334CCF2FAE49DFB22DCA36C11FC5E776" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D8A2F1D362888C154065E295E9551990" pageId="0" pageNumber="1" type="ecology">
<paragraph id="3982C0338216FC8AFDA4FC8E57FD568F" pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph id="23F90970292966D0E73D8F67A1702A3B" pageId="0" pageNumber="1">
<taxonomicName id="3FB1386640720678A7660D4099CF0039" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="F4BD236D7E18207C2EFBB686A1081986" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
grows in cloud forests, cloud forest edges and clearings, roadsides and montane scrub, from (600-)1,200 to 4,000 m elevation. Most specimens have been collected between 2,000 and 2,800 m elevation.
</paragraph>
</subSubSection>
<subSubSection id="913528974180F5720C4367BE3EEF2565" pageId="0" pageNumber="1" type="common names and uses">
<paragraph id="30DA008F891E49D49CBFEA97205EFFA2" pageId="0" pageNumber="1">Common names and uses.</paragraph>
<paragraph id="4A711F3D07D999D6D3EDD94DC964AE98" pageId="0" pageNumber="1">
Peru. Cusco: muyuqhaya (
<emphasis id="7B189F972C683AC56691975D1D62BE9B" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="0642718B0A9865377EC4A88213828802" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 5284
</emphasis>
). No uses recorded.
</paragraph>
</subSubSection>
<subSubSection id="271232C9B2482F5A8DF5EF54C3521F15" pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph id="B9D7408F13956742EFF878A04FC4B26B" pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
<paragraph id="E17309ECBF66D7E4AF67A8E6DC7E5FA7" pageId="0" pageNumber="1">
<emphasis id="CA78F192CAAD1F07659A574AF83D7EB9" bold="true" pageId="0" pageNumber="1">
(
<bibRefCitation id="F32F659BE4AC12452B8C1461E0620E40" author="Ibarrola, DA" journalOrPublisher="Nueva Serie" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/resources/redlistguidelines" refId="B116" refString="2022. . https://www.iucnredlist.org/resources/redlistguidelines" url="https://www.iucnredlist.org/resources/redlistguidelines" year="2022">IUCN 2022</bibRefCitation>
).
</emphasis>
Least Concern [LC]. EOO = 140,455 km2 [LC]; AOO = 340 km2 [EN].
<taxonomicName id="8B358E2E2064EBD53D01E04A33DF907C" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="A5DD06539382585CA5F29DBEA59E4022" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
is a weedy shrub of landslides and road edges and has a relatively wide distribution; it occurs around many of the protected archaeological sites in the region of Cusco, Peru.
</paragraph>
</subSubSection>
<subSubSection id="0AB1E7827CD9AAA0FD9B47EA7EB51CC7" pageId="0" pageNumber="1" type="discussion">
<paragraph id="3C11740DA9C02E85E92EE404FAFA6D9E" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="9720F288E95D0DC9CB9DCCCA44BEA23A" pageId="0" pageNumber="1">
<taxonomicName id="6AC3E298047F83CF59A70CDC3840B14D" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pallidum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="pallidum">
<emphasis id="5E1B9302D0E6920EEAD2AA9F0FB08885" italics="true" pageId="0" pageNumber="1">Solanum pallidum</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName id="EC4630EEBDF97F528F4FCB44BE376D1F" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
<emphasis id="0AA96FB51A1E0517D0665E2DB3D12F96" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
</taxonomicName>
in its large flowers and highly branched inflorescences usually on long peduncles but is easily distinguished from it by often dense pubescence composed of branched (dendritic) trichomes on all plant parts. The two taxa are somewhat sympatric, but
<taxonomicName id="255089D25611E49D1A07CCFA0697C09B" family="Solanaceae" kingdom="Plantae" lsidName="S. pallidum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pallidum">
<emphasis id="29ACDDEFB58ABCBB519CFED34F782390" italics="true" pageId="0" pageNumber="1">S. pallidum</emphasis>
</taxonomicName>
is confined to the eastern Andean slopes, while
<taxonomicName id="FCED5698EA85B68DB43AEA2FF9AB809C" family="Solanaceae" kingdom="Plantae" lsidName="S. cochabambense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="cochabambense">
<emphasis id="EE218A2073CD18DFA2B86882738FD74E" italics="true" pageId="0" pageNumber="1">S. cochabambense</emphasis>
</taxonomicName>
occurs on both slopes of the Andes in Peru. Molecular sequence data suggest the species are closely related (
<bibRefCitation id="939DB4F6F84113C21CCBE5902A138EAA" DOI="https://doi.org/10.1002/ajb2.1827" author="Gagnon, E" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" pagination="580 - 601" refId="B92" refString="Gagnon, E, Hilgenhof, R, Orejuela, A, McDonnell, A, Sablok, G, Aubriot, X, Giacomin, LL, Gouvea, Y, Bohs, L, Dodsworth, S, Martine, C, Poczai, P, Knapp, S, Saerkinen, T, 2022. Phylogenomic discordance suggests polytomies along the backbone of the large genus Solanum. American Journal of Botany 109 (4): 580 - 601, DOI: https://doi.org/10.1002/ajb2.1827" title="Phylogenomic discordance suggests polytomies along the backbone of the large genus Solanum." url="https://doi.org/10.1002/ajb2.1827" volume="109" year="2022">Gagnon et al. 2022</bibRefCitation>
).
