425 lines
56 KiB
XML
425 lines
56 KiB
XML
<document id="BEDAE6F6B418F6A5615540AAC53FFCD5" ID-CLB-Dataset="30508" ID-DOI="10.11646/zootaxa.4394.3.9" ID-GBIF-Dataset="50cee940-aa9d-410f-9217-425658d1d587" ID-ISSN="1175-5326" ID-Zenodo-Dep="1199848" ID-ZooBank="AD1DEC6F-7027-4134-AD94-CCE20E82E1CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521176754760" checkinUser="plazi" docAuthor="Kompier, Tom & Karube, Haruki" docDate="2018" docId="7C658792FFA7AE2CFF39E35CA8EFFDF4" docLanguage="en" docName="zootaxa.4394.3.9.pdf" docOrigin="Zootaxa 4394 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlorogomphus canhvang Kompier & Karube 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="442" masterDocId="805CFFEAFFA6AE29FFAEE14BA972FFEC" masterDocTitle="Chlorogomphus canhvang sp. nov. from Central Vietnam (Odonata: Chlorogomphidae)" masterLastPageNumber="442" masterPageNumber="437" pageNumber="438" updateTime="1698531130700" updateUser="ExternalLinkService">
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<mods:mods id="17EAE01D1F28CAB8AC8223736925F32E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="B05058DA6DD6CACE2E617DC9065FB72F">
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<mods:title id="69598930AEF6D1A6E514DDA3DB3D4DDC">Chlorogomphus canhvang sp. nov. from Central Vietnam (Odonata: Chlorogomphidae)</mods:title>
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<mods:name id="EF2E798EC5A7A66ED5D1E938B6CDFD56" type="personal">
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<mods:roleTerm id="50E251712D50594AA09C4A2CFBFA7E26">Author</mods:roleTerm>
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<mods:namePart id="5D8C47D99C9472C4239072CA14CA14FB">Kompier, Tom</mods:namePart>
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<mods:name id="469B411E9DEC0686B657D1E2354AE556" type="personal">
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<mods:roleTerm id="2249EA8C1C3C22A46B4359DD65670E0A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5D9F2E0DE03EBDF6D5A63130BB2EA8C9">Karube, Haruki</mods:namePart>
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<mods:typeOfResource id="E79BE9172C05350A6586FB52E7F22D1B">text</mods:typeOfResource>
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<mods:title id="53BAC2A1BEF8E03746D10E4937F6F401">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D8157258701722740611F255901EAE8B">
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<mods:date id="F9766C4F47F480159C10C6613B0ADE9C">2018</mods:date>
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<mods:number id="0E69A83A9F5405DE6B91E6A6F40A5252">2018-03-15</mods:number>
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<mods:number id="487CBA2270AB0309D2CE368976C41FD4">4394</mods:number>
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</mods:detail>
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<mods:detail id="A812224C6B8BE87DE4C44BC20F2C458F" type="issue">
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<mods:number id="7F4907AF6124D0D520B4EC79310AF7B8">3</mods:number>
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<mods:classification id="96AFD3691A3BCFC3D7A17F1597A268C4">journal article</mods:classification>
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<mods:identifier id="2F0373323376E9E5776D1CD977BCD14A" type="CLB-Dataset">30508</mods:identifier>
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<mods:identifier id="401F46191961C0299DA296978A193B62" type="DOI">10.11646/zootaxa.4394.3.9</mods:identifier>
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<mods:identifier id="501FA9F4903EEF848185EF8269EE82ED" type="GBIF-Dataset">50cee940-aa9d-410f-9217-425658d1d587</mods:identifier>
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<mods:identifier id="3C5363C7B82476472FA2EB6EFAEB83F6" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C1F96AFBFD8E86D330F14D827B07DFEF" type="Zenodo-Dep">1199848</mods:identifier>
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</mods:mods>
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<treatment id="7C658792FFA7AE2CFF39E35CA8EFFDF4" ID-DOI="http://doi.org/10.5281/zenodo.5976865" ID-GBIF-Taxon="142289408" ID-Zenodo-Dep="5976865" LSID="urn:lsid:plazi:treatment:7C658792FFA7AE2CFF39E35CA8EFFDF4" httpUri="http://treatment.plazi.org/id/7C658792FFA7AE2CFF39E35CA8EFFDF4" lastPageId="5" lastPageNumber="442" pageId="1" pageNumber="438">
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<subSubSection id="BCD6650FFFA7AE28FF39E35CA8D3FDBF" pageId="1" pageNumber="438" type="nomenclature">
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<paragraph id="F4733684FFA7AE28FF39E35CAB4FFDDD" blockId="1.[151,573,535,595]" box="[151,573,535,562]" pageId="1" pageNumber="438">
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<heading id="AF3B81E8FFA7AE28FF39E35CAB4FFDDD" bold="true" box="[151,573,535,562]" fontSize="11" level="1" pageId="1" pageNumber="438" reason="1">
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<emphasis id="C6B8EA96FFA7AE28FF39E35CAB4FFDDD" bold="true" box="[151,573,535,562]" pageId="1" pageNumber="438">
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<taxonomicName id="33CC4D07FFA7AE28FF39E35CA8ABFDDD" ID-CoL="9X2B2" authority="Kompier & Karube, 2018" authorityName="Kompier & Karube" authorityYear="2018" box="[151,473,535,561]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="canhvang" status="sp. nov.">
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<emphasis id="C6B8EA96FFA7AE28FF39E35CA8ABFDDD" bold="true" box="[151,473,535,561]" italics="true" pageId="1" pageNumber="438">Chlorogomphus canhvang</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="DD8B57EDFFA7AE28FE4EE353AB4FFDDD" box="[480,573,535,562]" pageId="1" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="F4733684FFA7AE28FF39E371A8D3FDBF" blockId="1.[151,573,535,595]" box="[151,417,570,595]" pageId="1" pageNumber="438">
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(
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<figureCitation id="6CF72A01FFA7AE28FF31E371A866FDBF" box="[159,276,570,595]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="1" pageNumber="438">Figs. 1d–f</figureCitation>
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,
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<figureCitation id="6CF72A01FFA7AE28FE8EE370A82BFDBF" box="[288,345,571,595]" captionStart="FIGURE 2" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[159,1405,214,1851]" captionTargetId="figure@3.[150,1436,213,1885]" captionTargetPageId="3" captionText="FIGURE 2. Comparison of anal appendages [a–b, e–f] and vesica spermalis [c–d, g–h] of Chlorogomphus auratus (Tam Dao NP, Vinh Phuc Prov.) [a–d] and C. canhvang, holotype [e–h]. (a, e) dorsal view; (b, f) lateral view; (c, g) lateral view; (d, h) ventral view of terminal segment." httpUri="https://zenodo.org/record/1199852/files/figure.png" pageId="1" pageNumber="438">2e–g</figureCitation>
