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<mods:title id="6F0E976F0049872AB5B2D06185B162BD">The family Lecithoceridae (Lepidoptera, Gelechioidea) from Bioko Island (Equatorial Guinea), with descriptions of five new species</mods:title>
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<mods:namePart id="C2F76122E6A4A66EECC1FF8C51DEF76D">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="10AA8452441BCAEDE1B8DDD38E1F5C08">Bioresource and Environmental Center, Incheon National University, Incheon, 22012 Korea.</mods:affiliation>
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<mods:namePart id="A21F7E4CD4AB82BD2A29A7B91AAAFF62">Karisch, Timm</mods:namePart>
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<mods:affiliation id="9ACAF9A930A616EDF87591D39754DD92">Museum für Naturkunde und Vorgeschichte, Askanische Strasse 32, 06842 Dessau-Rosslau, Germany. timm. karisch @ naturkunde. dessau. de, https: // orcid. org / 0000 - 0003 - 1281 - 7512</mods:affiliation>
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<heading id="D0BB6D78EF4E6851CEF8FB61FDDF9F98" bold="true" box="[151,526,1158,1185]" fontSize="11" level="1" pageId="7" pageNumber="588" reason="1">
<taxonomicName id="4C4CA197EF4E6851CEF8FB61FDDF9F98" ID-CoL="9FSGJ" authority="Park, 2018" authorityName="Park" authorityYear="2018" box="[151,526,1158,1185]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="bilobella">
<emphasis id="B9380606EF4E6851CEF8FB61FDDF9F98" bold="true" box="[151,526,1158,1185]" pageId="7" pageNumber="588">
<emphasis id="B9380606EF4E6851CEF8FB61FEAE9F99" bold="true" box="[151,383,1158,1185]" italics="true" pageId="7" pageNumber="588">Thubdora bilobella</emphasis>
<bibRefCitation id="EFDDA7E5EF4E6851CFE8FB60FDDF9F98" author="Park, K. - T." box="[391,526,1158,1185]" pageId="7" pageNumber="588" pagination="1085 - 1093" refId="ref5632" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park, 2018</bibRefCitation>
</emphasis>
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<paragraph id="8BF3DA14EF4E6851CEF8FB4CFEFB9FFE" blockId="7.[151,526,1158,1222]" box="[151,298,1195,1222]" pageId="7" pageNumber="588">
(
<figureCitation id="1377C691EF4E6851CEF0FB4CFEF39FFE" box="[159,290,1195,1222]" captionStart="FIGURE 6" captionStartId="8.[151,250,1854,1878]" captionTargetBox="[161,1414,387,1814]" captionTargetId="figure-111@8.[151,1436,381,1831]" captionTargetPageId="8" captionText="FIGURE 6. Thubdora bilobella Park, 2018: A, adult; B, male genitalia, slide no. 3824; C, aedeagus; D, abdomen, with closeup segment VII-VIII; E male genitalia (with right valva) of the holotype. Scale bar for the genitalia with aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5057137" httpUri="https://zenodo.org/record/5057137/files/figure.png" pageId="7" pageNumber="588">Figs. 6AE</figureCitation>
)
</paragraph>
</subSubSection>
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<treatmentCitation id="0AEDFC05EF4E6851CEF8FB08FDC29E3F" author="Park, K. - T." box="[151,531,1263,1287]" page="1089" pageId="7" pageNumber="588" year="2018">
<taxonomicName id="4C4CA197EF4E6851CEF8FB08FDC29E3F" ID-CoL="9FSGJ" authority="Park, 2018: 1089" authorityName="Park" authorityPageNumber="1089" authorityYear="2018" box="[151,531,1263,1287]" class="Insecta" family="Lecithoceridae" genus="Thubdora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="588" phylum="Arthropoda" rank="species" species="bilobella">
<emphasis id="B9380606EF4E6851CEF8FB08FE8A9E3F" box="[151,347,1263,1287]" italics="true" pageId="7" pageNumber="588">Thubdora bilobella</emphasis>
<bibRefCitation id="EFDDA7E5EF4E6851CF0DFB08FDC29E3F" author="Park, K. - T." box="[354,531,1263,1287]" pageId="7" pageNumber="588" pagination="1085 - 1093" refId="ref5632" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park, 2018: 1089</bibRefCitation>
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;
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<bibRefCitation id="EFDDA7E5EF4E6851CC72FB08FD289E3F" author="Park, K. - T. &amp; Mey, W. &amp; Koo, J. M. &amp; De Prins, J. &amp; Akite, P. &amp; Cho, S. W." box="[541,761,1263,1287]" pageId="7" pageNumber="588" pagination="451 - 487" refId="ref5861" refString="Park, K. - T., Mey, W., Koo, J. M., De Prins, J., Akite, P. &amp; Cho, S. W. (2020) Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo. Zootaxa, 4759 (4), 451 - 487. https: // doi. org / 10.11646 / zootaxa. 4759.4.1" type="journal article" year="2020">Park et al., 2020: 471</bibRefCitation>
</treatmentCitation>
. TL:
<collectingCountry id="F35B9A84EF4E6851CD41FB08FC4A9E3F" box="[814,923,1263,1287]" name="Cameroon" pageId="7" pageNumber="588">Cameroon</collectingCountry>
.