</paragraph>
<paragraph id="213E57AB7EAA1BCCD5032DA1475C32F7" pageId="0" pageNumber="1">
The collection used to describe
<taxonomicName id="41C0007107836E5DAA5AFA70C57FFEB1" family="Solanaceae" kingdom="Plantae" lsidName="S. pallidum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pallidum">
<emphasis id="2E4AAA4C7D247D935CECFB1D654AAA82" italics="true" pageId="0" pageNumber="1">S. pallidum</emphasis>
</taxonomicName>
(
<emphasis id="E871BE330E7C503E9C2C038353BB2AA3" italics="true" pageId="0" pageNumber="1">Bang 64</emphasis>
;
<bibRefCitation id="58F150C41136B592264A189841E4CE0C" author="Rusby, HH" journalOrPublisher="Memoirs of the Torrey Botanical Club" pageId="0" pageNumber="1" pagination="203 - 274" refId="B198" refString="Rusby, HH, 1895. On the collections of Mr. Miguel Bang in Bolivia. Part II. Memoirs of the Torrey Botanical Club 4: 203 - 274" title="On the collections of Mr. Miguel Bang in Bolivia. Part II." volume="4" year="1895">Rusby 1895</bibRefCitation>
) is a mixed gathering of
<taxonomicName id="C375A148B66BCF326B92AF1512F12502" family="Solanaceae" kingdom="Plantae" lsidName="S. pallidum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pallidum">
<emphasis id="0AF6525F123F86657DE021DA22DED14B" italics="true" pageId="0" pageNumber="1">S. pallidum</emphasis>
</taxonomicName>
and
<taxonomicName id="CDE619C08CB8A7227089D30E4C4253D8" family="Solanaceae" kingdom="Plantae" lsidName="S. gonocladum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="gonocladum">
<emphasis id="00891CD4DED3CBA5813C034ADAC6AFA8" italics="true" pageId="0" pageNumber="1">S. gonocladum</emphasis>
</taxonomicName>
(see
<taxonomicName id="3D5187847C6C65B858EFB8967C9E4B55" family="Solanaceae" kingdom="Plantae" lsidName="S. gonocladum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="gonocladum">
<emphasis id="BB44865C2D81E5ED1A37936AE772C359" italics="true" pageId="0" pageNumber="1">S. gonocladum</emphasis>
</taxonomicName>
). We have selected the sheet at NY (barcode 00172111) that has the protologue attached and is annotated by Rusby as the lectotype for
<taxonomicName id="EDBC7D85D1DD2D5FF834A91320475CD7" family="Solanaceae" kingdom="Plantae" lsidName="S. pallidum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pallidum">
<emphasis id="F966F8187B777A4AA7F8AEC139C4BE27" italics="true" pageId="0" pageNumber="1">S. pallidum</emphasis>
</taxonomicName>
. Care must be taken with assigned duplicates of
<emphasis id="57DFC154021399F24A4BA4EDC0E9C568" italics="true" pageId="0" pageNumber="1">Bang 64</emphasis>
as isolectotypes, some sheets correspond to
<taxonomicName id="F9458A7EC403784E5B4140BC8AD72CF5" family="Solanaceae" kingdom="Plantae" lsidName="S. bangii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="bangii">
<emphasis id="7FDC0933DBF6039A99D98EA1EBD472BF" italics="true" pageId="0" pageNumber="1">S. bangii</emphasis>
</taxonomicName>
, a synonym of
<taxonomicName id="2DE4D2AC4EF4E42D8A933FD029A66678" family="Solanaceae" kingdom="Plantae" lsidName="S. gonocladum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="gonocladum">
<emphasis id="E22E6E972BE7A867C2231F4A1C7EBBFC" italics="true" pageId="0" pageNumber="1">S. gonocladum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="6F145561F1C8C6A4D3D7892FB5694084" pageId="0" pageNumber="1">
In subsequent years