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,
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<figureCitation id="6CF72A01FFA7AE28FECAE371A8EBFDBF" box="[356,409,570,595]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="1" pageNumber="438">3b–f</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="BCD6650FFFA7AE28FF39E3C9AB97FC9E" pageId="1" pageNumber="438" type="materials_examined">
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||
<paragraph id="F4733684FFA7AE28FF39E3C9AC04FD53" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
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<bibRef id="2D6CA272FFA7AE28FF39E3C9AC04FD53" pageId="1" pageNumber="438" refId="ref784" type="book chapter">
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<materialsCitation id="44A43CD9FFA7AE28FF39E3C9A9CFFD53" ID-GBIF-Occurrence="1835647450" collectingDate="2016-05-06" collectorName="Along QL" latitude="17.488" location="Quang Binh Province" longLatPrecision="76" longitude="106.302" pageId="1" pageNumber="438" specimenCount="1" specimenCount-male="1" stateProvince="Quang Binh Province" typeStatus="holotype">
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<title id="86C8DDB2FFA7AE28FF39E3C9AB6EFD76" box="[151,540,642,667]" jopScore="0.16837364" pageId="1" pageNumber="438">
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||
<emphasis id="C6B8EA96FFA7AE28FF39E3C9A8F6FD77" bold="true" box="[151,388,642,667]" pageId="1" pageNumber="438">Materials examined</emphasis>
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.
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<typeStatus id="2B778826FFA7AE28FE3CE3C9A88DFD77" box="[402,511,642,667]" pageId="1" pageNumber="438" type="holotype">
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<emphasis id="C6B8EA96FFA7AE28FE3CE3C9A88DFD77" bold="true" box="[402,511,642,667]" pageId="1" pageNumber="438">Holotype</emphasis>
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||
</typeStatus>
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||
<specimenCount id="E2CAFD0DFFA7AE28FDABE3C8AB6EFD76" box="[517,540,643,666]" pageId="1" pageNumber="438" type="male">♂</specimenCount>
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||
</title>
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||
,
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<collectorName id="59395352FFA7AE28FD86E3C9ABECFD77" box="[552,670,642,667]" pageId="1" pageNumber="438">
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<author id="2C71DE61FFA7AE28FD86E3C9ABECFD77" box="[552,670,642,667]" fns="I" npo="LF" pageId="1" pageNumber="438">Along QL</author>
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</collectorName>
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15,
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<collectingRegion id="3608F866FFA7AE28FD6AE3C8AAC5FD77" box="[708,951,642,667]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="438">
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Quang
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<collectingRegion id="3608F866FFA7AE28FCB8E3C9AAC5FD77" box="[790,951,642,667]" inferredBySuffix="true" pageId="1" pageNumber="438">Binh Province</collectingRegion>
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</collectingRegion>
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(appr.
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<geoCoordinate id="91F85043FFA7AE28FBABE3C8AD1DFD76" box="[1029,1135,643,667]" direction="north" orientation="latitude" pageId="1" pageNumber="438" precision="55" value="17.488">17.488 N</geoCoordinate>
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,
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<geoCoordinate id="91F85043FFA7AE28FBD4E3C9AD82FD77" box="[1146,1264,642,667]" direction="east" orientation="longitude" pageId="1" pageNumber="438" precision="55" value="106.302">106.302 E</geoCoordinate>
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),
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<collectingDate id="9036E9ACFFA7AE28FAAAE3C9AC19FD77" box="[1284,1387,642,667]" pageId="1" pageNumber="438" value="2016-05-06">6.V.2016</collectingDate>
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, TK leg
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</materialsCitation>
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.
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<materialsCitation id="44A43CD9FFA7AE28FF69E3EDAA46FD53" ID-GBIF-Occurrence="1835647507" box="[199,820,678,703]" collectingDate="2017-05-17" collectorName="Along QL" latitude="17.488" location="Quang Binh Province" longLatPrecision="76" longitude="106.302" pageId="1" pageNumber="438" specimenCount="1" specimenCount-female="1" stateProvince="Quang Binh Province" typeStatus="paratype">
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||
<typeStatus id="2B778826FFA7AE28FF69E3EDA84DFD53" box="[199,319,678,703]" pageId="1" pageNumber="438" type="paratype">
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<emphasis id="C6B8EA96FFA7AE28FF69E3EDA84DFD53" bold="true" box="[199,319,678,703]" pageId="1" pageNumber="438">Paratypes</emphasis>
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</typeStatus>
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<specimenCount id="E2CAFD0DFFA7AE28FEE8E3ECA802FD52" box="[326,368,679,703]" pageId="1" pageNumber="438" type="female">1 ♀</specimenCount>
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, same location and collector,
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<collectingDate id="9036E9ACFFA7AE28FD10E3ECAA46FD53" box="[702,820,679,703]" pageId="1" pageNumber="438" value="2017-05-17">
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17.V.
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<year id="806477D7FFA7AE28FD58E3ECAA46FD53" box="[758,820,679,703]" pageId="1" pageNumber="438">2017</year>
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||
</collectingDate>
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</materialsCitation>
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||
;
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<pagination id="CD05C870FFA7AE28FC91E3ECAA3FFD53" box="[831,845,679,703]" pageId="1" pageNumber="438" type="pagination">1</pagination>
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♂, same date, location and collector as holotype.