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</paragraph>
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<subSubSection id="C356899FEF4E6851CEF8FAD2FD029E83" pageId="7" pageNumber="588" type="materials_examined">
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<emphasis id="B9380606EF4E6851CEF8FAD2FE539E77" bold="true" box="[151,386,1333,1359]" pageId="7" pageNumber="588">Material examined.</emphasis>
<materialsCitation id="3B24D049EF4E6851CFE4FAD2FCC59EA0" collectingDate="1994-08-18" collectionCode="T" country="Equatorial Guinea" county="Ose" elevation="1200" latitude="3.3750002" location="Island of Bioko" longLatPrecision="21" longitude="8.541667" municipality="Gran Caldera Volcanica de Luba" pageId="7" pageNumber="588" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4A119DEF4E6851CFE4FAD2FE7E9E77" box="[395,431,1333,1359]" pageId="7" pageNumber="588" type="male">1♂</specimenCount>
,
<collectingCountry id="F35B9A84EF4E6851CFD1FAD2FD5E9E77" box="[446,655,1333,1359]" name="Equatorial Guinea" pageId="7" pageNumber="588">Equatorial Guinea</collectingCountry>
,
<location id="8E938CCFEF4E6851CCF3FAD2FC9C9E77" LSID="urn:lsid:plazi:treatment:03E56B02EF4E6851CEF8FB61FC5B9DE3:8E938CCFEF4E6851CCF3FAD2FC9C9E77" box="[668,845,1333,1359]" country="Equatorial Guinea" county="Ose" latitude="3.3750002" longLatPrecision="21" longitude="8.541667" municipality="Gran Caldera Volcanica de Luba" name="Island of Bioko" pageId="7" pageNumber="588">Island of Bioko</location>
(former Fernando Póo),
<collectingMunicipality id="6B97406EEF4E6851CA04FAD2FF009E4B" pageId="7" pageNumber="588">Gran Caldera Volcanica de Luba</collectingMunicipality>
u
<collectingCounty id="6292A298EF4E6851CE80FABEFECC9E4B" box="[239,285,1369,1395]" pageId="7" pageNumber="588">Ose</collectingCounty>
, outer NNE slope about
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<quantity id="4CB477F1EF4E6851CC5AFABEFDBE9E4B" box="[565,623,1369,1395]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="588" unit="km" value="4.0">4 km</quantity>
SSW Ruiché
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,
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S of Luba
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, higher altitude primary rain forest,
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<elevation id="00613D27EF4E6851CB0BFABEFF7E9EA0" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="588" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<geoCoordinate id="EE78BCD3EF4E6851CED3FA9AFE919EA0" box="[188,320,1405,1432]" degrees="3" direction="north" minutes="22" orientation="latitude" pageId="7" pageNumber="588" precision="15" seconds="30" value="3.3750002">3°2230” N</geoCoordinate>
<geoCoordinate id="EE78BCD3EF4E6851CF27FA9AFE1A9EA0" box="[328,459,1405,1432]" degrees="8" direction="east" minutes="32" orientation="longitude" pageId="7" pageNumber="588" precision="15" seconds="30" value="8.541667">8°3230” E</geoCoordinate>
,
<date id="FFF2FCD4EF4E6851CFB7FA9AFDB49EAF" box="[472,613,1405,1431]" pageId="7" pageNumber="588" value="1994-08-18">
<collectingDate id="EFB6053CEF4E6851CFB7FA9AFDB49EAF" box="[472,613,1405,1431]" pageId="7" pageNumber="588" value="1994-08-18">18 viii 1994</collectingDate>
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, at light, leg.