<bibRefCitation id="976E3064B0FAF805ED6B937FC3165A01" DOI="https://doi.org/10.2307/2478226" author="Rusby, HH" journalOrPublisher="Bulletin of the Torrey Botanical Club" pageId="0" pageNumber="1" pagination="188 - 200" refId="B200" refString="Rusby, HH, 1899. An enumeration of the plants collected by H.H. Rusby in South America, 1895-1896. XXVII. Bulletin of the Torrey Botanical Club 26: 188 - 200, DOI: https://doi.org/10.2307/2478226" title="An enumeration of the plants collected by H. H. Rusby in South America, 1895 - 1896. XXVII." url="https://doi.org/10.2307/2478226" volume="26" year="1899">Rusby (1899</bibRefCitation>
,
<bibRefCitation id="38514B17449761F53D70E7650B6BAE08" author="Rusby, HH" journalOrPublisher="Bulletin of the New York Botanical Garden" pageId="0" pageNumber="1" pagination="309 - 470" refId="B201" refString="Rusby, HH, 1907. An enumeration of the plants collected in Bolivia by Miguel Bang. Part 4. Bulletin of the New York Botanical Garden 4: 309 - 470" title="An enumeration of the plants collected in Bolivia by Miguel Bang. Part 4." volume="4" year="1907">1907</bibRefCitation>
) described several names we here recognise as synonyms of
<taxonomicName id="1FEAE6F8F647FD6A1AF71FC229EBDEDF" family="Solanaceae" kingdom="Plantae" lsidName="S. pallidum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="pallidum">
<emphasis id="DAF9D6612E6709F308689390DA58CE17" italics="true" pageId="0" pageNumber="1">S. pallidum</emphasis>
</taxonomicName>
but cited no herbaria in the protologues We have lectotypified all of these with the specimens in NY (where the specimens that Rusby used are held) that best correspond to the protologues and are most complete (see above).
</paragraph>
<paragraph id="4DCE2C3A6D0686B782A55ECB47E038C1" pageId="0" pageNumber="1">
<bibRefCitation id="55265D92B1514277BC7589F5F4DB32A0" DOI="https://doi.org/10.1002/fedr.19120103305" author="Bitter, G" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="0" pageNumber="1" pagination="529 - 565" refId="B16" refString="Bitter, G, 1912a. Solana nova vel minus cognita I. Repertorium Specierum Novarum Regni Vegetabilis 10 (33-36): 529 - 565, DOI: https://doi.org/10.1002/fedr.19120103305" title="Solana nova vel minus cognita I." url="https://doi.org/10.1002/fedr.19120103305" volume="10" year="1912 a">Bitter (1912a)</bibRefCitation>
described
<taxonomicName id="0D6BDD8BF75F97ECEF6EE9B20C6E06F1" family="Solanaceae" kingdom="Plantae" lsidName="S. buchtienii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="buchtienii">
<emphasis id="F0A08F8A1369D8C4AA5DD172F60C664D" italics="true" pageId="0" pageNumber="1">S. buchtienii</emphasis>
</taxonomicName>
citing
<emphasis id="C2D28FE14669AE3149F5C907C893ED9F" italics="true" pageId="0" pageNumber="1">Buchtien 765</emphasis>
with the date of 12 Feb 1907 but no herbarium; of the duplicates we have seen, only that in HBG (HBG-511410) has that number and the correct date; we designate it here as the lectotype. Other collections (e.g., NY barcode 00139077 and US barcode 00027486, acc. # 700088) do not have the same collecting date (date of collection Nov 1910) and so are not part of the same gathering and thus are not isolectotypes. In the same publication Bitter described
<taxonomicName id="5A003A2580F2545674B1F3860C4F10DB" family="Solanaceae" kingdom="Plantae" lsidName="S. scotinonectarium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="scotinonectarium">
<emphasis id="0646A5E3B9993B0D9F18D041B2791776" italics="true" pageId="0" pageNumber="1">S. scotinonectarium</emphasis>
</taxonomicName>
citing