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</bibRef>
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</paragraph>
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<paragraph id="F4733684FFA7AE28FF69E381A8E6FCC7" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
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<materialsCitation id="44A43CD9FFA7AE28FF69E381AD80FD0E" ID-GBIF-Occurrence="1835647505" box="[199,1266,714,739]" country="Japan" location="Kanagawa" pageId="1" pageNumber="438" specimenCount="1" stateProvince="Kanagawa" typeStatus="holotype">
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The
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<typeStatus id="2B778826FFA7AE28FF57E381A829FD0F" box="[249,347,714,739]" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
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will be deposited at the
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<collectingRegion id="3608F866FFA7AE28FDC3E380AB91FD0F" box="[621,739,715,739]" country="Japan" name="Kanagawa" pageId="1" pageNumber="438">Kanagawa</collectingRegion>
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Prefectural Museum of Natural History,
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<collectingCountry id="8CDB7614FFA7AE28FB01E380AD80FD0E" box="[1199,1266,715,738]" name="Japan" pageId="1" pageNumber="438">Japan</collectingCountry>
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</materialsCitation>
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.
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<materialsCitation id="44A43CD9FFA7AE28FB53E380A8E2FCC7" ID-GBIF-Occurrence="1835647506" collectionCode="RMNH" collectorName="Rijksmuseum van Natuurlijke Historie & Leiden" country="Netherlands" location="Center" pageId="1" pageNumber="438" specimenCount="1" stateProvince="Center" typeStatus="paratype">
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<typeStatus id="2B778826FFA7AE28FB53E380AC18FD0F" box="[1277,1386,715,739]" pageId="1" pageNumber="438" type="paratype">Paratypes</typeStatus>
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will be deposited at Naturalis Biodiversity
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<collectingRegion id="3608F866FFA7AE28FDCBE3A4ABC2FCEB" box="[613,688,751,775]" country="France" name="Centre" pageId="1" pageNumber="438">Center</collectingRegion>
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||
(formerly
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<collectorName id="59395352FFA7AE28FC9BE3A5AD8DFCEB" box="[821,1279,750,775]" pageId="1" pageNumber="438">Rijksmuseum van Natuurlijke Historie</collectorName>
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||
),
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<collectorName id="59395352FFA7AE28FAB6E3A5AC1AFCEB" box="[1304,1384,750,775]" pageId="1" pageNumber="438">Leiden</collectorName>
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||
,
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<collectingCountry id="8CDB7614FFA7AE28FAD4E3A5A86CFCC7" name="Netherlands" pageId="1" pageNumber="438">the Netherlands</collectingCountry>
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||
(
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<collectionCode id="92DDAE41FFA7AE28FE83E258A8FAFCC7" box="[301,392,787,811]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="438">RMNH</collectionCode>
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||
)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="F4733684FFA7AE28FF69E27DAB97FC9E" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<materialsCitation id="44A43CD9FFA7AE28FF69E27DAB97FC9E" ID-GBIF-Occurrence="1835647504" collectingDate="2016-05-04" latitude="16.064" location="Along National Road" longLatPrecision="76" longitude="107.485" pageId="1" pageNumber="438" specimenCount="1" specimenCount-female="1" stateProvince="Thua Thien-Hue Province">
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Additional observations:
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<location id="F113605FFFA7AE28FE4DE27DABA1FCA3" LSID="urn:lsid:plazi:treatment:7C658792FFA7AE2CFF39E35CA8EFFDF4:F113605FFFA7AE28FE4DE27DABA1FCA3" box="[483,723,822,847]" latitude="16.064" longLatPrecision="76" longitude="107.485" name="Along National Road" pageId="1" pageNumber="438" stateProvince="Thua Thien-Hue Province">Along National Road</location>
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14,
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<collectingRegion id="3608F866FFA7AE28FCABE27DAD4FFCA3" box="[773,1085,822,847]" country="Vietnam" name="Thua Thien-Hue" pageId="1" pageNumber="438">
|
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Thua
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<collectingRegion id="3608F866FFA7AE28FCE8E27DAD4FFCA3" box="[838,1085,822,847]" inferredBySuffix="true" pageId="1" pageNumber="438">Thien—Hue Province</collectingRegion>
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</collectingRegion>
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(appr.
|
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<geoCoordinate id="91F85043FFA7AE28FB22E27DAD85FCA2" box="[1164,1271,822,847]" direction="north" orientation="latitude" pageId="1" pageNumber="438" precision="55" value="16.064">16.064 N</geoCoordinate>
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,
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<geoCoordinate id="91F85043FFA7AE28FAADE27DAC0BFCA3" box="[1283,1401,822,847]" direction="east" orientation="longitude" pageId="1" pageNumber="438" precision="55" value="107.485">107.485 E</geoCoordinate>
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||
),
|
||
<specimenCount id="E2CAFD0DFFA7AE28FA20E27CA9DEFC9E" pageId="1" pageNumber="438" type="female">1 ♀</specimenCount>
|
||
caught, photographed and released,
|
||
<collectingDate id="9036E9ACFFA7AE28FDE8E211ABDFFC9F" box="[582,685,858,883]" pageId="1" pageNumber="438" value="2016-05-04">
|
||
<date id="80721044FFA7AE28FDE8E211ABDFFC9F" box="[582,685,858,883]" pageId="1" pageNumber="438" value="2016-05-04">4.V.2016</date>
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</collectingDate>
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, TK.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="BCD6650FFFA7AE28FF69E235AAB7FC33" pageId="1" pageNumber="438" type="etymology">
|
||
<paragraph id="F4733684FFA7AE28FF69E235AAB7FC33" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
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||
<emphasis id="C6B8EA96FFA7AE28FF69E235A834FC7B" bold="true" box="[199,326,894,919]" pageId="1" pageNumber="438">Etymology</emphasis>
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. The specific name
|
||