<collectionCode id="ED5D42D1EF4E6851CD6DFA99FCC59EA0" box="[770,788,1406,1432]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="588">T</collectionCode>
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. Karisch, gen. slide no. 3824/Karisch;
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<specimenCount id="9D4A119DEF4E6851CAA0FA9AFB229EAF" box="[1231,1267,1405,1431]" pageId="7" pageNumber="588" type="male">1♂</specimenCount>
, same locality,
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<collectingDate id="EFB6053CEF4E6851CEF8FA46FEF09E83" box="[151,289,1441,1467]" pageId="7" pageNumber="588" value="1994-08-20">20.viii.1994</collectingDate>
</date>
, in coll. MNVD, coll.
<collectorName id="26B9BFC2EF4E6851CC72FA46FDA59E83" box="[541,628,1441,1467]" pageId="7" pageNumber="588">Karisch</collectorName>
(
<collectionCode id="ED5D42D1EF4E6851CCECFA46FD169E83" box="[643,711,1441,1467]" pageId="7" pageNumber="588">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C356899FEF4E6851CEA8FA22FC729DAC" pageId="7" pageNumber="588" type="discussion">
<paragraph id="8BF3DA14EF4E6851CEA8FA22FBB99D57" blockId="7.[151,1437,1333,1756]" pageId="7" pageNumber="588">
<emphasis id="B9380606EF4E6851CEA8FA22FEEA9EE7" bold="true" box="[199,315,1477,1503]" pageId="7" pageNumber="588">Remarks.</emphasis>
This species was originally described from
<collectingCountry id="F35B9A84EF4E6851CD42FA22FC739EE7" box="[813,930,1477,1503]" name="Cameroon" pageId="7" pageNumber="588">Cameroon</collectingCountry>
(
<bibRefCitation id="EFDDA7E5EF4E6851CDDEFA22FBFE9EE7" author="Park, K. - T." box="[945,1071,1477,1503]" pageId="7" pageNumber="588" pagination="1085 - 1093" refId="ref5632" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park, 2018</bibRefCitation>
) and also reported from
<collectingCountry id="F35B9A84EF4E6851CB2AFA22FA4D9EE7" box="[1349,1436,1477,1503]" name="Uganda" pageId="7" pageNumber="588">Uganda</collectingCountry>
by
<bibRefCitation id="EFDDA7E5EF4E6851CED5FA0EFE509D3B" author="Park, K. - T. &amp; Mey, W. &amp; Koo, J. M. &amp; De Prins, J. &amp; Akite, P. &amp; Cho, S. W." box="[186,385,1513,1540]" pageId="7" pageNumber="588" pagination="451 - 487" refId="ref5861" refString="Park, K. - T., Mey, W., Koo, J. M., De Prins, J., Akite, P. &amp; Cho, S. W. (2020) Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo. Zootaxa, 4759 (4), 451 - 487. https: // doi. org / 10.11646 / zootaxa. 4759.4.1" type="journal article" year="2020">Park et al. (2020)</bibRefCitation>
. This species can be distinguished from its allies by the shape of the uncus, the valva, and the aedeagus in the male genitalia. Little differences from the
<typeStatus id="54F764B6EF4E6851CD42F9E9FC8C9D10" box="[813,861,1550,1576]" pageId="7" pageNumber="588">type</typeStatus>
specimen of the species (
<figureCitation id="1377C691EF4E6851CAEDF9EAFB099D10" box="[1154,1240,1549,1576]" captionStart="FIGURE 5" captionStartId="6.[151,250,1980,2004]" captionTargetBox="[152,1434,631,1944]" captionTargetId="figure-157@6.[151,1436,628,1956]" captionTargetPageId="6" captionText="FIGURE 5. Thubdora angustiala Park &amp; Karisch, sp. nov.: A, adult, holotype; B, dorsal surface of head and thorax; C, female genitalia, slide no, 3827/Karisch. Scale bar for the genitalia with aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5057133" httpUri="https://zenodo.org/record/5057133/files/figure.png" pageId="7" pageNumber="588">Fig. 5E</figureCitation>
) are found in the male genitalia, with the uncus less directed outwardly and the triangular caudal plate of juxta smaller, However, those differences are considered to be infraspecific variation. A further study is needed.
