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<document id="9C8A634BC92CF8E7F782FE2E7FFEBAF3" ID-DOI="10.5281/zenodo.202515" ID-GBIF-Dataset="0c035728-2d80-4f1d-93f8-6478a00ad5cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="202515" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460271096384" checkinUser="plazi" docAuthor="Vogel, Gernot &amp; Luo, Jian" docDate="2011" docId="7C7ABE6E3873307F24A5F925FC17F829" docLanguage="en" docName="zt02807p040.pdf" docOrigin="Zootaxa 2807" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lycodon gongshan Vogel &amp; Luo, 2011, spec. nov." docType="treatment" docVersion="9" lastPageNumber="37" masterDocId="8043C616387730772432FFF8FFD8FF91" masterDocTitle="A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae)" masterLastPageNumber="40" masterPageNumber="29" pageNumber="33" updateTime="1698246841238" updateUser="plazi">
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<mods:title id="5D02C199C6D866312DF12C9ECEC1BC48">A new species of the genus Lycodon (Boie, 1826) from the southwestern mountains of China (Squamata: Colubridae)</mods:title>
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<mods:namePart id="7E0E41304C24BF842CB2855577536911">Vogel, Gernot</mods:namePart>
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<mods:namePart id="AD03516BD28F08EEF4DBACB8462B836A">Luo, Jian</mods:namePart>
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<mods:title id="B44929948FD7B3855F43185F1E43BCD0">Zootaxa</mods:title>
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<mods:date id="BC5DE4E5D455833BDC42F0E2757D0BF9">2011</mods:date>
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<treatment id="7C7ABE6E3873307F24A5F925FC17F829" ID-DOI="http://doi.org/10.5281/zenodo.5620317" ID-GBIF-Taxon="119395182" ID-Zenodo-Dep="5620317" LSID="urn:lsid:plazi:treatment:7C7ABE6E3873307F24A5F925FC17F829" httpUri="http://treatment.plazi.org/id/7C7ABE6E3873307F24A5F925FC17F829" lastPageId="8" lastPageNumber="37" pageId="4" pageNumber="33">
<subSubSection id="BCC95CF33873307324A5F925FED2F888" pageId="4" pageNumber="33" type="nomenclature">
<paragraph id="F46C0F783873307324A5F925FE33F967" blockId="4.[151,491,1757,1817]" box="[151,491,1757,1782]" pageId="4" pageNumber="33">
<heading id="AF24B8143873307324A5F925FE33F967" bold="true" box="[151,491,1757,1782]" fontSize="10" level="2" pageId="4" pageNumber="33" reason="2">
<emphasis id="C6A7D36A3873307324A5F925FE33F967" bold="true" box="[151,491,1757,1782]" pageId="4" pageNumber="33">
<taxonomicName id="33D374FB3873307324A5F925FEAAF967" box="[151,370,1757,1782]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="33" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A3873307324A5F925FEAAF967" bold="true" box="[151,370,1757,1782]" italics="true" pageId="4" pageNumber="33">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E1138733073254BF926FE33F967" box="[377,491,1758,1782]" pageId="4" pageNumber="33" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="F46C0F783873307324A5F8F9FED2F888" blockId="4.[151,491,1757,1817]" box="[151,266,1793,1817]" pageId="4" pageNumber="33">
(
<figureCitation id="6CE813FD3873307324ADF8F9FEDBF888" box="[159,259,1793,1817]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,714,737]" captionStartId-1="5.[151,250,1619,1642]" captionTargetBox-0="[151,1436,290,702]" captionTargetBox-1="[151,1436,806,1598]" captionTargetId-0="figure@5.[151,1436,290,702]" captionTargetId-1="figure@5.[151,1436,806,1598]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 3. Dorsal view of the preserved Holotype of Lycodon gongshan spec. nov., KIZ 730034, TL 963 mm. A) Dorsal view, B) ventral view. Photographs by Gernot Vogel." captionText-1="FIGURE 4 Lateral view of right side of the head of the preserved holotype of Lycodon gongshan spec. nov., KIZ 730034. Photograph by Gernot Vogel." httpUri-0="https://zenodo.org/record/202518/files/figure.png" httpUri-1="https://zenodo.org/record/202519/files/figure.png" pageId="4" pageNumber="33">Figs 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33873307224A5F8B2FEDEFF66" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33" type="materials_examined">
<paragraph id="F46C0F783873307324A5F8B2FB0FF814" blockId="4.[151,1437,1865,2033]" pageId="4" pageNumber="33">
<emphasis id="C6A7D36A3873307324A5F8B2FED3F8F3" bold="true" box="[151,267,1866,1890]" pageId="4" pageNumber="33">
<typeStatus id="2B68B1DA3873307324A5F8B2FEDEF8F3" box="[151,262,1866,1890]" pageId="4" pageNumber="33" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="92C297BD387330732520F8B2FE99F8F0" box="[274,321,1866,1889]" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="33">KIZ</collectionCode>
730034 adult male (tail dissected), from “Bapo,
<taxonomicName id="33D374FB38733073275FF8B1FC39F8F0" box="[877,993,1865,1889]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
,
<collectingCountry id="8CC44FE83873307327DCF8B1FBEEF8F0" box="[1006,1078,1865,1889]" name="China" pageId="4" pageNumber="33">China</collectingCountry>
”, today Dulongjiang Township,
<taxonomicName id="33D374FB3873307324A5F895FED1F814" box="[151,265,1901,1925]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
County, Yunnan Province,
<collectingCountry id="8CC44FE838733073260EF895FCB7F814" box="[572,879,1901,1925]" name="China" pageId="4" pageNumber="33">Peoples Republic of China</collectingCountry>
(
<figureCitation id="6CE813FD38733073274CF895FC3DF814" box="[894,997,1901,1925]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="5.[151,250,714,737]" captionStartId-1="5.[151,250,1619,1642]" captionTargetBox-0="[151,1436,290,702]" captionTargetBox-1="[151,1436,806,1598]" captionTargetId-0="figure@5.[151,1436,290,702]" captionTargetId-1="figure@5.[151,1436,806,1598]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="FIGURE 3. Dorsal view of the preserved Holotype of Lycodon gongshan spec. nov., KIZ 730034, TL 963 mm. A) Dorsal view, B) ventral view. Photographs by Gernot Vogel." captionText-1="FIGURE 4 Lateral view of right side of the head of the preserved holotype of Lycodon gongshan spec. nov., KIZ 730034. Photograph by Gernot Vogel." httpUri-0="https://zenodo.org/record/202518/files/figure.png" httpUri-1="https://zenodo.org/record/202519/files/figure.png" pageId="4" pageNumber="33">Figs 34</figureCitation>
). Collector unknown.
</paragraph>
<paragraph id="F46C0F783873307224F7F86AFEDEFF66" blockId="4.[151,1437,1865,2033]" lastBlockId="5.[151,1436,151,247]" lastPageId="5" lastPageNumber="34" pageId="4" pageNumber="33">
<emphasis id="C6A7D36A3873307324F7F86AFE69F83B" bold="true" box="[197,433,1937,1962]" pageId="4" pageNumber="33">
<typeStatus id="2B68B1DA3873307324F7F86AFEE5F83B" box="[197,317,1938,1962]" pageId="4" pageNumber="33" type="paratype">Paratypes</typeStatus>
(n = 3).
</emphasis>
<collectionCode id="92C297BD387330732585F86AFE3DF838" box="[439,485,1938,1961]" httpUri="http://grbio.org/cool/ncht-ribt" name="Kunming Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="33">KIZ</collectionCode>
730008, juvenile male (tail dissected), from “Bapo,
<taxonomicName id="33D374FB387330732043F869FB3DF838" box="[1137,1253,1937,1961]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
,
<collectingCountry id="8CC44FE83873307320CBF869FA98F838" box="[1273,1344,1937,1961]" name="China" pageId="4" pageNumber="33">China</collectingCountry>
”, today Dulongjiang Township,
<taxonomicName id="33D374FB387330732582F84DFDFAF85C" box="[432,546,1973,1997]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
County, Yunnan Province,
<collectingCountry id="8CC44FE838733073275BF84DFB76F85C" box="[873,1198,1973,1997]" name="China" pageId="4" pageNumber="33">Peoples Republic of China</collectingCountry>
; collector unknown.
<collectionCode id="92C297BD3873307324A5F822FF0CF860" box="[151,212,2010,2033]" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="4" pageNumber="33">HNU</collectionCode>
200609001, adult male, from Dulongjiang Township,
<taxonomicName id="33D374FB387330732709F821FC75F860" box="[827,941,2009,2033]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
County, Yunnan Province,
<collectingCountry id="8CC44FE83873307220D1F821FEFAFF3E" lastPageId="5" lastPageNumber="34" name="China" pageId="4" pageNumber="33">Peoples Republic of China</collectingCountry>
; collected by Qingbai Hou.
<collectionCode id="92C297BD387230722650FF60FD47FF3E" box="[610,671,152,175]" httpUri="http://grbio.org/cool/s4qq-4c7f" name="Hunan Normal University" pageId="5" pageNumber="34">HNU</collectionCode>
200505002, adult female, from Xiaoheishan, Longjiang Township, Longling County, Yunnan Province,
<collectingCountry id="8CC44FE838723072260AFF44FCACFF45" box="[568,884,188,212]" name="China" pageId="5" pageNumber="34">Peoples Republic of China</collectingCountry>
; collected by Yan Hengmei, Liang Hongbin and Guo Keji.
</paragraph>
</subSubSection>
<caption id="A0AC5FF03872307224A5FD32FD76FD6E" httpUri="https://zenodo.org/record/202518/files/figure.png" pageId="5" pageNumber="34" targetBox="[151,1436,290,702]" targetPageId="5">
<paragraph id="F46C0F783872307224A5FD32FD76FD6E" blockId="5.[151,1436,714,767]" pageId="5" pageNumber="34">
<emphasis id="C6A7D36A3872307224A5FD32FECDFD71" bold="true" box="[151,277,714,737]" pageId="5" pageNumber="34">FIGURE 3.</emphasis>
Dorsal view of the preserved Holotype of
<taxonomicName id="33D374FB3872307226EAFD33FC42FD70" box="[728,922,715,737]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="34" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A3872307226EAFD33FC42FD70" box="[728,922,715,737]" italics="true" pageId="5" pageNumber="34">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387230722790FD33FBD3FD71" box="[930,1035,715,736]" pageId="5" pageNumber="34" rank="species">
<emphasis id="C6A7D36A387230722790FD33FBD3FD71" bold="true" box="[930,1035,715,736]" pageId="5" pageNumber="34">spec. nov.</emphasis>
</taxonomicNameLabel>
, KIZ 730034, TL 963 mm. A) Dorsal view, B) ventral view. Photographs by Gernot Vogel.