<emphasis id="16DA1246FACD50053FC35DC611B6D624" italics="true" pageId="0" pageNumber="1">Bang 31 pro parte</emphasis>
from &quot;herb. Berol.&quot; and
<emphasis id="B3EE16F776F91430ADE1FB8CB5589BDC" italics="true" pageId="0" pageNumber="1">Buchtien 332 pro parte</emphasis>
from &quot;herb. Buchtien! Vratisl.!&quot;. He often used different duplicates of the same collection to describe different taxa based on minor leaf shape and size differences; he also cited
<emphasis id="944C79DF25A0E2159D470E42EE14FAAD" italics="true" pageId="0" pageNumber="1">Buchtien 332 pro parte</emphasis>
as the basis for
<taxonomicName id="F66880D01473C27706C462AB5B4CDA77" family="Solanaceae" kingdom="Plantae" lsidName="S. subauriferum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="subauriferum">
<emphasis id="FAF6C9D92020FA814E4FA8C7C11E1373" italics="true" pageId="0" pageNumber="1">S. subauriferum</emphasis>
</taxonomicName>
without citing a herbarium but saying &quot;una cum
<taxonomicName id="239EC296E41A04C2F308B719ED8EF1A0" family="Solanaceae" kingdom="Plantae" lsidName="S. scotinonectarium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="scotinonectarium">
<emphasis id="FD192481A9CFA53312352D2C2B4EC972" italics="true" pageId="0" pageNumber="1">S. scotinonectarium</emphasis>
</taxonomicName>
&quot;.
<emphasis id="67C35CCADB19D588B49EC37D2646B6CA" italics="true" pageId="0" pageNumber="1">Bang 31</emphasis>
at &quot;herb. Berol&quot; also forms the basis for his
<taxonomicName id="6F0907696A858A8596F6B3CCFA45832A" family="Solanaceae" kingdom="Plantae" lsidName="S. irenaeum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="irenaeum">
<emphasis id="9031883F319A4BDF58519EAFB2C3F68E" italics="true" pageId="0" pageNumber="1">S. irenaeum</emphasis>
</taxonomicName>
(
<bibRefCitation id="21777F90723C64488F6BED2FACF3D2EA" DOI="https://doi.org/10.1002/fedr.19120103305" author="Bitter, G" journalOrPublisher="Repertorium Specierum Novarum Regni Vegetabilis" pageId="0" pageNumber="1" pagination="529 - 565" refId="B16" refString="Bitter, G, 1912a. Solana nova vel minus cognita I. Repertorium Specierum Novarum Regni Vegetabilis 10 (33-36): 529 - 565, DOI: https://doi.org/10.1002/fedr.19120103305" title="Solana nova vel minus cognita I." url="https://doi.org/10.1002/fedr.19120103305" volume="10" year="1912 a">Bitter 1912a</bibRefCitation>
), a synonym of
<taxonomicName id="672796A1745F30414F91D6BEDCE8723D" family="Solanaceae" kingdom="Plantae" lsidName="S. polytrichostylum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="polytrichostylum">
<emphasis id="BDEF55CD52A4CFB3019A196AFB34BEA1" italics="true" pageId="0" pageNumber="1">S. polytrichostylum</emphasis>
</taxonomicName>
. A sheet in US (barcode 00027813, acc. # 1175818) bears both the numbers 322 and 332 and is annotated &quot;
<taxonomicName id="0936BA9FA7BB5070CF65EE3111766706" family="Solanaceae" kingdom="Plantae" lsidName="S. subauriferum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="subauriferum">S. subauriferum</taxonomicName>
Bitt. n. sp.&quot; by Buchtien; we are interpreting the number 322 as an error to be corrected. In view of
<normalizedToken id="DEE9972EAD35D76187E58D5B84514052" originalValue="Bitters">Bitter's</normalizedToken>
practice of using different sheets of the same gathering to describe different species, the destruction of the WRSL sheet of
<emphasis id="E026345123315C9E13F001D540DE274E" italics="true" pageId="0" pageNumber="1">Buchtien 332</emphasis>