<taxonomicName id="33CC4D07FFA7AE28FD81E235ABEEFC7B" authorityName="Kompier & Karube" authorityYear="2018" box="[559,668,894,919]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA7AE28FD81E235ABEEFC7B" box="[559,668,894,919]" italics="true" pageId="1" pageNumber="438">canhvang</emphasis>
|
||
</taxonomicName>
|
||
means “golden-winged” in Vietnamese (cánh vàng) in reference to the golden-winged appearance of the female and as a sister taxon to
|
||
<taxonomicName id="33CC4D07FFA7AE28FC08E2E9ADCAFC57" authorityName="Martin" authorityYear="1910" box="[934,1208,930,955]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA7AE28FC08E2E9ADCAFC57" box="[934,1208,930,955]" italics="true" pageId="1" pageNumber="438">Chlorogomphus auratus</emphasis>
|
||
</taxonomicName>
|
||
, the name of which also refers to the golden wings of the female. An adjective in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BCD6650FFFA7AE2DFF69E2A1A87DFE3C" lastPageId="4" lastPageNumber="441" pageId="1" pageNumber="438" type="description">
|
||
<paragraph id="F4733684FFA7AE28FF69E2A1ADF0FB37" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E2A1A8ADFBEF" bold="true" box="[199,479,1002,1027]" pageId="1" pageNumber="438">
|
||
Description of
|
||
<typeStatus id="2B778826FFA7AE28FED7E2A1A8ADFBEF" box="[377,479,1002,1027]" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="C6B8EA96FFA7AE28FE41E2A1AB5EFBEF" box="[495,556,1002,1027]" italics="true" pageId="1" pageNumber="438">Head</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FD93E2A1ABFFFBEF" box="[573,653,1002,1027]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="1" pageNumber="438">Fig. 3f</figureCitation>
|
||
). Labium yellow, labial palps golden-brown. Labrum black with two small pale yellow spots in V-shape centrally below dorsal margin. Anteclypeus black with lower and upper margin bordered by pale yellow, narrowly connected over center. Postclypeus pale yellow with two small brown spots. Frons black, covered in black setae, in lateral view not noticeably protruding, in dorsal view anterior margin only slightly rounded covered in black setae; postfrons with thin greenish yellow line along anterior margin. Vertex conically raised, posterior surface slightly concave. Antennae black. Occiput black, forming a triangular pyramid, apex close to posterior margin, covered in long black setae. Compound eyes bright green.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA7AE28FF69E5ADA837FA63" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E5ADA86AFB13" box="[199,280,1254,1279]" italics="true" pageId="1" pageNumber="438">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FE88E5ADA80BFB13" box="[294,377,1254,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="1" pageNumber="438">Fig. 1d</figureCitation>
|
||
). Prothorax matte black, anterior margin pale yellow and with small pale yellow area on lateral margins and center of posterior margin. Synthorax black with yellow markings as follows: A thin greenish-yellow dorsal stripe and a thin yellow antehumeral stripe on both sides, and a broad yellow band over the spiracle and covering most of the metepisternum. Ventral margin of metepimeron narrowly yellow. Coxae, legs, spines and claws all black.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA7AE28FF69E4D7AADEFA17" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E4D7A878FA5F" box="[199,266,1436,1459]" italics="true" pageId="1" pageNumber="438">Wings</emphasis>
|
||
. Hyaline with black veins and apices edged narrowly dark brown. Nodal index 18–26: 28–
|
||
<quantity id="33349B61FFA7AE28FAAFE4D0AC45FA5F" box="[1281,1335,1434,1459]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="1" pageNumber="438" unit="in" value="18.0">18 in</quantity>
|
||
FW, 20– 23:
|
||
<quantity id="33349B61FFA7AE28FF19E4F4A86DFA3B" box="[183,287,1470,1495]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="1" pageNumber="438" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">21–22 in</quantity>
|
||
HW. Triangles 3-celled in FW and 3–4 celled in HW. Anal loop 18–19 celled, anal triangle 3-celled. Membranule grey. Pterostigma black, covering 3.5–4 underlying cells.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA7AE28FF69E74DAAA4F943" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E74DA843F9F3" box="[199,305,1542,1567]" italics="true" pageId="1" pageNumber="438">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FEEEE74DA8E2F9F3" box="[320,400,1542,1567]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="1" pageNumber="438">Fig. 1d</figureCitation>
|
||
). S1 black, but ventral third yellow; S2 black with broad yellow band laterally, from anteroventral corner over auricle towards posterior margin, which is narrowly black. S3 black with large anterolateral yellow markings that just connect to each other anteriorly over dorsal carina. S4–7 black with paired antero-dorsal spots, small on S4–6, but larger on S7. S8–10 black. The dorsal side of S10 curving downward in between the cerci to form a small nipple-like protrusion, not visible in lateral view (
|
||
<figureCitation id="6CF72A01FFA7AE28FCD8E7DDAABBF943" box="[886,969,1686,1711]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="1" pageNumber="438">Fig. 3d</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA7AE28FF69E7F1AA6AF8F7" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E7F1A8DCF93F" box="[199,430,1722,1747]" italics="true" pageId="1" pageNumber="438">Secondary genitalia</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FE6CE7F1AB6BF93F" box="[450,537,1722,1747]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="1" pageNumber="438">Fig. 3c</figureCitation>
|
||
). Black. Anterior lamina with undulating anterior margin and small inward directed apical hook, keeled posterior margin of posterior hamule in lateral view about 1.8 times as long as anterior lamina, thin, with apical hook weekly directed anteriorly.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA7AE28FF69E66CA895F86B" blockId="1.[151,1437,642,2035]" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E66CA8F5F8D3" box="[199,391,1830,1855]" italics="true" pageId="1" pageNumber="438">Vesica spermalis</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FE35E66DAB62F8D3" box="[411,528,1830,1855]" captionStart="FIGURE 2" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[159,1405,214,1851]" captionTargetId="figure@3.[150,1436,213,1885]" captionTargetPageId="3" captionText="FIGURE 2. Comparison of anal appendages [a–b, e–f] and vesica spermalis [c–d, g–h] of Chlorogomphus auratus (Tam Dao NP, Vinh Phuc Prov.) [a–d] and C. canhvang, holotype [e–h]. (a, e) dorsal view; (b, f) lateral view; (c, g) lateral view; (d, h) ventral view of terminal segment." httpUri="https://zenodo.org/record/1199852/files/figure.png" pageId="1" pageNumber="438">
|
||
Fig.