</paragraph>
</caption>
<caption id="A0AC5FF03872307224A5F9ABFE40F919" httpUri="https://zenodo.org/record/202519/files/figure.png" pageId="5" pageNumber="34" targetBox="[151,1436,806,1598]" targetPageId="5">
<paragraph id="F46C0F783872307224A5F9ABFE40F919" blockId="5.[151,1436,1619,1672]" pageId="5" pageNumber="34">
<emphasis id="C6A7D36A3872307224A5F9ABFED4F9F8" bold="true" box="[151,268,1619,1642]" pageId="5" pageNumber="34">FIGURE 4</emphasis>
Lateral view of right side of the head of the preserved holotype of
<taxonomicName id="33D374FB38723072279EF9ACFBB3F9FB" box="[940,1131,1620,1642]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="34" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A38723072279EF9ACFBB3F9FB" box="[940,1131,1620,1642]" italics="true" pageId="5" pageNumber="34">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387230722043F9ACFB0FF9F8" box="[1137,1239,1620,1641]" pageId="5" pageNumber="34" rank="species">
<emphasis id="C6A7D36A387230722043F9ACFB0FF9F8" bold="true" box="[1137,1239,1620,1641]" pageId="5" pageNumber="34">spec. nov.</emphasis>
</taxonomicNameLabel>
, KIZ 730034. Photograph by Gernot Vogel.
</paragraph>
</caption>
<subSubSection id="BCC95CF33872307124F7F94DFB57FD41" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34" type="diagnosis">
<paragraph id="F46C0F783872307224F7F94DFA4CF810" blockId="5.[151,1437,1717,2030]" pageId="5" pageNumber="34">
<emphasis id="C6A7D36A3872307224F7F94DFEE7F95F" bold="true" box="[197,319,1717,1742]" pageId="5" pageNumber="34">Diagnosis.</emphasis>
A species of the genus
<taxonomicName id="33D374FB387230722679F94DFD73F95C" box="[587,683,1717,1741]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="C6A7D36A387230722679F94DFD73F95C" box="[587,683,1717,1741]" italics="true" pageId="5" pageNumber="34">Lycodon</emphasis>
</taxonomicName>
characterized by: (1) a loreal scale entering the orbit; (2) 17 dorsal scale rows anteriorly and at midbody; (3) upper and vertebral dorsal rows (612) are keeled; (4) 210216 ventrals in males, and 215 ventrals in the single female known; (5) 9596 subcaudals in males, and 92 subcaudals in the single female; (6) relative tail length ca.
<quantity id="332BA29D38723072260CF8D9FD3FF8A8" box="[574,743,1825,1849]" metricMagnitude="-3" metricUnit="m" metricValue="5.8801" metricValueMax="5.8928" metricValueMin="5.8674" pageId="5" pageNumber="34" unit="in" value="0.2315" valueMax="0.232" valueMin="0.231">0.2310.232 in</quantity>
males, and
<quantity id="332BA29D38723072275FF8D9FC11F8A8" box="[877,969,1825,1849]" metricMagnitude="-3" metricUnit="m" metricValue="5.461" pageId="5" pageNumber="34" unit="in" value="0.215">0.215 in</quantity>
the female; (7) 8 supralabials with SL 4 6 touching the orbit; (7) 3240 pale bands on a dark body; (8) breadth of the anteriormost band 1 dorsal scale middorsally, 34 ventrals, midventrally; (9) belly with discreet bands throughout, and (10) dark ventrals on the throat.
</paragraph>
<paragraph id="F46C0F783872307124F7F875FDBBFED2" blockId="5.[151,1437,1717,2030]" lastBlockId="6.[151,1437,155,2019]" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
This species can be recognized by its long tail (tail length/total length
<quantity id="332BA29D3872307227E8F875FB5BF834" box="[986,1155,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="5.8801" metricValueMax="5.8928" metricValueMin="5.8674" pageId="5" pageNumber="34" unit="in" value="0.2315" valueMax="0.232" valueMin="0.231">0.2310.232 in</quantity>
males,
<quantity id="332BA29D3872307220E8F875FAEEF834" box="[1242,1334,1933,1957]" metricMagnitude="-3" metricUnit="m" metricValue="5.461" pageId="5" pageNumber="34" unit="in" value="0.215">0.215 in</quantity>
females, vs.
<quantity id="332BA29D3872307224F2F849FEB2F858" box="[192,362,1969,1993]" metricMagnitude="-3" metricUnit="m" metricValue="5.3721" metricValueMax="5.715" metricValueMin="5.0292" pageId="5" pageNumber="34" unit="in" value="0.2115" valueMax="0.225" valueMin="0.198">0.1980.225 in</quantity>
males,
<quantity id="332BA29D3872307225F7F849FDB7F858" box="[453,623,1969,1993]" metricMagnitude="-3" metricUnit="m" metricValue="5.2324" metricValueMax="5.5626" metricValueMin="4.9022" pageId="5" pageNumber="34" unit="in" value="0.206" valueMax="0.219" valueMin="0.193">0.1930.219 in</quantity>
females of
<taxonomicName id="33D374FB3872307226CAF84AFCA3F858" box="[760,891,1969,1993]" class="Squamata" family="Colubridae" genus="Ophites" higherTaxonomySource="CoL" kingdom="Animalia" pageId="5" pageNumber="34" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3872307226CAF84AFCDFF858" box="[760,775,1970,1993]" italics="true" pageId="5" pageNumber="34">L</emphasis>
.
<emphasis id="C6A7D36A38723072272AF849FCA3F858" box="[792,891,1969,1993]" italics="true" pageId="5" pageNumber="34">fasciatus</emphasis>
</taxonomicName>
), and the high number of subcaudals (
<quantity id="332BA29D387230722104F849FA44F858" box="[1334,1436,1969,1993]" metricMagnitude="0" metricUnit="m" metricValue="2.4257" metricValueMax="2.4384" metricValueMin="2.413" pageId="5" pageNumber="34" unit="in" value="95.5" valueMax="96.0" valueMin="95.0">9596 in</quantity>
males, and
<quantity id="332BA29D387230722525F82DFE89F87C" box="[279,337,2005,2029]" metricMagnitude="0" metricUnit="m" metricValue="2.3368" pageId="5" pageNumber="34" unit="in" value="92.0">92 in</quantity>
females). Also, its anteriormost pale band is very narrow, being 1 dorsal scale broad middorsally in all 4 known specimens, vs. 15 dorsals (
<emphasis id="C6A7D36A38713071264AFF66FD5EFF28" box="[632,646,158,185]" italics="true" pageId="6" pageNumber="35">x</emphasis>
=
<quantity id="332BA29D387130712695FF63FD3EFF22" box="[679,742,155,179]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" pageId="6" pageNumber="35" unit="in" value="2.5">2.5 in</quantity>
<specimenCount id="E2D5C4F13871307126DFFF63FC5EFF22" box="[749,902,155,179]" pageId="6" pageNumber="35" type="generic">62 specimens</specimenCount>
) in the
<taxonomicName id="33D374FB3871307127EAFF64FB80FF22" box="[984,1112,155,179]" class="Squamata" family="Colubridae" genus="Ophites" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3871307127EAFF64FB80FF22" box="[984,1112,155,179]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
group, with the anteriormost band starting at ventral 47 (vs. usually ventral
<quantity id="332BA29D387130712682FF47FCD7FF46" box="[688,783,191,215]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="4.064" metricValueMin="2.54" pageId="6" pageNumber="35" unit="in" value="13.0" valueMax="16.0" valueMin="10.0">10-16 in</quantity>
<taxonomicName id="33D374FB387130712724FF39FC40FF49" box="[790,920,192,216]" class="Squamata" family="Colubridae" genus="Ophites" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387130712724FF39FC40FF49" box="[790,920,192,216]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
and in species of the
<taxonomicName id="33D374FB3871307120BEFF39FAD4FF49" box="[1164,1292,192,216]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="C6A7D36A3871307120BEFF39FAD4FF49" box="[1164,1292,192,216]" italics="true" pageId="6" pageNumber="35">L. ruhstrati</emphasis>
</taxonomicName>
group). Furthermore, the new species has a dark throat, whereas in
<taxonomicName id="33D374FB387130712731FF1DFC5BFF6D" box="[771,899,228,252]" class="Squamata" family="Colubridae" genus="Ophites" higherTaxonomySource="CoL" kingdom="Animalia" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387130712731FF1DFC5BFF6D" box="[771,899,228,252]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
usually the throat is white (except in some individuals from parts of Yunnan. Vogel
<emphasis id="C6A7D36A387130712601FEF1FDBCFEB1" box="[563,612,264,288]" italics="true" pageId="6" pageNumber="35">et al</emphasis>
. unpublished data). Detailed comparisons with other species of the genus
<taxonomicName id="33D374FB3871307124A5FED4FF2FFED5" box="[151,247,300,324]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="genus">
<emphasis id="C6A7D36A3871307124A5FED4FF2FFED5" box="[151,247,300,324]" italics="true" pageId="6" pageNumber="35">Lycodon</emphasis>
</taxonomicName>
appear below in the Discussion.