and the Buchtien Herbarium being held at US (
<bibRefCitation id="43452CCBC786AB1818EE65BE98A2AA67" author="Morton, CV" journalOrPublisher="Plant Science Bulletin" pageId="0" pageNumber="1" pagination="1 - 4" publicationUrl="https://cms.botany.org/psbarchive/issue/1966-v12-no-2.html" refId="B167" refString="Morton, CV, Stern, WL, 1966. The history of the United States national herbarium. Plant Science Bulletin 12 (2): 1 - 4, https://cms.botany.org/psbarchive/issue/1966-v12-no-2.html" title="The history of the United States national herbarium." url="https://cms.botany.org/psbarchive/issue/1966-v12-no-2.html" volume="12" year="1966">Morton and Stern 1966</bibRefCitation>
), we have designated the US sheet as the lectotype of both
<taxonomicName id="C0CFA08DEDFAE19A887D9DBF3E396EBF" family="Solanaceae" kingdom="Plantae" lsidName="S. scotinonectarium" order="Solanales" pageId="0" pageNumber="1" rank="species" species="scotinonectarium">
<emphasis id="4CDC161F220633C295987E3C260669C7" italics="true" pageId="0" pageNumber="1">S. scotinonectarium</emphasis>
</taxonomicName>
and
<taxonomicName id="9EE577E46E03FAB68AB354FF4189079A" family="Solanaceae" kingdom="Plantae" lsidName="S. subauriferum" order="Solanales" pageId="0" pageNumber="1" rank="species" species="subauriferum">
<emphasis id="99A33839F3A08F2A07C223CF9B9128DB" italics="true" pageId="0" pageNumber="1">S. subauriferum</emphasis>
</taxonomicName>
making these names homotypic.
</paragraph>
<paragraph id="CDD6432A082F47F41894D527D70120F0" pageId="0" pageNumber="1">
<taxonomicName id="1C62F5D1C28BCBE78AAA3C862773DAE0" authorityName="Bitter" authorityYear="1913" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum sandianum" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="sandianum">
<emphasis id="1F581B8A7D6F564A1E1DAE60770143E9" italics="true" pageId="0" pageNumber="1">Solanum sandianum</emphasis>
</taxonomicName>
was described (
<bibRefCitation id="2CBDD5CA4E7BD2A6DAD6EAEBF0E72342" author="Bitter, G" editor="Urban, I" journalOrPublisher="Botanisches Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 50, Beiblatt 111" pageId="0" pageNumber="1" pagination="58 - 67" refId="B19" refString="Bitter, G, 1913. Solana peruv., aequat., boliv. In: Urban, I, Ed., Plantae novae andinae imprimis Weberbauerianae VI. Botanisches Jahrbuecher fur Systematik, Pflanzengeschichte und Pflanzengeographie 50, Beiblatt 111: 58 - 67" title="Solana peruv., aequat., boliv." volumeTitle="Plantae novae andinae imprimis Weberbauerianae VI." year="1913">Bitter 1913</bibRefCitation>
) citing a collection made by August Weberbauer in southern Peru (
<emphasis id="9C958149E01418C53915E70ED57602F0" italics="true" pageId="0" pageNumber="1">Weberbauer 930</emphasis>
) held in Berlin (F. neg. 2636). That specimen is destroyed and we have found no duplicates, even at MOL where
<normalizedToken id="7097F2101B7EEFC5D901A3911238DEBE" originalValue="Weberbauers">Weberbauer's</normalizedToken>
original herbarium is held. We thus neotypify this name with a modern collection made near the type locality of Cuyocuyo in Sandia Province (
<emphasis id="A3870AD25D341E8C556726B5AB98B83C" italics="true" pageId="0" pageNumber="1">
<normalizedToken id="9248A5E19CC41D44027E4BD01CA5D159" originalValue="Särkinen">Saerkinen</normalizedToken>
et al. 4055
</emphasis>
) held in the Peruvian National Herbarium (USM).
</paragraph>
</subSubSection>
</treatment>
</document>