</paragraph>
<paragraph id="8BF3DA14EF4E6851CEA8F99EFC729DAC" blockId="7.[151,1437,1333,1756]" box="[199,931,1657,1684]" pageId="7" pageNumber="588">
<emphasis id="B9380606EF4E6851CEA8F99EFEBD9DAB" box="[199,364,1657,1683]" italics="true" pageId="7" pageNumber="588">Male genitalia</emphasis>
(
<figureCitation id="1377C691EF4E6851CF14F99EFDD09DAC" box="[379,513,1657,1684]" captionStart="FIGURE 6" captionStartId="8.[151,250,1854,1878]" captionTargetBox="[161,1414,387,1814]" captionTargetId="figure-111@8.[151,1436,381,1831]" captionTargetPageId="8" captionText="FIGURE 6. Thubdora bilobella Park, 2018: A, adult; B, male genitalia, slide no. 3824; C, aedeagus; D, abdomen, with closeup segment VII-VIII; E male genitalia (with right valva) of the holotype. Scale bar for the genitalia with aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5057137" httpUri="https://zenodo.org/record/5057137/files/figure.png" pageId="7" pageNumber="588">Figs. 6C, D</figureCitation>
): See also
<bibRefCitation id="EFDDA7E5EF4E6851CC17F99EFD269DAB" author="Park, K. - T." box="[632,759,1657,1683]" pageId="7" pageNumber="588" pagination="1085 - 1093" refId="ref5632" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park (2018</bibRefCitation>
, Figs. 15, 15a).
</paragraph>
</subSubSection>
<subSubSection id="C356899FEF4E6851CEA8F97AFC5B9DE3" pageId="7" pageNumber="588" type="distribution">
<paragraph id="8BF3DA14EF4E6851CEA8F97AFC5B9DE3" blockId="7.[151,1437,1333,1756]" pageId="7" pageNumber="588">
<emphasis id="B9380606EF4E6851CEA8F97AFEB09D8F" bold="true" box="[199,353,1693,1719]" pageId="7" pageNumber="588">Distribution.</emphasis>
<collectingCountry id="F35B9A84EF4E6851CF09F97AFDE39D8F" box="[358,562,1693,1719]" name="Equatorial Guinea" pageId="7" pageNumber="588">Equatorial Guinea</collectingCountry>
(new record; Isl. Bioko). The species was originally described from
<collectingCountry id="F35B9A84EF4E6851CB47F97AFA4C9D8F" box="[1320,1437,1693,1719]" name="Cameroon" pageId="7" pageNumber="588">Cameroon</collectingCountry>
(
<bibRefCitation id="EFDDA7E5EF4E6851CEF0F926FECD9DE3" author="Park, K. - T." box="[159,284,1729,1755]" pageId="7" pageNumber="588" pagination="1085 - 1093" refId="ref5632" refString="Park, K. - T. (2018) A new genus Thubdora Park, gen. nov. and seven new species of the subfamily Torodorinae (Lepidoptera, Lecithoceridae) from Cameroon and Rep. of Congo, Africa. Journal of Asia-Pacific Entomology, 21, 1085 - 1093. https: // doi. org / 10.1016 / j. aspen. 2018.07.018" type="journal article" year="2018">Park, 2018</bibRefCitation>
) and was also reported from
<collectingCountry id="F35B9A84EF4E6851CC0DF926FD689DE3" box="[610,697,1729,1755]" name="Uganda" pageId="7" pageNumber="588">Uganda</collectingCountry>
(
<bibRefCitation id="EFDDA7E5EF4E6851CCA8F926FCAF9DE3" author="Park, K. - T. &amp; Mey, W. &amp; Koo, J. M. &amp; De Prins, J. &amp; Akite, P. &amp; Cho, S. W." box="[711,894,1729,1756]" pageId="7" pageNumber="588" pagination="451 - 487" refId="ref5861" refString="Park, K. - T., Mey, W., Koo, J. M., De Prins, J., Akite, P. &amp; Cho, S. W. (2020) Fourteen new species of the genus Thubdora Park, 2018 (Lepidoptera: Lecithoceridae) from Uganda, and three new combinations in Ptilothyris Walsingham, 1897 from DR Congo. Zootaxa, 4759 (4), 451 - 487. https: // doi. org / 10.11646 / zootaxa. 4759.4.1" type="journal article" year="2020">Park et al. 2020</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>