</paragraph>
<paragraph id="F46C0F783871307124F7FEB7FB57FD41" blockId="6.[151,1437,155,2019]" pageId="6" pageNumber="35">
<emphasis id="C6A7D36A3871307124F7FEB7FEB8FEF9" bold="true" box="[197,352,335,360]" pageId="6" pageNumber="35">Comparison.</emphasis>
Superficially,
<taxonomicName id="33D374FB38713071263BFEA8FD38FEF9" box="[521,736,336,360]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A38713071263BFEA8FD38FEF9" box="[521,736,336,360]" italics="true" pageId="6" pageNumber="35">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E113871307126D8FEA8FC87FEF9" box="[746,863,336,360]" pageId="6" pageNumber="35" rank="species">
<emphasis id="C6A7D36A3871307126D8FEA8FC87FEF9" bold="true" box="[746,863,336,360]" pageId="6" pageNumber="35">spec. nov.</emphasis>
</taxonomicNameLabel>
resembles other species of the
<taxonomicName id="33D374FB3871307120FDFEA8FA43FEF9" box="[1231,1435,336,360]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3871307120FDFEA8FA43FEF9" box="[1231,1435,336,360]" italics="true" pageId="6" pageNumber="35">Lycodon fasciatus</emphasis>
</taxonomicName>
group. It differs from
<taxonomicName id="33D374FB3871307125BBFE8DFDD1FE1D" box="[393,521,372,396]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3871307125BBFE8DFDD1FE1D" box="[393,521,372,396]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
by its much longer tail, more ventrals in males (210216 vs. 190211), more subcaudals in males (9496 vs. 7490) and females (92 vs. 6788). In
<taxonomicName id="33D374FB3871307127A3FE61FBC5FE21" box="[913,1053,408,432]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan">
<emphasis id="C6A7D36A3871307127A3FE61FC78FE21" box="[913,928,409,432]" italics="true" pageId="6" pageNumber="35">L</emphasis>
.
<emphasis id="C6A7D36A387130712782FE60FBC5FE21" box="[944,1053,408,432]" italics="true" pageId="6" pageNumber="35">gongshan</emphasis>
</taxonomicName>
in all eight occurrences only four lower labials touch the anterior chin shields, whereas in
<taxonomicName id="33D374FB38713071272EFE45FC78FE45" box="[796,928,444,468]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A38713071272EFE45FCF4FE45" box="[796,812,445,468]" italics="true" pageId="6" pageNumber="35">L</emphasis>
.
<emphasis id="C6A7D36A38713071270EFE44FC78FE45" box="[828,928,444,468]" italics="true" pageId="6" pageNumber="35">fasciatus</emphasis>
</taxonomicName>
this is usually five, and only rarely four (on both sides only in five of
<specimenCount id="E2D5C4F138713071258EFE27FD82FE66" box="[444,602,479,503]" pageId="6" pageNumber="35" type="generic">64 specimens</specimenCount>
). The anteriormost band lies closer to the head in
<taxonomicName id="33D374FB3871307120A1FE19FAC7FE69" box="[1171,1311,480,504]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan">
<emphasis id="C6A7D36A3871307120A1FE19FAC7FE69" box="[1171,1311,480,504]" italics="true" pageId="6" pageNumber="35">L. gongshan</emphasis>
</taxonomicName>
(first band starts at ventral 47 vs.usually
<quantity id="332BA29D3871307125CBFDFBFD88FD8A" box="[505,592,515,539]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" metricValueMax="4.572" metricValueMin="2.032" pageId="6" pageNumber="35" unit="in" value="13.0" valueMax="18.0" valueMin="8.0">818 in</quantity>
<taxonomicName id="33D374FB38713071266AFDFDFD02FD8D" box="[600,730,516,540]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A38713071266AFDFDFD02FD8D" box="[600,730,516,540]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
, rarely
<quantity id="332BA29D387130712707FDFBFCA0FD8A" box="[821,888,515,539]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="6" pageNumber="35" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5-8 in</quantity>
five of
<specimenCount id="E2D5C4F13871307127E6FDFBFBAAFD8A" box="[980,1138,515,539]" pageId="6" pageNumber="35" type="generic">67 specimens</specimenCount>
), and is very narrow, only one dorsal scale broad in the four available specimens, vs. usually
<quantity id="332BA29D3871307127AEFDDFFC22FDAE" box="[924,1018,551,575]" metricMagnitude="-2" metricUnit="m" metricValue="9.525" metricValueMax="15.24" metricValueMin="3.81" pageId="6" pageNumber="35" unit="in" value="3.75" valueMax="6.0" valueMin="1.5">1.56 in</quantity>
<taxonomicName id="33D374FB387130712031FDD1FB5EFDD1" box="[1027,1158,552,576]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387130712031FDD1FB5EFDD1" box="[1027,1158,552,576]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
(rarely one dorsal scale broad, 5 of
<specimenCount id="E2D5C4F1387130712525FDB3FE6BFDF2" box="[279,435,587,611]" pageId="6" pageNumber="35" type="generic">67 specimens</specimenCount>
). The number of subcaudals and tail length are sexually dimorphic in the new species, but not in
<taxonomicName id="33D374FB3871307124D2FD89FEBAFD19" box="[224,354,624,648]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3871307124D2FD89FEBAFD19" box="[224,354,624,648]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
(
<tableCitation id="B9513AC3387130712542FD97FE11FD16" box="[368,457,623,647]" captionStart="TABLE 2" captionStartId="3.[151,235,727,748]" captionTargetBox="[151,1436,807,1028]" captionTargetPageId="3" captionText="TABLE 2. The comparison of distinguishing characters of the holotype of Ophites fasciatus with the new species and Lycodon" httpUri="http://table.plazi.org/id/A0AC5FF03874307424A5FD2FFA44FD7C" pageId="6" pageNumber="35" tableUuid="A0AC5FF03874307424A5FD2FFA44FD7C">Table 2</tableCitation>
). Lastly,
<taxonomicName id="33D374FB387130712602FD89FD63FD19" box="[560,699,624,648]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A387130712602FD89FD63FD19" box="[560,699,624,648]" italics="true" pageId="6" pageNumber="35">L. gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E113871307126F1FD89FCF6FD19" box="[707,814,625,648]" pageId="6" pageNumber="35" rank="species">spec. nov</taxonomicNameLabel>
reaches a larger size than
<taxonomicName id="33D374FB38713071206AFD89FB01FD19" box="[1112,1241,624,648]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A38713071206AFD89FB01FD19" box="[1112,1241,624,648]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
: two of the three available males of the new species are considerably larger than the largest of the
<specimenCount id="E2D5C4F138713071201EFD6BFB49FD3A" box="[1068,1169,659,683]" pageId="6" pageNumber="35" type="male">32 males</specimenCount>
of
<taxonomicName id="33D374FB38713071208AFD6DFAE1FD3D" box="[1208,1337,660,684]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A38713071208AFD6DFAE1FD3D" box="[1208,1337,660,684]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
at hand; the single female of the new species is larger than the 36 examined females of
<taxonomicName id="33D374FB387130712035FD41FB50FD41" box="[1031,1160,696,720]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387130712035FD41FB50FD41" box="[1031,1160,696,720]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33871307124F7FD24FE67FC39" pageId="6" pageNumber="35" type="reference_group">
<paragraph id="F46C0F783871307124F7FD24FE67FC39" blockId="6.[151,1437,155,2019]" pageId="6" pageNumber="35">
<taxonomicName id="33D374FB3871307124F7FD24FD8EFD62" authority="Werner, 1922" authorityName="Werner" authorityYear="1922" box="[197,598,731,756]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="C6A7D36A3871307124F7FD24FE60FD65" box="[197,440,732,756]" italics="true" pageId="6" pageNumber="35">Dinodon yunnanensis</emphasis>
<bibRefCitation id="904272893871307125F2FD24FD8EFD62" author="Werner" box="[448,598,731,755]" pageId="6" pageNumber="35" refString="Werner, F. (1922) Neue Reptilien aus Sud-China, gesammelt von Dr. H. Handel-Mazzetti. Anzeiger der Akademie der Wissenschaften in Wien, 59, 220 - 222." type="journal article" year="1922">Werner, 1922</bibRefCitation>
</taxonomicName>
was described from Yunnanfu, now Kunming, Yunnan Province,
<collectingCountry id="8CC44FE8387130712108FD23FA58FD62" box="[1338,1408,731,755]" name="China" pageId="6" pageNumber="35">China</collectingCountry>
. It is currently regarded as a synonym of
<taxonomicName id="33D374FB387130712674FCF8FCC9FC89" box="[582,785,768,792]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387130712674FCF8FCC9FC89" box="[582,785,768,792]" italics="true" pageId="6" pageNumber="35">Lycodon fasciatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="90427289387130712712FCF8FC7CFC89" author="Pope" box="[800,932,768,792]" pageId="6" pageNumber="35" refString="Pope, C. H. (1935) The reptiles of China. Turtles, crocodilians, snakes, lizards. Natural History of central Asia, X. American Museum of Natural History, New York, i-lii + 604 pp. + 27 Pls." type="book" year="1935">Pope, 1935</bibRefCitation>
;
<bibRefCitation id="90427289387130712782FCF8FB4FFC89" author="Zhao" box="[944,1175,768,792]" pageId="6" pageNumber="35" refString="Zhao, E. M. &amp; Adler, K. (1993) Herpetology of China. Society for the Study of Amphibians and Reptiles, Athens, Ohio, Contributions to Herpetology, number 10. 522 pp. + 48 pls." type="book" year="1993">Zhao &amp; Adler, 1993</bibRefCitation>
).