|
||
<quantity id="33349B61FFA7AE28FE7AE66DA882F8D3" box="[468,496,1830,1855]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="1" pageNumber="438" unit="g" value="2.0">2g</quantity>
|
||
–h
|
||
</figureCitation>
|
||
). Similar to that of
|
||
<taxonomicName id="33CC4D07FFA7AE28FCAFE66DAD67F8D2" authorityName="Martin" authorityYear="1910" box="[769,1045,1830,1855]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="438" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA7AE28FCAFE66DAD67F8D2" box="[769,1045,1830,1855]" italics="true" pageId="1" pageNumber="438">Chlorogomphus auratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FB84E66DADEFF8D3" box="[1066,1181,1830,1855]" captionStart="FIGURE 2" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[159,1405,214,1851]" captionTargetId="figure@3.[150,1436,213,1885]" captionTargetPageId="3" captionText="FIGURE 2. Comparison of anal appendages [a–b, e–f] and vesica spermalis [c–d, g–h] of Chlorogomphus auratus (Tam Dao NP, Vinh Phuc Prov.) [a–d] and C. canhvang, holotype [e–h]. (a, e) dorsal view; (b, f) lateral view; (c, g) lateral view; (d, h) ventral view of terminal segment." httpUri="https://zenodo.org/record/1199852/files/figure.png" pageId="1" pageNumber="438">Fig. 2c–d</figureCitation>
|
||
), but second segment broader, with stout spine, the last segment somewhat more robust, its ventral plate somewhat more slender and more deeply notched apically.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA7AE2BFF69E6D9ABDFFEF0" blockId="1.[151,1437,642,2035]" lastBlockId="2.[151,1436,151,717]" lastPageId="2" lastPageNumber="439" pageId="1" pageNumber="438">
|
||
<emphasis id="C6B8EA96FFA7AE28FF69E6D9A8FEF847" box="[199,396,1938,1963]" italics="true" pageId="1" pageNumber="438">Anal appendages</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA7AE28FE31E6D9AB7DF847" box="[415,527,1938,1963]" captionStart="FIGURE 2" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[159,1405,214,1851]" captionTargetId="figure@3.[150,1436,213,1885]" captionTargetPageId="3" captionText="FIGURE 2. Comparison of anal appendages [a–b, e–f] and vesica spermalis [c–d, g–h] of Chlorogomphus auratus (Tam Dao NP, Vinh Phuc Prov.) [a–d] and C. canhvang, holotype [e–h]. (a, e) dorsal view; (b, f) lateral view; (c, g) lateral view; (d, h) ventral view of terminal segment." httpUri="https://zenodo.org/record/1199852/files/figure.png" pageId="1" pageNumber="438">Fig. 2e–f</figureCitation>
|
||
). Black. Cerci about as long as S9, in lateral view broad at base, their dorsal margin almost straight, only slightly curving apically, and with a small blunt hump directed dorsad at their anterior end, their ventral margin more strongly curved, concave, their apex blunt. In dorsal view cerci widely separated, slightly diverging, subapically bifurcate, the outer branch shorter and directed somewhat ventrad, the inner branch apically slightly hooked laterally. The epiproct deeply incised, U-shaped in dorsal view, its branches virtually parallel-sided, minimally longer than cerci, in lateral view thickest in anterior half, curved upward and ending in a double tooth on the dorsal margin of their apex.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA4AE2BFF69E063AAC5FEAC" blockId="2.[151,1436,151,717]" box="[199,951,295,320]" pageId="2" pageNumber="439">Measurements (in mm). HW 55.6; abdomen incl. appendages 59.6.</paragraph>
|
||
<paragraph id="F4733684FFA4AE2BFF69E007AA81FE41" blockId="2.[151,1436,151,717]" pageId="2" pageNumber="439">
|
||
<emphasis id="C6B8EA96FFA4AE2BFF69E007A8DBFE89" bold="true" box="[199,425,332,357]" pageId="2" pageNumber="439">Variation in males</emphasis>
|
||
.
|
||
<typeStatus id="2B778826FFA4AE2BFE12E006AB6CFE89" box="[444,542,333,357]" pageId="2" pageNumber="439" type="paratype">Paratype</typeStatus>
|
||
very similarly marked as
|
||
<typeStatus id="2B778826FFA4AE2BFCF8E007AAC9FE89" box="[854,955,332,357]" pageId="2" pageNumber="439" type="holotype">holotype</typeStatus>
|
||
, but somewhat smaller: HW
|
||
<quantity id="33349B61FFA4AE2BFABBE007AC1BFE88" box="[1301,1385,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="439" unit="mm" value="52.0">52 mm</quantity>
|
||
and abdomen incl. appendages
|
||
<quantity id="33349B61FFA4AE2BFE68E024AB6AFE6B" box="[454,536,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="439" unit="mm" value="52.0">52 mm</quantity>
|
||
. Nodal index 16–29:28–
|
||
<quantity id="33349B61FFA4AE2BFC82E024AA16FE64" box="[812,868,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="2" pageNumber="439" unit="in" value="16.0">16 in</quantity>
|
||
FW, 20–23:23–
|
||
<quantity id="33349B61FFA4AE2BFBB4E03BAD20FE64" box="[1050,1106,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="2" pageNumber="439" unit="in" value="19.0">19 in</quantity>
|
||
HW. Triangles 4-celled in all wings. Anal loop 16–18 celled. Pterostigma covering 4–4.5 underlying cells.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA4AE2BFF69E0FCA805FE19" blockId="2.[151,1436,151,717]" pageId="2" pageNumber="439">
|
||
<emphasis id="C6B8EA96FFA4AE2BFF69E0FCAB3EFE3C" bold="true" box="[199,588,439,464]" pageId="2" pageNumber="439">
|
||
Description of female
|
||
<typeStatus id="2B778826FFA4AE2BFE4FE0FCAB3EFE3C" box="[481,588,439,464]" pageId="2" pageNumber="439" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="C6B8EA96FFA4AE2BFDCFE0FCABECFE3C" box="[609,670,439,464]" italics="true" pageId="2" pageNumber="439">Head</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA4AE2BFD1AE0FCAA5DFE3C" box="[692,815,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="2" pageNumber="439">Fig. 3b, e</figureCitation>
|
||
). As male, but vertex larger, postero-dorsal surface distinctly V-shaped.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA4AE2BFF69E0B4ABD3FDD1" blockId="2.[151,1436,151,717]" pageId="2" pageNumber="439">
|
||
<emphasis id="C6B8EA96FFA4AE2BFF69E0B4A86AFDF4" box="[199,280,511,536]" italics="true" pageId="2" pageNumber="439">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA4AE2BFE86E0B4A80AFDF4" box="[296,376,511,536]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="2" pageNumber="439">Fig. 1e</figureCitation>
|
||
). Prothorax as male but yellow spots on center of posterior margin larger. Synthorax as male. Coxae brown, legs, spines and claws all black.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA4AE2BFF69E302A820FD21" blockId="2.[151,1436,151,717]" pageId="2" pageNumber="439">
|
||
<emphasis id="C6B8EA96FFA4AE2BFF69E302A879FD8C" box="[199,267,585,608]" italics="true" pageId="2" pageNumber="439">Wings</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA4AE2BFEB4E30CA818FD8C" box="[282,362,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="2" pageNumber="439">Fig. 1e</figureCitation>
|
||
). Tinted golden, only minimally on antenodal part, strongest on postnodal part, with black veins and apices edged broadly dark brown, nodal index 18–28:30–
|
||
<quantity id="33349B61FFA4AE2BFCE1E327AAFAFD69" box="[847,904,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="2" pageNumber="439" unit="in" value="18.0">18 in</quantity>
|
||
FW, 21–24:23–
|
||
<quantity id="33349B61FFA4AE2BFB93E327AD04FD69" box="[1085,1142,620,645]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="2" pageNumber="439" unit="in" value="21.0">21 in</quantity>
|
||
HW. Triangles 5–6 celled in all wings. Anal loop 25–26 celled, anal triangle 3-celled. Membranule grey. Pterostigma black, covering 4–4.5 underlying cells.