<collectingCountry id="8CC44FE8387130712099FCF8FB12FC89" box="[1195,1226,768,792]" name="American Samoa" pageId="6" pageNumber="35">As</collectingCountry>
with
<emphasis id="C6A7D36A387130712139FCF8FF38FCAD" italics="true" pageId="6" pageNumber="35">
<taxonomicName id="33D374FB387130712139FCF8FF05FCAD" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">Lycodon fasciatus</taxonomicName>
,
</emphasis>
the
<typeStatus id="2B68B1DA387130712524FCDBFEA0FCAA" box="[278,376,803,827]" pageId="6" pageNumber="35" type="holotype">holotype</typeStatus>
of
<taxonomicName id="33D374FB387130712590FCDCFD40FCAD" box="[418,664,804,828]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="C6A7D36A387130712590FCDCFD40FCAD" box="[418,664,804,828]" italics="true" pageId="6" pageNumber="35">Dinodon yunnanensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="33D374FB387130712701FCDDFC18FCAD" box="[819,960,804,828]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A387130712701FCDDFC18FCAD" box="[819,960,804,828]" italics="true" pageId="6" pageNumber="35">L. gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E113871307127FBFCDCFBE6FCAD" box="[969,1086,804,828]" pageId="6" pageNumber="35" rank="species">
<emphasis id="C6A7D36A3871307127FBFCDCFBE6FCAD" bold="true" box="[969,1086,804,828]" pageId="6" pageNumber="35">spec. nov.</emphasis>
</taxonomicNameLabel>
in the number of ventrals and subcaudals, in the relative tail length, and in having fewer bands on the body + tail (23+7 vs. 3240 +
<quantity id="332BA29D387130712128FCB0FAA6FCF1" box="[1306,1406,840,864]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="6" pageNumber="35" unit="in" value="14.0" valueMax="16.0" valueMin="12.0">1216 in</quantity>
<taxonomicName id="33D374FB3871307121B7FCB1FEDCFC15" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A3871307121B7FCB1FEDCFC15" italics="true" pageId="6" pageNumber="35">L. gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E1138713071253FFC94FE5AFC15" box="[269,386,876,900]" pageId="6" pageNumber="35" rank="species">
<emphasis id="C6A7D36A38713071253FFC94FE5AFC15" bold="true" box="[269,386,876,900]" pageId="6" pageNumber="35">spec. nov.</emphasis>
</taxonomicNameLabel>
). A comparison of
<taxonomicName id="33D374FB387130712657FC95FD2AFC15" box="[613,754,876,900]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A387130712657FC95FD2AFC15" box="[613,754,876,900]" italics="true" pageId="6" pageNumber="35">L. gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E113871307126CEFC94FCA8FC15" box="[764,880,876,900]" pageId="6" pageNumber="35" rank="species">
<emphasis id="C6A7D36A3871307126CEFC94FCA8FC15" bold="true" box="[764,880,876,900]" pageId="6" pageNumber="35">spec. nov.</emphasis>
</taxonomicNameLabel>
with species of the
<taxonomicName id="33D374FB387130712069FC95FB07FC15" box="[1115,1247,876,900]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387130712069FC95FB07FC15" box="[1115,1247,876,900]" italics="true" pageId="6" pageNumber="35">L. fasciatus</emphasis>
</taxonomicName>
and
<taxonomicName id="33D374FB387130712128FC95FA44FC15" box="[1306,1436,876,900]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="ruhstrati">
<emphasis id="C6A7D36A387130712128FC95FA44FC15" box="[1306,1436,876,900]" italics="true" pageId="6" pageNumber="35">L. ruhstrati</emphasis>
</taxonomicName>
groups is given in Table 4.
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33871307124F7FC4CFE5FFB82" pageId="6" pageNumber="35" type="etymology">
<paragraph id="F46C0F783871307124F7FC4CFE5FFB82" blockId="6.[151,1437,155,2019]" pageId="6" pageNumber="35">
<emphasis id="C6A7D36A3871307124F7FC4CFE93FC5D" bold="true" box="[197,331,948,972]" pageId="6" pageNumber="35">Etymology.</emphasis>
The specific name refers to the area of origin of this species,
<taxonomicName id="33D374FB387130712027FC4BFB5FFC5A" box="[1045,1159,947,971]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
County. We suggest the following common names:
<taxonomicName id="33D374FB3871307125E3FC20FD9BFC61" box="[465,579,984,1008]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
Baihuanshe (Chinese),
<taxonomicName id="33D374FB387130712767FC20FC1FFC61" box="[853,967,984,1008]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
Wolf Snake (English),
<taxonomicName id="33D374FB3871307120E4FC20FA9FFC61" box="[1238,1351,984,1008]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
Wolfszahnnatter (
<collectingCountry id="8CC44FE8387130712528FC03FEA3FB82" box="[282,379,1019,1043]" name="Germany" pageId="6" pageNumber="35">German</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33871307124F7FBE7FC53F8C3" pageId="6" pageNumber="35" type="description">
<paragraph id="F46C0F783871307124F7FBE7FCC5F89B" blockId="6.[151,1437,155,2019]" pageId="6" pageNumber="35">
<emphasis id="C6A7D36A3871307124F7FBE7FDCCFBA9" bold="true" box="[197,532,1055,1080]" pageId="6" pageNumber="35">
Description of the
<typeStatus id="2B68B1DA387130712594FBD8FDD7FBA9" box="[422,527,1056,1080]" pageId="6" pageNumber="35" type="holotype">holotype</typeStatus>
.
</emphasis>
<emphasis id="C6A7D36A387130712629FBD8FDACFBA9" box="[539,628,1056,1080]" italics="true" pageId="6" pageNumber="35">Habitus</emphasis>
. Body elongate, somewhat laterally compressed; head flattened anteriorly, well distinct from neck; snout depressed and elongated; nostril oval, large. Eye moderate, pupil vertically elliptic. SVL
<quantity id="332BA29D38713071251DFB90FE57FB11" box="[303,399,1128,1152]" metricMagnitude="-1" metricUnit="m" metricValue="7.4" pageId="6" pageNumber="35" unit="mm" value="740.0">740 mm</quantity>
; TaL
<quantity id="332BA29D3871307125FCFB90FDF6FB11" box="[462,558,1128,1152]" metricMagnitude="-1" metricUnit="m" metricValue="2.23" pageId="6" pageNumber="35" unit="mm" value="223.0">223 mm</quantity>
; TL
<quantity id="332BA29D387130712651FB90FD1CFB11" box="[611,708,1128,1152]" metricMagnitude="-1" metricUnit="m" metricValue="9.629999999999999" pageId="6" pageNumber="35" unit="mm" value="963.0">963 mm</quantity>
.
<emphasis id="C6A7D36A3871307126FCFB90FCADFB11" box="[718,885,1128,1152]" italics="true" pageId="6" pageNumber="35">Body scalation</emphasis>
. 212 ventrals (+ 1 preventral), 96 subcaudals, all paired. Anal entire. Dorsal scales in 171715 rows, the 6 upper rows feebly keeled. Vertebral row not enlarged. No apical pit detected.
<emphasis id="C6A7D36A387130712591FB48FD96FB59" box="[419,590,1200,1224]" italics="true" pageId="6" pageNumber="35">Head scalation</emphasis>
. Rostral triangular, hardly visible from above; nasal vertically divided by furrow below and above nostril; two small internasals, widely in contact with each other and prefrontals; two large prefrontals, longer and wider than internasals; rather small, more or less triangular frontal; 2 large parietals, each bordered by three large scales, 2 upper temporals and larger paraparietal posteriorly; 1/1 supraocular; 1/1 preocular, above posterior part of loreal; 2/2 subequal postoculars; 1/1 loreal, rather elongate and narrow, entering orbit, in contact with SL 2 and 3, preocular, prefrontal and posterior part of nasal; 8/8 SL, SL
<quantity id="332BA29D387130712066FA9BFB43FAEA" box="[1108,1179,1379,1403]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="6" pageNumber="35" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">12 in</quantity>
contact with nasal, SL 35 entering orbit, 6th SL largest; 2+2/2+3 temporals, lower anterior temporal slightly broader than upper; 8/8 infralabials,
<collectingCountry id="8CC44FE8387130712510FA57FEE4FA57" box="[290,316,1455,1478]" name="Iceland" pageId="6" pageNumber="35">IL</collectingCountry>
<quantity id="332BA29D387130712571FA56FE52FA57" box="[323,394,1454,1478]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="6" pageNumber="35" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">14 in</quantity>
contact with first pair of chin shields anterior and posterior chin shields about same length on right, on left anterior larger than posterior, anterior pair wider.
<emphasis id="C6A7D36A38713071274FFA2AFB75FA7B" box="[893,1197,1490,1514]" italics="true" pageId="6" pageNumber="35">Coloration in preservative</emphasis>
. Body and tail dark brown, with 37 crossbands on body and 15 on tail, anteriorly narrow, becoming wider posteriorly; crossbands about 1 dorsal scale broad middorsally, widening ventrolaterally to about 34 dorsals broad; first crossband beginning at level of ventral 5; second crossband 4 scales behind first; crossbands on body darker centered with dark brown speckles; crossbands on tail also darkly speckled. Head uniformly blackish-brown; no collar. Underside of head dark on anterior third, otherwise cream; throat cream with dark, cloudy speckles on first four ventrals. Venter dark with clear, rather irregular cream bands, 2-3 ventrals broad, with 24 ventrals between each band. Within bands some ventrals dark on one half and cream on other, especially on posterior of body. Under surface of tail banded as body venter, clear cream bands about 3 subcaudals broad.
</paragraph>
<paragraph id="F46C0F783871307124F7F8EEFC53F8C3" blockId="6.[151,1437,155,2019]" pageId="6" pageNumber="35">
<emphasis id="C6A7D36A3871307124F7F8EEFEE1F8BE" bold="true" box="[197,313,1814,1839]" pageId="6" pageNumber="35">Variation.</emphasis>
The
<typeStatus id="2B68B1DA38713071254AF8EFFE3EF8BE" box="[376,486,1815,1839]" pageId="6" pageNumber="35" type="paratype">paratypes</typeStatus>
, agree in most respects with the description of the
<typeStatus id="2B68B1DA387130712009F8EEFB79F8BF" box="[1083,1185,1814,1838]" pageId="6" pageNumber="35" type="holotype">holotype</typeStatus>
. A comparison of the most important morphological characters is summarized in
<tableCitation id="B9513AC3387130712703F8C2FC5EF8C3" box="[817,902,1850,1874]" captionStart="TABLE 3" captionStartId="7.[151,235,152,173]" captionTargetBox="[151,1436,232,968]" captionTargetPageId="7" captionText="TABLE 3. Scalation of the four type specimens of Lycodon gongshan spec. nov. * = holotype. 1: small part of the tail tip missing." httpUri="http://table.plazi.org/id/A0AC5FF03870307024A5FF60FF66FF5D" pageId="6" pageNumber="35" tableUuid="A0AC5FF03870307024A5FF60FF66FF5D">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33871307124F5F8A6FEB1F82F" pageId="6" pageNumber="35" type="distribution">
<paragraph id="F46C0F783871307124F5F8A6FEB1F82F" blockId="6.[151,1437,155,2019]" pageId="6" pageNumber="35">
<emphasis id="C6A7D36A3871307124F5F8A6FEB8F8E6" bold="true" box="[199,352,1886,1911]" pageId="6" pageNumber="35">Distribution.</emphasis>
<collectingCountry id="8CC44FE8387130712558F8A6FE69F8E7" box="[362,433,1886,1910]" name="China" pageId="6" pageNumber="35">China</collectingCountry>
.
<taxonomicName id="33D374FB38713071258CF8A6FD4DF8E7" box="[446,661,1886,1910]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="35" phylum="Chordata" rank="species" species="gongshan">
<emphasis id="C6A7D36A38713071258CF8A6FD4DF8E7" box="[446,661,1886,1910]" italics="true" pageId="6" pageNumber="35">Lycodon gongshan</emphasis>
</taxonomicName>
is presently only known from the
<taxonomicName id="33D374FB387130712019F8A6FB45F8E7" box="[1067,1181,1886,1910]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
County and Longling County areas, in Yunnan Province,
<collectingCountry id="8CC44FE838713071260FF87AFC5BF80B" box="[573,899,1922,1946]" name="China" pageId="6" pageNumber="35">Peoples Republic of China</collectingCountry>
. Both localities are situated in the Hengduan Mountains (
<figureCitation id="6CE813FD38713071252FF85EFE83F82F" box="[285,347,1958,1982]" captionStart="FIGURE 5" captionStartId="8.[151,250,1300,1323]" captionTargetBox="[151,1436,267,1273]" captionTargetId="figure@8.[151,1436,267,1273]" captionTargetPageId="8" captionText="FIGURE 5. Map showing the distribution of Lycodon gongshan spec. nov. (red squares 1 - 2), Lycodon fasciatus (green squares 3 - 7), “ Dinodon yunnanensis ” (blue square 8), and Lycodon synaptor (yellow square 9). 1. “ Bapo ”, now Dulongjiang Township, Gongshan County, Yunnan Province, Peoples Republic of China [Type locality of Lycodon gongshan spec. nov.]; 2. Xiaoheishan, Longjiang Township, Longling County, Yunnan Province, Peoples Republic of China; 3. “ Ponsee ”, now Bangxi, Yingjiang County, Yunnan Province, Peoples Republic of China [former type locality of Ophites fasciatus]; 4. “ Mogok ”, Mandalay Division, Myanmar [New type locality of Lycodon fasciatus]; 5. Tuantian, Tengchong County, Yunnan Province, Peoples Republic of China; 6. Husa, Longchuan County, Yunnan Province, Peoples Republic of China; 7. Ruili City, Yunnan Province, Peoples Republic of China; 8. “ Yunnanfu ”, now Kunming City, Yunnan Province, Peoples Republic of China [Type locality of Dinodon yunnanensis]; 9. “ Tongchuan ”, now Dongchuan District, Yunnan Province, Peoples Republic of China [Type locality of Lycodon synaptor]. “ For Lycodon fasciatus only specimens from the Gongshan area are shown in the map. ”" httpUri="https://zenodo.org/record/202520/files/figure.png" pageId="6" pageNumber="35">Fig 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33871307024F7F832FDA1FB74" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35" type="biology_ecology">
<paragraph id="F46C0F783871307124F7F832FC2FF873" blockId="6.[151,1437,155,2019]" box="[197,1015,1994,2019]" pageId="6" pageNumber="35">
<emphasis id="C6A7D36A3871307124F7F832FEFCF872" bold="true" box="[197,292,1994,2019]" pageId="6" pageNumber="35">Biology.</emphasis>
There is no information available on the biology of this species.
</paragraph>
<caption id="A0AC5FF03870307024A5FF60FF66FF5D" ID-Table-UUID="A0AC5FF03870307024A5FF60FF66FF5D" httpUri="http://table.plazi.org/id/A0AC5FF03870307024A5FF60FF66FF5D" pageId="7" pageNumber="36" targetBox="[151,1436,232,968]" targetIsTable="true" targetPageId="7">
<paragraph id="F46C0F783870307024A5FF60FF66FF5D" blockId="7.[151,1436,152,204]" pageId="7" pageNumber="36">
<emphasis id="C6A7D36A3870307024A5FF60FEDCFF3C" bold="true" box="[151,260,152,173]" pageId="7" pageNumber="36">TABLE 3.</emphasis>
Scalation of the four type specimens of
<taxonomicName id="33D374FB3870307026A9FF60FC82FF3F" box="[667,858,152,174]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A3870307026A9FF60FC82FF3F" box="[667,858,152,174]" italics="true" pageId="7" pageNumber="36">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387030702752FF60FC1FFF3C" box="[864,967,152,173]" pageId="7" pageNumber="36" rank="species">
<emphasis id="C6A7D36A387030702752FF60FC1FFF3C" bold="true" box="[864,967,152,173]" pageId="7" pageNumber="36">spec. nov.</emphasis>
</taxonomicNameLabel>
* = holotype. 1: small part of the tail tip missing.
</paragraph>
</caption>
<paragraph id="F46C0F783870307024ADFF10FEC3FC59" pageId="7" pageNumber="36">
<table id="86D3FDD83870CF8824A5FF10FA44FC59" box="[151,1436,232,968]" gridcols="5" gridrows="6" pageId="7" pageNumber="36">
<tr id="4AE30D3A3870CF8824A5FF10FA44FE34" box="[151,1436,232,421]" gridrow="0" pageId="7" pageNumber="36">
<th id="093264463870CF8824A5FF10FE7AFE34" box="[151,418,232,421]" gridcol="0" gridrow="0" pageId="7" pageNumber="36">Characters Sex SVL TaL Rel TL</th>
<th id="093264463870CF8825EEFF10FDBEFE34" box="[476,614,232,421]" gridcol="1" gridrow="0" pageId="7" pageNumber="36">KIZ 730034* adult male 740 223 0.232</th>
<th id="093264463870CF882686FF10FCE5FE34" box="[692,829,232,421]" gridcol="2" gridrow="0" pageId="7" pageNumber="36">KIZ 730008 juvenile male 236 71 0.231</th>
<th id="093264463870CF8827A4FF10FB9EFE34" box="[918,1094,232,421]" gridcol="3" gridrow="0" pageId="7" pageNumber="36">HNU 200609001 adult male 691 237 -1</th>
<th id="093264463870CF8820A3FF10FA44FE34" box="[1169,1436,232,421]" gridcol="4" gridrow="0" pageId="7" pageNumber="36">HNU 200505002 adult female 598 164 0.215</th>
</tr>
<tr id="4AE30D3A3870CF8824A5FE42FA44FE5E" box="[151,1436,442,463]" gridrow="1" pageId="7" pageNumber="36">
<th id="093264463870CF8824A5FE42FE7AFE5E" box="[151,418,442,463]" gridcol="0" gridrow="1" pageId="7" pageNumber="36">ASR</th>
<td id="093264463870CF8825EEFE42FDBEFE5E" box="[476,614,442,463]" gridcol="1" gridrow="1" pageId="7" pageNumber="36">17</td>
<td id="093264463870CF882686FE42FCE5FE5E" box="[692,829,442,463]" gridcol="2" gridrow="1" pageId="7" pageNumber="36">17</td>
<td id="093264463870CF8827A4FE42FB9EFE5E" box="[918,1094,442,463]" gridcol="3" gridrow="1" pageId="7" pageNumber="36">17</td>
<td id="093264463870CF8820A3FE42FA44FE5E" box="[1169,1436,442,463]" gridcol="4" gridrow="1" pageId="7" pageNumber="36">17</td>
</tr>
<tr id="4AE30D3A3870CF8824A5FE1BFA44FDDC" box="[151,1436,483,589]" gridrow="2" pageId="7" pageNumber="36">
<th id="093264463870CF8824A5FE1BFE7AFDDC" box="[151,418,483,589]" gridcol="0" gridrow="2" pageId="7" pageNumber="36">MSR VEN SC</th>
<td id="093264463870CF8825EEFE1BFDBEFDDC" box="[476,614,483,589]" gridcol="1" gridrow="2" pageId="7" pageNumber="36">17 212 96</td>
<td id="093264463870CF882686FE1BFCE5FDDC" box="[692,829,483,589]" gridcol="2" gridrow="2" pageId="7" pageNumber="36">17 216 95</td>
<td id="093264463870CF8827A4FE1BFB9EFDDC" box="[918,1094,483,589]" gridcol="3" gridrow="2" pageId="7" pageNumber="36">17 210 94 1</td>
<td id="093264463870CF8820A3FE1BFA44FDDC" box="[1169,1436,483,589]" gridcol="4" gridrow="2" pageId="7" pageNumber="36">17 215 92</td>
</tr>
<tr id="4AE30D3A3870CF8824A5FD9AFA44FD58" box="[151,1436,610,713]" gridrow="3" pageId="7" pageNumber="36">
<th id="093264463870CF8824A5FD9AFE7AFD58" box="[151,418,610,713]" gridcol="0" gridrow="3" pageId="7" pageNumber="36">Lo enters eye Dorsal bands Tail bands</th>
<td id="093264463870CF8825EEFD9AFDBEFD58" box="[476,614,610,713]" gridcol="1" gridrow="3" pageId="7" pageNumber="36">yes 37 15</td>
<td id="093264463870CF882686FD9AFCE5FD58" box="[692,829,610,713]" gridcol="2" gridrow="3" pageId="7" pageNumber="36">yes 40 16</td>
<td id="093264463870CF8827A4FD9AFB9EFD58" box="[918,1094,610,713]" gridcol="3" gridrow="3" pageId="7" pageNumber="36">yes 38 16</td>
<td id="093264463870CF8820A3FD9AFA44FD58" box="[1169,1436,610,713]" gridcol="4" gridrow="3" pageId="7" pageNumber="36">yes 32 13</td>