|
||
</paragraph>
|
||
<caption id="A0B3660CFFA4AE2BFF39E6DAAAEFF808" httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="2" pageNumber="439" startId="2.[151,250,1937,1959]" targetBox="[151,1436,761,1912]" targetPageId="2">
|
||
<paragraph id="F4733684FFA4AE2BFF39E6DAAAEFF808" blockId="2.[151,1436,1937,2020]" pageId="2" pageNumber="439">
|
||
<emphasis id="C6B8EA96FFA4AE2BFF39E6DAA861F84A" bold="true" box="[151,275,1937,1959]" pageId="2" pageNumber="439">FIGURE 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="33CC4D07FFA4AE2BFE24E6DAAB0DF84B" authorityName="Martin" authorityYear="1910" box="[394,639,1937,1959]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="439" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA4AE2BFE24E6DAAB0DF84B" box="[394,639,1937,1959]" italics="true" pageId="2" pageNumber="439">Chlorogomphus auratus</emphasis>
|
||
</taxonomicName>
|
||
[a–c] and
|
||
<taxonomicName id="33CC4D07FFA4AE2BFD42E6D9AA1EF84B" authorityName="Kompier & Karube" authorityYear="2018" box="[748,876,1937,1959]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="439" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA4AE2BFD42E6D9AA1EF84B" box="[748,876,1937,1959]" italics="true" pageId="2" pageNumber="439">C. canhvang</emphasis>
|
||
</taxonomicName>
|
||
[d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A0B3660CFFA5AE2AFF39E6DAA89FF808" httpUri="https://zenodo.org/record/1199852/files/figure.png" pageId="3" pageNumber="440" startId="3.[151,250,1937,1959]" targetBox="[159,1405,214,1851]" targetPageId="3">
|
||
<paragraph id="F4733684FFA5AE2AFF39E6DAA89FF808" blockId="3.[151,1436,1937,2020]" pageId="3" pageNumber="440">
|
||
<emphasis id="C6B8EA96FFA5AE2AFF39E6DAA861F84A" bold="true" box="[151,275,1937,1959]" pageId="3" pageNumber="440">FIGURE 2.</emphasis>
|
||
Comparison of anal appendages [a–b, e–f] and vesica spermalis [c–d, g–h] of
|
||
<taxonomicName id="33CC4D07FFA5AE2AFB96E6DAAC5CF84B" authorityName="Martin" authorityYear="1910" box="[1080,1326,1937,1959]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA5AE2AFB96E6DAAC5CF84B" box="[1080,1326,1937,1959]" italics="true" pageId="3" pageNumber="440">Chlorogomphus auratus</emphasis>
|
||
</taxonomicName>
|
||
(Tam Dao NP, Vinh Phuc Prov.) [a–d] and
|
||
<taxonomicName id="33CC4D07FFA5AE2AFE4AE6FBAB17F829" authorityName="Kompier & Karube" authorityYear="2018" box="[484,613,1967,1989]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="440" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA5AE2AFE4AE6FBAB17F829" box="[484,613,1967,1989]" italics="true" pageId="3" pageNumber="440">C. canhvang</emphasis>
|
||
</taxonomicName>
|
||
, holotype [e–h]. (a, e) dorsal view; (b, f) lateral view; (c, g) lateral view; (d, h) ventral view of terminal segment.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F4733684FFA2AE2DFF69E1DCAAE2FEF0" blockId="4.[151,1437,151,464]" pageId="4" pageNumber="441">
|
||
<emphasis id="C6B8EA96FFA2AE2DFF69E1DCA843FF5C" box="[199,305,151,176]" italics="true" pageId="4" pageNumber="441">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="6CF72A01FFA2AE2DFEEEE1DCA8FDFF5C" box="[320,399,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="4" pageNumber="441">Fig. 1e</figureCitation>
|
||
). Black with yellow markings as follows: S1 with small dorsal spot and yellow ventral half; S2 with broad band laterally, extending from antero-ventral corner over auricle towards posterior margin, which is narrowly black, and large antero-dorsal spot. S3 with anterior half almost completely yellow, but for ventral margin. S4–7 with partially fused antero-dorsal spots, largest on S7.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA2AE2DFF69E063A829FEAC" blockId="4.[151,1437,151,464]" box="[199,347,295,320]" pageId="4" pageNumber="441">
|
||
<emphasis id="C6B8EA96FFA2AE2DFF69E063A876FED3" box="[199,260,296,319]" italics="true" pageId="4" pageNumber="441">Cerci</emphasis>
|
||
. Black.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA2AE2DFF69E006AAF0FE89" blockId="4.[151,1437,151,464]" box="[199,898,332,357]" pageId="4" pageNumber="441">Measurements (in mm). Hindwing 53; abdomen incl. cerci 55.</paragraph>
|
||
<paragraph id="F4733684FFA2AE2DFF69E024A87DFE3C" blockId="4.[151,1437,151,464]" pageId="4" pageNumber="441">
|
||
<emphasis id="C6B8EA96FFA2AE2DFF69E024A8C4FE64" bold="true" box="[199,438,367,392]" pageId="4" pageNumber="441">Variation in females</emphasis>
|
||
. Female photographed from
|
||
<collectingRegion id="3608F866FFA2AE2DFD54E024AAB4FE64" box="[762,966,367,392]" country="Vietnam" name="Thua Thien-Hue" pageId="4" pageNumber="441">Thua Thien—Hue</collectingRegion>
|
||
Prov. (
|
||
<figureCitation id="6CF72A01FFA2AE2DFBBBE024AD12FE64" box="[1045,1120,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="4" pageNumber="441">Fig. 1f</figureCitation>
|
||
) with 31 Ax in left FW. The golden colour of the wings much more intense, also on antenodal part of wings, with dark spot on nodus and at wing base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A0B3660CFFA2AE2DFF39E7BAADEDF8A8" httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="4" pageNumber="441" startId="4.[151,250,1777,1799]" targetBox="[170,1422,531,1765]" targetPageId="4">
|
||
<paragraph id="F4733684FFA2AE2DFF39E7BAADEDF8A8" blockId="4.[151,1436,1777,1860]" pageId="4" pageNumber="441">