</tr>
<tr id="4AE30D3A3870CF8824A5FD25FA44FCD4" box="[151,1436,733,837]" gridrow="4" pageId="7" pageNumber="36">
<th id="093264463870CF8824A5FD25FE7AFCD4" box="[151,418,733,837]" gridcol="0" gridrow="4" pageId="7" pageNumber="36">First band at VEN no Broad base [VEN] Broad vertebral [Dorsals]</th>
<td id="093264463870CF8825EEFD25FDBEFCD4" box="[476,614,733,837]" gridcol="1" gridrow="4" pageId="7" pageNumber="36">5 4 1</td>
<td id="093264463870CF882686FD25FCE5FCD4" box="[692,829,733,837]" gridcol="2" gridrow="4" pageId="7" pageNumber="36">4 3.5 1</td>
<td id="093264463870CF8827A4FD25FB9EFCD4" box="[918,1094,733,837]" gridcol="3" gridrow="4" pageId="7" pageNumber="36">6 3 1</td>
<td id="093264463870CF8820A3FD25FA44FCD4" box="[1169,1436,733,837]" gridcol="4" gridrow="4" pageId="7" pageNumber="36">7 4 1</td>
</tr>
<tr id="4AE30D3A3870CF8824A5FC56FA44FC59" box="[151,1436,942,968]" gridrow="5" pageId="7" pageNumber="36" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="093264463870CF8824A5FC56FE7AFC59" box="[151,418,942,968]" gridcol="0" gridrow="5" pageId="7" pageNumber="36">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="F46C0F783870307024A5FC0DFDA1FB74" blockId="7.[151,1437,1013,2009]" pageId="7" pageNumber="36">
There are four specimens of
<taxonomicName id="33D374FB3870307025DAFC0DFD6DFB9C" box="[488,693,1013,1037]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3870307025DAFC0DFD6DFB9C" box="[488,693,1013,1037]" italics="true" pageId="7" pageNumber="36">Lycodon fasciatus</emphasis>
</taxonomicName>
available, collected in the Hengduan Mountains. All stem from Yunnan Province. These are KIZ
<date id="806D29B8387030702639FBE1FD99FBA0" box="[523,577,1049,1073]" pageId="7" pageNumber="36" value="1974-02">74 II</date>
0 263 from Tuantian, Tengchong County, KIZ
<date id="806D29B838703070207AFBE1FBABFBA0" box="[1096,1139,1049,1073]" pageId="7" pageNumber="36" value="1974-01">74 I</date>
0 145 from Husa, Longchuan County, KIZ
<date id="806D29B8387030702555FBC5FE40FBC4" box="[359,408,1085,1109]" pageId="7" pageNumber="36" value="1974-01">74 I</date>
0 0 35 from Gongwa, Longchuan County, and KIZ
<date id="806D29B83870307027C1FBC5FBF5FBC4" box="[1011,1069,1085,1109]" pageId="7" pageNumber="36" value="1974-02">74 II</date>
0 262 from Tengchong County. Longchuan, Tengchong and Yingjiang are located in the southwest of Gaoligong Mountain, in the Hengduan Mountains. The first three specimens are females, the last is male. All of them are typical
<taxonomicName id="33D374FB3870307020B7FB7EFADFFB0C" box="[1157,1287,1157,1181]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A3870307020B7FB7EFADFFB0C" box="[1157,1287,1157,1181]" italics="true" pageId="7" pageNumber="36">L. fasciatus</emphasis>
</taxonomicName>
and the morphology is well within the range of this species. Thus,
<taxonomicName id="33D374FB387030702730FB52FC5BFB50" box="[770,899,1193,1217]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387030702730FB52FC5BFB50" box="[770,899,1193,1217]" italics="true" pageId="7" pageNumber="36">L. fasciatus</emphasis>
</taxonomicName>
is sympatric (but perhaps not syntopic) with
<taxonomicName id="33D374FB3870307021B7FB52FEDCFB74" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A3870307021B7FB52FEDCFB74" italics="true" pageId="7" pageNumber="36">L. gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387030702539FB35FEA5FB74" box="[267,381,1229,1253]" pageId="7" pageNumber="36" rank="species">
<emphasis id="C6A7D36A387030702539FB35FEA5FB74" bold="true" box="[267,381,1229,1253]" pageId="7" pageNumber="36">spec. nov.</emphasis>
</taxonomicNameLabel>
in at least some areas.
</paragraph>
</subSubSection>
<subSubSection id="BCC95CF33870307F24F7FB09FC17F829" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36" type="reference_group">
<paragraph id="F46C0F783870307024F7FB09FEB8F994" blockId="7.[151,1437,1013,2009]" pageId="7" pageNumber="36">
<taxonomicName id="33D374FB3870307024F7FB09FE42FA98" box="[197,410,1265,1289]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A3870307024F7FB09FE42FA98" box="[197,410,1265,1289]" italics="true" pageId="7" pageNumber="36">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387030702591FB09FDCFFA98" box="[419,535,1265,1289]" pageId="7" pageNumber="36" rank="species">
<emphasis id="C6A7D36A387030702591FB09FDCFFA98" bold="true" box="[419,535,1265,1289]" pageId="7" pageNumber="36">spec. nov.</emphasis>
</taxonomicNameLabel>
seems to be endemic to the Hengduan Mountains, a range of about
<quantity id="332BA29D387030702115FB09FAA6FA98" box="[1319,1406,1265,1289]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="36" unit="km" value="500.0">500 km</quantity>
in length, reaching
<quantity id="332BA29D387030702561FAEDFE68FABC" box="[339,432,1301,1325]" metricMagnitude="3" metricUnit="m" metricValue="6.74" pageId="7" pageNumber="36" unit="m" value="6740.0">6,740 m</quantity>
a.s.l. This range is already known as a centre of endemism and high biodiversity (
<bibRefCitation id="90427289387030702160FAEDFF0BFAC0" author="Venell" pageId="7" pageNumber="36" refString="Venell, T. (2006) Conservation Biology in the Hengduan Mountains sub-alpine conifer forests. Unpublished report, Brandeis University, Waltham, USA, 40 pp." type="book" year="2006">Venell 2006</bibRefCitation>
;
<bibRefCitation id="904272893870307024ECFAC2FEA6FAC0" author="Fu" box="[222,382,1337,1361]" pageId="7" pageNumber="36" refString="Fu, C., Wang, J., Pu, Z., Zhang, S., Chen, H., Zhao, B., Chen, J. &amp; Wu, J. (2007) Elevational gradients of diversity for lizards and snakes in the Hengduan Mountains, China. Biodiversity and Conservation, 16, 707 - 726." type="journal article" year="2007">
Fu
<emphasis id="C6A7D36A387030702531FAC2FEECFAC0" box="[259,308,1337,1361]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2007
</bibRefCitation>
), but still seems to be relatively underexplored for reptiles and amphibians (
<bibRefCitation id="904272893870307020E0FAC1FA49FAC0" author="Stotz" box="[1234,1425,1337,1361]" pageId="7" pageNumber="36" refString="Stotz, D. F., Harris, E. J., Moskovits, D. K., Hao, K., Yi, S. &amp; Adelmann, G. W. (edts) (2003) China, Yunnan, Southern Gaoligongshan. Rapid biological inventories No 4. The Field Museum, Chicago, Illinois, 146 pp." type="book" year="2003">
Stotz
<emphasis id="C6A7D36A387030702126FAC2FA9EFAC0" box="[1300,1350,1337,1361]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2003
</bibRefCitation>
). In 2003, UNESCO included the area “Three Parallel Rivers of Yunnan” in the World Heritage List. One reason was explained as follows: “Northwest Yunnan is the area of richest biodiversity in
<collectingCountry id="8CC44FE838703070203AFA79FB93FA08" box="[1032,1099,1409,1433]" name="China" pageId="7" pageNumber="36">China</collectingCountry>
and may be the most biologically diverse temperate region on Earth. The property encompasses most of the natural habitats in the Hengduan Mountains, one of the world's most important remaining areas for the conservation of the Earth's biodiversity” (
<bibRefCitation id="904272893870307024ACFA15FE8CF994" author="UNESCO" box="[158,340,1517,1541]" pageId="7" pageNumber="36" refString="UNESCO (2003) Decisions adopted by the 27 th Session of the World heritage Committee in 2003, Paris, UNESCO World Heritage Centre, 179 pp." type="book" year="2003">UNESCO 2003</bibRefCitation>
).
</paragraph>
<paragraph id="F46C0F783870307F24F7F9E9FC73FF45" blockId="7.[151,1437,1013,2009]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36">
Previously, 23 reptile species were regarded as endemic to the Hengduan Mountains (
<bibRefCitation id="904272893870307020B6F9E9FAB8F9B8" author="Zhao" box="[1156,1376,1553,1578]" pageId="7" pageNumber="36" refString="Zhao, E. &amp; Yang, D. (Eds.) (1997) The Series of the Scientific Expedition to the Hengduan Mountains, Qinghai-Xizang Plateau. Amphibians and Reptiles of the Hengduan Mountains Region. The Comprehensive Scientific Expedition to the Qinghai-Xizang Plateau, Chinese Academy of Science. Beijing (Science Press), i-xiv + 303 pp., pls I-VIII [in Chinese]." type="book" year="1997">Zhao &amp; Yang 1997</bibRefCitation>
), and there are ten endemic snake species here, namely:
<taxonomicName id="33D374FB3870307026F9F9CDFB8FF9DC" authority="Werner, 1922" authorityName="Werner" authorityYear="1922" box="[715,1111,1589,1613]" class="Reptilia" family="Viperidae" genus="Gloydius" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="monticola">
<emphasis id="C6A7D36A3870307026F9F9CDFC7FF9DC" box="[715,935,1589,1613]" italics="true" pageId="7" pageNumber="36">Gloydius monticola</emphasis>
(
<bibRefCitation id="90427289387030702787F9CDFB97F9DC" author="Werner" box="[949,1103,1589,1613]" pageId="7" pageNumber="36" refString="Werner, F. (1922) Neue Reptilien aus Sud-China, gesammelt von Dr. H. Handel-Mazzetti. Anzeiger der Akademie der Wissenschaften in Wien, 59, 220 - 222." type="journal article" year="1922">Werner, 1922</bibRefCitation>
)
</taxonomicName>
(Yunnan Province),
<taxonomicName id="33D374FB38703070210CF9CDFE70F9E0" authority="Jiang, 1977" authorityName="Jiang" authorityYear="1977" class="Reptilia" family="Viperidae" genus="Ovophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="zayuensis">
<emphasis id="C6A7D36A38703070210CF9CDFEDBF9E0" italics="true" pageId="7" pageNumber="36">Ovophis zayuensis</emphasis>
(Jiang, 1977)
</taxonomicName>
(Tibet Autonomous Region),
<taxonomicName id="33D374FB38703070273DF9A1FBAEF9E0" box="[783,1142,1625,1649]" class="Reptilia" family="Viperidae" genus="Protobothrops" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="xiangchengensis">
<emphasis id="C6A7D36A38703070273DF9A1FBAEF9E0" box="[783,1142,1625,1649]" italics="true" pageId="7" pageNumber="36">Protobothrops xiangchengensis</emphasis>
</taxonomicName>
(Zhao, Jiang, &amp; Huang, 1978) (Sichuan and Yunnan provinces),
<taxonomicName id="33D374FB387030702656F985FBF7F904" authority="Hu &amp; Zhao, 1966" authorityName="Hu &amp; Zhao" authorityYear="1966" box="[612,1071,1661,1685]" class="Reptilia" family="Xenodermatidae" genus="Achalinus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="meiguensis">
<emphasis id="C6A7D36A387030702656F985FC82F904" box="[612,858,1661,1685]" italics="true" pageId="7" pageNumber="36">Achalinus meiguensis</emphasis>
Hu &amp; Zhao, 1966
</taxonomicName>
(Sichuan, Guizhou and Yunnan provinces),
<taxonomicName id="33D374FB38703070252FF959FD00F928" authority="Chernov, 1962" authorityName="Chernov" authorityYear="1962" box="[285,728,1697,1721]" class="Reptilia" family="Colubridae" genus="Calamaria" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="C6A7D36A38703070252FF959FDF2F928" box="[285,554,1697,1721]" italics="true" pageId="7" pageNumber="36">Calamaria yunnanensis</emphasis>
Chernov, 1962
</taxonomicName>
(Yunnan Province),
<taxonomicName id="33D374FB3870307027F7F959FA44F928" authority="Stejneger, 1929" authorityName="Stejneger" authorityYear="1929" box="[965,1436,1697,1721]" class="Reptilia" family="Colubridae" genus="Euprepiophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="perlaceus">
<emphasis id="C6A7D36A3870307027F7F959FB09F928" box="[965,1233,1697,1721]" italics="true" pageId="7" pageNumber="36">Euprepiophis perlaceus</emphasis>
(Stejneger, 1929)
</taxonomicName>
(Sichuan Province),
<taxonomicName id="33D374FB38703070254DF93DFCB2F94C" authority="Zhao &amp; Jiang, 1981" authorityName="Zhao &amp; Jiang" authorityYear="1981" box="[383,874,1733,1757]" class="Reptilia" family="Colubridae" genus="Oligodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="multizonatus">
<emphasis id="C6A7D36A38703070254DF93DFD58F94C" box="[383,640,1733,1757]" italics="true" pageId="7" pageNumber="36">Oligodon multizonatus</emphasis>
Zhao &amp; Jiang, 1981
</taxonomicName>
(Sichuan, Shaanxi and Gansu provinces),
<taxonomicName id="33D374FB38703070217FF93EFE03F890" authority="Guo &amp; Deng, 2009" authorityName="Guo &amp; Deng" authorityYear="2009" class="Reptilia" family="Pareatidae" genus="Pareas" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="nigriceps">
<emphasis id="C6A7D36A38703070217FF93EFF27F890" italics="true" pageId="7" pageNumber="36">Pareas nigriceps</emphasis>
Guo &amp; Deng, 2009
</taxonomicName>
(Yunnan Province),
<taxonomicName id="33D374FB3870307026F3F911FADBF890" authority="Jiang &amp; Zhao, 1983" authorityName="Jiang &amp; Zhao" authorityYear="1983" box="[705,1283,1769,1793]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="pentasupralabialis">
<emphasis id="C6A7D36A3870307026F3F911FBC7F890" box="[705,1055,1769,1793]" italics="true" pageId="7" pageNumber="36">Rhabdophis pentasupralabialis</emphasis>
Jiang &amp; Zhao, 1983
</taxonomicName>
(Sichuan and Yunnan provinces), and
<taxonomicName id="33D374FB387030702598F8F5FC33F8B4" authority="Guo, Liu, Feng &amp; He, 2008" authorityName="Guo, Liu, Feng &amp; He" authorityYear="2008" box="[426,1003,1805,1830]" class="Reptilia" family="Colubridae" genus="Thermophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="species" species="zhaoermii">
<emphasis id="C6A7D36A387030702598F8F5FD70F8B4" box="[426,680,1805,1829]" italics="true" pageId="7" pageNumber="36">Thermophis zhaoermii</emphasis>
Guo, Liu, Feng &amp; He, 2008
</taxonomicName>
(Sichuan Province) (
<bibRefCitation id="904272893870307020EEF8F5FA49F8B4" author="Luo" box="[1244,1425,1805,1829]" pageId="7" pageNumber="36" refString="Luo, J., Gao, H. Y., Liu, Y. M., Xu, L. &amp; Luo, Y. (2010) A revised checklist and distribution of Chinese snakes. Acta Herpetologica Sinica, 12, 67 - 91 [in Chinese]." type="journal article" year="2010">
Luo
<emphasis id="C6A7D36A387030702120F8F6FA9DF8B4" box="[1298,1349,1805,1829]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2010
</bibRefCitation>
). Several endemic subspecies of snakes have also been described from this area, and the following are sometimes regarded as valid but should be checked for their status:
<taxonomicName id="33D374FB387030702725F8ADFA88F8FC" authority="Zhao &amp; Kou, 1987" authorityName="Zhao &amp; Kou" authorityYear="1987" box="[791,1360,1877,1901]" class="Reptilia" family="Colubridae" genus="Sibynophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="subSpecies" species="chinensis" subSpecies="miyiensis">
<emphasis id="C6A7D36A387030702725F8ADFBA9F8FC" box="[791,1137,1877,1901]" italics="true" pageId="7" pageNumber="36">Sibynophis chinensis miyiensis</emphasis>
Zhao &amp; Kou, 1987
</taxonomicName>
(Sichuan and Yunnan provinces),
<taxonomicName id="33D374FB3870307025D6F881FC2AF800" authority="Yang &amp; Su, 1984" authorityName="Yang &amp; Su" authorityYear="1984" box="[484,1010,1913,1938]" class="Reptilia" family="Colubridae" genus="Elaphe" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="subSpecies" species="carinata" subSpecies="deqinensis">
<emphasis id="C6A7D36A3870307025D6F881FCC5F800" box="[484,797,1913,1937]" italics="true" pageId="7" pageNumber="36">Elaphe carinata deqinensis</emphasis>
Yang &amp; Su, 1984
</taxonomicName>
(Yunnan Province),
<taxonomicName id="33D374FB3870307020D6F881FDA6F824" authority="Zhao &amp; Jiang, 1981" authorityName="Zhao &amp; Jiang" authorityYear="1981" class="Reptilia" family="Colubridae" genus="Macropisthodon" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="subSpecies" species="rudis" subSpecies="multiprefrontalis">
<emphasis id="C6A7D36A3870307020D6F881FE4CF824" italics="true" pageId="7" pageNumber="36">Macropisthodon rudis multiprefrontalis</emphasis>
Zhao &amp; Jiang, 1981
</taxonomicName>
(Sichuan Province), and
<taxonomicName id="33D374FB387030702792F865FA44F824" authority="Zhao, 1995" authorityName="Zhao" authorityYear="1995" box="[928,1436,1949,1973]" class="Reptilia" family="Viperidae" genus="Ovophis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="36" phylum="Chordata" rank="subSpecies" species="monticola" subSpecies="zhaokentangi">
<emphasis id="C6A7D36A387030702792F865FACAF824" box="[928,1298,1949,1973]" italics="true" pageId="7" pageNumber="36">Ovophis monticola zhaokentangi</emphasis>
Zhao, 1995
</taxonomicName>
(Yunnan Province) (
<bibRefCitation id="9042728938703070254AF839FDF3F848" author="Luo" box="[376,555,1985,2009]" pageId="7" pageNumber="36" refString="Luo, J., Gao, H. Y., Liu, Y. M., Xu, L. &amp; Luo, Y. (2010) A revised checklist and distribution of Chinese snakes. Acta Herpetologica Sinica, 12, 67 - 91 [in Chinese]." type="journal article" year="2010">
Luo
<emphasis id="C6A7D36A38703070259DF83AFE38F848" box="[431,480,1985,2009]" italics="true" pageId="7" pageNumber="36">et al</emphasis>
. 2010
</bibRefCitation>
). Having in mind that several subspecies of Oriental snakes proved to be valid species in recent years (e.g.,
<bibRefCitation id="90427289387F307F25E9FF60FD32FF3E" author="Wuster" box="[475,746,151,175]" pageId="8" pageNumber="37" refString="Wuster, W. &amp; Thorpe, R. S. (1992) Asiatic cobras: population systematics of the Naja naja species complex (Serpentes: Elapidae) in India and Central Asia. Herpetologica, 48 (1), 69 - 85." type="journal article" year="1992">Wüster &amp; Thorpe, 1992</bibRefCitation>
;
<bibRefCitation id="90427289387F307F26C4FF6FFC31FF3E" author="Vogel" box="[758,1001,151,175]" pageId="8" pageNumber="37" refString="Vogel, G. &amp; David, P. (2007) On the taxonomy of the Xenochrophis piscator complex (Serpentes, Natricidae). In: M. Vences, J. Kohler, T. Ziegler &amp; W. Bohme (Eds.), Herpetologia Bonnensis II. Proceedings of the 13 th Congress of the Societas Europaea Herpetologica, Bonn, 241 - 246." type="book chapter" year="2007">Vogel &amp; David, 2007</bibRefCitation>
,
<bibRefCitation id="90427289387F307F27C6FF6FFADAFF3E" author="Vogel" box="[1012,1282,151,175]" pageId="8" pageNumber="37" refString="Vogel, G. &amp; Van Rooijen, J. (2011) Contributions to a Review of the Dendrelaphis pictus (Gmelin, 1789) Complex (Serpentes: Colubridae) - 3. The Indian Forms, with the Description of a New Species from the Western Ghats. Journal of Herpetology, 45 (1), in print." type="journal article" year="2011">
Vogel &amp; Rooijen, (
<quantity id="332BA29D387F307F20FBFF6FFADAFF3E" box="[1225,1282,151,175]" metricMagnitude="1" metricUnit="m" metricValue="5.10794" pageId="8" pageNumber="37" unit="in" value="2011.0">2011</quantity>
</bibRefCitation>
in print), it is expected that some of these may eventually receive full species status.