|
||
<emphasis id="C6B8EA96FFA2AE2DFF39E7BAA865F8EA" bold="true" box="[151,279,1777,1799]" pageId="4" pageNumber="441">FIGURE 3.</emphasis>
|
||
Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posterior margin of S10 [d], and frontal view of head [e–f]. (a)
|
||
<taxonomicName id="33CC4D07FFA2AE2DFC57E644AD9CF8CA" authorityName="Martin" authorityYear="1910" box="[1017,1262,1807,1830]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA2AE2DFC57E644AD9CF8CA" box="[1017,1262,1807,1830]" italics="true" pageId="4" pageNumber="441">Chlorogomphus auratus</emphasis>
|
||
</taxonomicName>
|
||
, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e)
|
||
<emphasis id="C6B8EA96FFA2AE2DFDDFE664AB8AF8A8" box="[625,760,1838,1860]" italics="true" pageId="4" pageNumber="441">
|
||
<taxonomicName id="33CC4D07FFA2AE2DFDDFE664AB86F8A8" authorityName="Kompier & Karube" authorityYear="2018" box="[625,756,1838,1860]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="canhvang">C. canhvang</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype; (c–d, f)
|
||
<emphasis id="C6B8EA96FFA2AE2DFC1DE664AD48F8A8" box="[947,1082,1838,1860]" italics="true" pageId="4" pageNumber="441">
|
||
<taxonomicName id="33CC4D07FFA2AE2DFC1DE664AD44F8A8" authorityName="Kompier & Karube" authorityYear="2018" box="[947,1078,1838,1860]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="canhvang">C. canhvang</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="BCD6650FFFA2AE2CFF69E63AA8EFFDF4" lastPageId="5" lastPageNumber="442" pageId="4" pageNumber="441" type="diagnosis">
|
||
<paragraph id="F4733684FFA2AE2CFF69E63AAC45FEAC" blockId="4.[151,1437,1905,2038]" lastBlockId="5.[151,1436,151,536]" lastPageId="5" lastPageNumber="442" pageId="4" pageNumber="441">
|
||
<emphasis id="C6B8EA96FFA2AE2DFF69E63AA8A2F866" bold="true" box="[199,464,1905,1930]" pageId="4" pageNumber="441">Differential diagnosis.</emphasis>
|
||
The male is similar to the male of
|
||
<taxonomicName id="33CC4D07FFA2AE2DFCDEE63AADF0F865" authorityName="Martin" authorityYear="1910" box="[880,1154,1905,1930]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA2AE2DFCDEE63AADF0F865" box="[880,1154,1905,1930]" italics="true" pageId="4" pageNumber="441">Chlorogomphus auratus</emphasis>
|
||
</taxonomicName>
|
||
, but easily separated by details of the anal appendages (
|
||
<figureCitation id="6CF72A01FFA2AE2DFE57E6DEABEBF842" box="[505,665,1941,1966]" captionStart="FIGURE 2" captionStartId="3.[151,250,1937,1959]" captionTargetBox="[159,1405,214,1851]" captionTargetId="figure@3.[150,1436,213,1885]" captionTargetPageId="3" captionText="FIGURE 2. Comparison of anal appendages [a–b, e–f] and vesica spermalis [c–d, g–h] of Chlorogomphus auratus (Tam Dao NP, Vinh Phuc Prov.) [a–d] and C. canhvang, holotype [e–h]. (a, e) dorsal view; (b, f) lateral view; (c, g) lateral view; (d, h) ventral view of terminal segment." httpUri="https://zenodo.org/record/1199852/files/figure.png" pageId="4" pageNumber="441">Fig. 2a–b, e–f</figureCitation>
|
||
) and protusion on posterior margin of S10 (
|
||
<figureCitation id="6CF72A01FFA2AE2DFB24E6DEADACF842" box="[1162,1246,1941,1966]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="4" pageNumber="441">Fig. 3d</figureCitation>
|
||
). The epiproct is much deeper incised in
|
||
<taxonomicName id="33CC4D07FFA2AE2DFE07E6F1AB4BF83E" authorityName="Kompier & Karube" authorityYear="2018" box="[425,569,1977,2002]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA2AE2DFE07E6F1AB4BF83E" box="[425,569,1977,2002]" italics="true" pageId="4" pageNumber="441">C. canhvang</emphasis>
|
||
</taxonomicName>
|
||
, forming two parallel branches as long as the distance between them, more slender than in
|
||
<taxonomicName id="33CC4D07FFA2AE2DFEEAE695A8C8F819" authorityName="Martin" authorityYear="1910" box="[324,442,2014,2037]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="441" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA2AE2DFEEAE695A8C8F819" box="[324,442,2014,2037]" italics="true" pageId="4" pageNumber="441">C. auratus</emphasis>
|
||
</taxonomicName>
|
||
and with the apical teeth directed more dorsally. In addition, the protrusion between the cerci formed by the posterior margin of S10 is only small and not visible in lateral view in
|
||
<taxonomicName id="33CC4D07FFA3AE2CFB20E1D3AC69FF5C" authorityName="Kompier & Karube" authorityYear="2018" box="[1166,1307,151,176]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA3AE2CFB20E1D3AC69FF5C" box="[1166,1307,151,176]" italics="true" pageId="5" pageNumber="442">C. canhvang</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is clearly visible and stout in
|
||
<taxonomicName id="33CC4D07FFA3AE2CFE72E1F6AB23FF38" authorityName="Martin" authorityYear="1910" box="[476,593,189,212]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA3AE2CFE72E1F6AB23FF38" box="[476,593,189,212]" italics="true" pageId="5" pageNumber="442">C. auratus</emphasis>
|
||
</taxonomicName>
|
||
. The shape of the female vertex is also different between the two species. It is postero-dorsally rounded in
|
||