</paragraph>
<caption id="A0AC5FF0387F307F24A5FAECFA52F9C6" httpUri="https://zenodo.org/record/202520/files/figure.png" pageId="8" pageNumber="37" targetBox="[151,1436,267,1273]" targetPageId="8">
<paragraph id="F46C0F78387F307F24A5FAECFA52F9C6" blockId="8.[151,1437,1300,1623]" pageId="8" pageNumber="37">
<emphasis id="C6A7D36A387F307F24A5FAECFECAFABB" bold="true" box="[151,274,1300,1323]" pageId="8" pageNumber="37">FIGURE 5.</emphasis>
Map showing the distribution of
<taxonomicName id="33D374FB387F307F2656FAEDFCFBFABA" box="[612,803,1301,1323]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A387F307F2656FAEDFCFBFABA" box="[612,803,1301,1323]" italics="true" pageId="8" pageNumber="37">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387F307F271AFAEDFC57FABB" box="[808,911,1301,1322]" pageId="8" pageNumber="37" rank="species">
<emphasis id="C6A7D36A387F307F271AFAEDFC57FABB" bold="true" box="[808,911,1301,1322]" pageId="8" pageNumber="37">spec. nov.</emphasis>
</taxonomicNameLabel>
(red squares 1-2),
<taxonomicName id="33D374FB387F307F207FFAEDFADBFABA" box="[1101,1283,1301,1323]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387F307F207FFAEDFADBFABA" box="[1101,1283,1301,1323]" italics="true" pageId="8" pageNumber="37">Lycodon fasciatus</emphasis>
</taxonomicName>
(green squares 3-7),
<emphasis id="C6A7D36A387F307F24E6FACEFE0BFADD" box="[212,467,1334,1356]" italics="true" pageId="8" pageNumber="37">
<taxonomicName id="33D374FB387F307F24E9FACEFE14FADD" box="[219,460,1334,1356]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="yunnanensis">Dinodon yunnanensis</taxonomicName>
</emphasis>
(blue square 8), and
<taxonomicName id="33D374FB387F307F26F8FACEFC5EFADD" box="[714,902,1334,1356]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="synaptor">
<emphasis id="C6A7D36A387F307F26F8FACEFC5EFADD" box="[714,902,1334,1356]" italics="true" pageId="8" pageNumber="37">Lycodon synaptor</emphasis>
</taxonomicName>
(yellow square 9). 1. “Bapo”, now Dulongjiang Township,
<taxonomicName id="33D374FB387F307F2534FAAFFEB5FAFC" box="[262,365,1367,1389]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
County, Yunnan Province, Peoples Republic of China [Type locality of
<taxonomicName id="33D374FB387F307F207AFAAFFADFFAFC" box="[1096,1287,1367,1389]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A387F307F207AFAAFFADFFAFC" box="[1096,1287,1367,1389]" italics="true" pageId="8" pageNumber="37">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387F307F213FFAA0FAABFAFC" box="[1293,1395,1368,1389]" pageId="8" pageNumber="37" rank="species">
<emphasis id="C6A7D36A387F307F213FFAA0FAABFAFC" bold="true" box="[1293,1395,1368,1389]" pageId="8" pageNumber="37">spec. nov.</emphasis>
</taxonomicNameLabel>
]; 2. Xiaoheishan, Longjiang Township, Longling County, Yunnan Province, Peoples Republic of China; 3. “Ponsee”, now Bangxi, Yingjiang County, Yunnan Province, Peoples Republic of China [former type locality of
<taxonomicName id="33D374FB387F307F207EFA62FB27FA21" box="[1100,1279,1434,1456]" class="Squamata" family="Colubridae" genus="Ophites" higherTaxonomySource="CoL" kingdom="Animalia" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387F307F207EFA62FB27FA21" box="[1100,1279,1434,1456]" italics="true" pageId="8" pageNumber="37">Ophites fasciatus</emphasis>
</taxonomicName>
]; 4. “Mogok”, Mandalay Division, Myanmar [New type locality of
<taxonomicName id="33D374FB387F307F268DFA44FCA0FA43" box="[703,888,1468,1490]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387F307F268DFA44FCA0FA43" box="[703,888,1468,1490]" italics="true" pageId="8" pageNumber="37">Lycodon fasciatus</emphasis>
</taxonomicName>
]; 5. Tuantian, Tengchong County, Yunnan Province, Peoples Republic of China; 6. Husa, Longchuan County, Yunnan Province, Peoples Republic of China; 7. Ruili City, Yunnan Province, Peoples Republic of China; 8. “Yunnanfu”, now Kunming City, Yunnan Province, Peoples Republic of China [Type locality of
<taxonomicName id="33D374FB387F307F2538F9D8FE30F9A7" box="[266,488,1568,1590]" class="Reptilia" family="Colubridae" genus="Dinodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="C6A7D36A387F307F2538F9D8FE30F9A7" box="[266,488,1568,1590]" italics="true" pageId="8" pageNumber="37">Dinodon yunnanensis</emphasis>
</taxonomicName>
]; 9. “Tongchuan”, now Dongchuan District, Yunnan Province, Peoples Republic of China [Type locality of
<taxonomicName id="33D374FB387F307F2577F9B9FE23F9C6" box="[325,507,1601,1623]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="synaptor">
<emphasis id="C6A7D36A387F307F2577F9B9FE23F9C6" box="[325,507,1601,1623]" italics="true" pageId="8" pageNumber="37">Lycodon synaptor</emphasis>
</taxonomicName>
]. “For
<taxonomicName id="33D374FB387F307F2677F9B9FD25F9C6" box="[581,765,1601,1623]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="fasciatus">Lycodon fasciatus</taxonomicName>
only specimens from the
<taxonomicName id="33D374FB387F307F2030F9B9FBB1F9C6" box="[1026,1129,1601,1623]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">Gongshan</taxonomicName>
area are shown in the map.”
</paragraph>
</caption>
<paragraph id="F46C0F78387F307F24F7F979FC17F829" blockId="8.[151,1437,1664,1976]" pageId="8" pageNumber="37">
<collectingCountry id="8CC44FE8387F307F24F7F979FF3CF908" box="[197,228,1665,1689]" name="American Samoa" pageId="8" pageNumber="37">As</collectingCountry>
already mentioned (
<bibRefCitation id="90427289387F307F25FEF978FD1CF909" author="Vogel" box="[460,708,1664,1688]" pageId="8" pageNumber="37" refString="Vogel, G. &amp; David, P. (2010) A new species of the genus Lycodon (Boie, 1826) from Yunnan Province, China (Serpentes: Colubridae). Bonn zoological Bulletin, 57, 289 - 296." type="journal article" year="2010">Vogel &amp; David, 2010</bibRefCitation>
), there are still several lineages combined under the name
<emphasis id="C6A7D36A387F307F216FF97AFEEDF92C" italics="true" pageId="8" pageNumber="37">
<taxonomicName id="33D374FB387F307F216FF97AFEE8F92C" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="fasciatus">Lycodon fasciatus</taxonomicName>
.
</emphasis>
This fact explains the large ranges of some characters in
<taxonomicName id="33D374FB387F307F27F7F95DFB49F92C" box="[965,1169,1701,1725]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" sensu="lato" species="fasciatus">
<emphasis id="C6A7D36A387F307F27F7F95DFB49F92C" box="[965,1169,1701,1725]" italics="true" pageId="8" pageNumber="37">Lycodon fasciatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387F307F20ABF95EFAD6F92C" box="[1177,1294,1701,1725]" pageId="8" pageNumber="37" sensu="lato">sensu lato</taxonomicNameLabel>
. Thus, some characters of
<taxonomicName id="33D374FB387F307F2506F931FDD2F970" box="[308,522,1737,1761]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="gongshan" status="sp. nov.">
<emphasis id="C6A7D36A387F307F2506F931FDD2F970" box="[308,522,1737,1761]" italics="true" pageId="8" pageNumber="37">Lycodon gongshan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DD946E11387F307F2626F931FD52F970" box="[532,650,1737,1761]" pageId="8" pageNumber="37" rank="species">
<emphasis id="C6A7D36A387F307F2626F931FD52F970" bold="true" box="[532,650,1737,1761]" pageId="8" pageNumber="37">spec. nov.</emphasis>
</taxonomicNameLabel>
might currently appear less diagnostic than they actually are. More material, especially from remote Chinese areas, is needed for a better understanding of the
<taxonomicName id="33D374FB387F307F209BF916FAF3F894" box="[1193,1323,1773,1797]" class="Reptilia" family="Colubridae" genus="Lycodon" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="37" phylum="Chordata" rank="species" species="fasciatus">
<emphasis id="C6A7D36A387F307F209BF916FAF3F894" box="[1193,1323,1773,1797]" italics="true" pageId="8" pageNumber="37">L. fasciatus</emphasis>
</taxonomicName>
complex. This is another example of a supposedly widespread species that, upon closer examination, proves to be a species complex. A similar situation in other taxa would represent a greater underestimation of diversity in the ophidian fauna of the Oriental region than currently appreciated, a topic that requires further investigation. Centers of endemicity can be overlooked by inaccurate and/or imprecise taxonomies, which may lead to dire consequences in nature conservation, especially for taxa less well studied than vertebrates.
</paragraph>
</subSubSection>
</treatment>
</document>