<taxonomicName id="33CC4D07FFA3AE2CFE43E1ABAB16FF14" authorityName="Martin" authorityYear="1910" box="[493,612,224,248]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA3AE2CFE43E1ABAB16FF14" box="[493,612,224,248]" italics="true" pageId="5" pageNumber="442">C. auratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="6CF72A01FFA3AE2CFDDDE194ABB6FF14" box="[627,708,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="5" pageNumber="442">Fig. 3a</figureCitation>
|
||
), but V-shaped in
|
||
<taxonomicName id="33CC4D07FFA3AE2CFC3EE1ABAD6CFF14" authorityName="Kompier & Karube" authorityYear="2018" box="[912,1054,223,248]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA3AE2CFC3EE1ABAD6CFF14" box="[912,1054,223,248]" italics="true" pageId="5" pageNumber="442">C. canhvang</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="6CF72A01FFA3AE2CFB83E194ADF3FF14" box="[1069,1153,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1777,1799]" captionTargetBox="[170,1422,531,1765]" captionTargetId="figure@4.[170,1424,511,1765]" captionTargetPageId="4" captionText="FIGURE 3. Comparison of antero-dorsal view of female vertex [a–b], left lateral view of male secondary genitalia [c], posterior view of male posteriormargin of S10[d], and frontalview of head[e–f]. (a) Chlorogomphus auratus, 22-V-2015, Cuc Phuong National Park, Ninh Binh Prov.; (b, e) C. canhvang, paratype; (c–d, f) C. canhvang, holotype." httpUri="https://zenodo.org/record/1199854/files/figure.png" pageId="5" pageNumber="442">Fig. 3b</figureCitation>
|
||
). The antero-dorsal spots on S4–7 appear to be more similar in size in
|
||
<taxonomicName id="33CC4D07FFA3AE2CFD09E04EAA52FEF0" authorityName="Martin" authorityYear="1910" box="[679,800,261,284]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA3AE2CFD09E04EAA52FEF0" box="[679,800,261,284]" italics="true" pageId="5" pageNumber="442">C. auratus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="6CF72A01FFA3AE2CFC9CE04FAAF4FEF1" box="[818,902,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="5" pageNumber="442">Fig. 1b</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="33CC4D07FFA3AE2CFB8AE04EADC6FEF1" authorityName="Kompier & Karube" authorityYear="2018" box="[1060,1204,260,285]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA3AE2CFB8AE04EADC6FEF1" box="[1060,1204,260,285]" italics="true" pageId="5" pageNumber="442">C. canhvang</emphasis>
|
||
</taxonomicName>
|
||
the spots on S7 are distinctly larger than those on S4–6 (
|
||
<figureCitation id="6CF72A01FFA3AE2CFD9DE06CABEEFEAC" box="[563,668,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1937,1959]" captionTargetBox="[151,1436,761,1912]" captionTargetId="figure@2.[151,1436,761,1912]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Chlorogomphus auratus [a–c] and C. canhvang [d–f]. (a) male, 23-VII-2017, Xuan Son National Park, Phu Tho Province; (b–c) female, 22-V-2015, Cuc Phuong National Park, Ninh Binh Province; (d) holotype male; (e) paratype female; (f) female, 4-V-2016, Thua Thien—Hue Province (posed for photo)." httpUri="https://zenodo.org/record/1199850/files/figure.png" pageId="5" pageNumber="442">Fig. 1e–f</figureCitation>
|
||
), although the validity of this character needs further study.
|
||
</paragraph>
|
||
<paragraph id="F4733684FFA3AE2CFF69E007A8EFFDF4" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="442">
|
||
<emphasis id="C6B8EA96FFA3AE2CFF69E007A8B3FE89" bold="true" box="[199,449,332,357]" pageId="5" pageNumber="442">Habitat and ecology.</emphasis>
|
||
<taxonomicName id="33CC4D07FFA3AE2CFE65E007AB87FE89" authorityName="Kompier & Karube" authorityYear="2018" box="[459,757,332,357]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="canhvang">
|
||
<emphasis id="C6B8EA96FFA3AE2CFE65E007AB87FE89" box="[459,757,332,357]" italics="true" pageId="5" pageNumber="442">Chlorogomphus canhvang</emphasis>
|
||
</taxonomicName>
|
||
was observed at small rocky streams under dense primary forest cover in low mountains (
|
||
<quantity id="33349B61FFA3AE2CFE59E024AB02FE64" box="[503,624,367,392]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="5" pageNumber="442" unit="m" value="400.0" valueMax="500.0" valueMin="300.0">300–500m</quantity>
|
||
asl). The males flew on sunny days, but females were also active under cloudy conditions. Although the same locations were also visited several times in June under good and sunny conditions, no specimens could be located, so it is likely that the flight period of the species is during April–May, but does not extend into June. By contrast,
|
||
<taxonomicName id="33CC4D07FFA3AE2CFDD4E096AB82FE18" authorityName="Martin" authorityYear="1910" box="[634,752,477,500]" class="Insecta" family="Chlorogomphidae" genus="Chlorogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="442" phylum="Arthropoda" rank="species" species="auratus">
|
||
<emphasis id="C6B8EA96FFA3AE2CFDD4E096AB82FE18" box="[634,752,477,500]" italics="true" pageId="5" pageNumber="442">C. auratus</emphasis>
|
||
</taxonomicName>
|
||
flies at least from early May to late July in northern
|
||
<collectingCountry id="8CDB7614FFA3AE2CFA93E097ACEEFE19" box="[1341,1436,476,501]" name="Vietnam" pageId="5" pageNumber="442">Vietnam</collectingCountry>
|
||
(T. Kompier
|
||
<emphasis id="C6B8EA96FFA3AE2CFE86E34AA8FFFDF4" box="[296,397,511,536]" italics="true" pageId="5" pageNumber="442">pers. obs</emphasis>
|
||
.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |